Coryssiphus cinerascens Simon, 1903a: 32 Dippenaar-Schoeman et al ., 2021: 9 Coryssiphus unicolor Simon, 1903a: 32 Dippenaar-Schoeman et al ., 2021: 11 A revision of the genus Coryssiphus Simon, 1903, with its transfer to Systariinae (Araneae: Miturgidae) Bosselaers, Jan Zootaxa 2024 2024-02-20 5415 1 153 168 YSTG Simon, 1903 Simon 1903 [151,610,151,178] Arachnida Liocranidae Coryssiphus Animalia Araneae 9 162 Arthropoda species cinerascens   Figs 1B, 2E–G, 5, 6C–E, 8      Coryssiphus cinerascens Simon, 1903a: 32( holotype♂:  SOUTH AFRICA:“Cap! Prom. Bonae Spei!” (MNHN 18821);  Dippenaar-Schoeman et al., 2021: 9, 4 figs.     Coryssiphus unicolor Simon, 1903a: 32( holotype ♀:  SOUTH AFRICA:“Cap! Prom. Bonae Spei!” (MNHN 13003);  Dippenaar-Schoeman et al., 2021: 11. syn. nov.   Additional material examined.   SOUTH AFRICA:  Western Cape: 1♂, Cape Peninsula, Constantia, Vlakkenberg, fynbos near summit,  16 September–4 October 1992, leg. B. Heydenrych( RMCA 174990); 1♂, Samedata as previous but  4–12 October 1992, leg. Heydenrych( RMCA 174995);  1♀, Same data as previous but  19 October–8 November 1992( RMCA 177006);  1♀, Cape Province, Houtbaai, Tierbos, in litter,  December 1960, leg. N. Leleup( RMCA 131824);   FIGURE 6.A, B.  Coryssiphus praeustus, female RMCA 131898 (A), female RMCA 247653 (B, left), female RMCA 131872 (B, right). C–E.  C. cinerascens, female RMCA 177006 (C, left),  C. unicolor, holotype (C, right), male holotype (D), female RMCA 131824 (e). A. Vulva, dorsal; B, C. Epigynes. D. palp, ventral (left) and retrolateral (right). E. Vulva, dorsal. Scale bars: A, E: 100 µm; B, C: 250 µm; D: 0.5 mm. Abbreviations: ID, Insemination duct; Sss, Sickle-shaped sclerite; ST1, Spermatheca 1; ST2, Spermatheca 2 (“bursa”).  Additional data on material only examined from photographs.   SOUTH AFRICA:  Western Cape: 2♀, Cape Town, Kirstenbosch Botanical Gardens, 33°55’04’’S, 18°25’36’’E,  214 ma.s.l., leaf litter in Afromontane forest,  11 December 2012, leg. J.A. Neethling( NCA 2013/673); 1♂, Samelocality 33°59.216’S, 18°25.742’E,  165 ma.s.l., sifting litter, Afromontane forest,  4 October 2022, leg. C. Haddad, D. Harms, R. Booysen& J. Neethling( NMBA 18840);  1♀, Cape Town, Westernslope of Table Mountain,  6 November 1949, leg. B. Malkin( CAS 9033195); 3 imm. 2♀, Stellenbosch, Jonkershoek Nature Reserve, 33°59.584’S, 18°57.358’E,  405 ma.s.l., sifting litter, Afromontane forest,  4 October 2022, leg. C. Haddad, D. Harms, R. Booysen& J. Neethling( NMBA 18876).   Diagnosis.  Coryssiphus cinerascensis close to  C. praeustus, but differs from it by the smaller body size, the slightly shorter, sturdier legs, especially the tibiae of legs and palps ( Tables 1, 2), the shorter cymbium of the male palp ( Figs 5C, D, 6D), the epigyne that has a trapezoidal instead of bell-shaped central depression ( Figs 6C, 8E), and the ST2 that are oriented antero-laterally instead of almost longitudinally ( Figs 6E, 8G).   Description.Male ( holotypeunless mentioned otherwise). Somatic morphology as described for the genus. Total l (n = 3) 3.2–3.3; carapace l 1.6, w 1.3; sternum l 0.9, w 0.9. Fovea situated 1.1 from front, fovea l 0.3 ( Fig. 5A). Eye arrangement: AER w 0.34, PER w 0.47, MOQ frontal end 0.20, posterior end 0.26, longitudinal l 0.24. Legs relatively short and sturdy ( Table 1). Leg spination as in Fig. 2E; ventral side of ti I with 6 vsp, ti II with 5 vsp. Male palp with cylindrical ti, cy about twice as long as diameter of bulbus ( Table 2, Figs 5C, 6D). Leg measurements: fe pa ti mt ta Total I 1.47 0.66 1.60 1.18 0.71 5.62 II 1.24 0.58 1.24 0.92 0.60 4.58 III 0.97 0.39 0.79 0.87 0.58 3.60 IV 1.42 0.60 1.47 1.29 0.71 5.50 Female ( holotypeof  C. unicolorunless otherwise stated). Somatic morphology as described for the genus. Total l 4.7–5.2 (n = 3), carapace l 1.7, w 1.5; sternum l 1.0, w 0.9. Fovea situated 1.1 from front, fovea l 0.3 ( Fig. 8A). Eye arrangement: AER w 0.45, PER w 0.55, MOQ frontal end 0.21, posterior end 0.28, longitudinal l 0.26. Legs relatively short and sturdy ( Table 1). Leg spination as in Fig. 2F, G; ventral side of ti I with 6–7 vsp, ti II with 6 vsp. Epigyne with trapezoidal central depression and sclerotised lateral rims that diverge over their entire length ( Figs 6C, 8E). Long axis of ST2 oriented in antero-lateral direction ( Figs 6E, 8G). Leg measurements: fe pa ti mt ta Total I 1.53 0.71 1.60 1.05 0.74 5.62 II 1.39 0.63 1.37 0.92 0.68 5.00 III 1.26 0.53 1.05 0.95 0.63 4.42 IV 1.71 0.66 1.66 1.47 0.82 6.31   Distribution.At present only known from the CapePeninsula and Stellenbosch, South Africa, but most probably present within a radius of at least 60 kmfrom the latter (Haddad, pers. comm.). 4526325307 1992-09-16 1992-10-04 1992-09-16 RMCA B. Heydenrych South Africa Peninsula Vlakkenberg Constantia 9 162 RMCA 174990 1 Western Cape 4526325310 1992-10-04 1992-11-08 1992-10-04 RMCA Heydenrych South Africa Peninsula Same Constantia 9 162 RMCA 174995, RMCA 177006 2 2 Cape 4526325305 1992-10-04 1992-11-08 1992-10-04 RMCA Heydenrych South Africa Peninsula Same Constantia 9 162 RMCA 174995, RMCA 177006 2 2 Cape 4526325311 1992-10-04 1992-11-08 1992-10-04 RMCA Heydenrych South Africa Peninsula Same Constantia 9 162 RMCA 174995, RMCA 177006 2 2 Cape 4526325302 2012-12-11 NCA Kirstenbosch Botanical Gardens South Africa Town 214 -33.917778 Western Cape 19 18.426666 Western 10 163 NCA 2013 2 2 Western Cape 4526325312 2012-12-11 NCA Kirstenbosch Botanical Gardens South Africa Town 214 -33.917778 Western Cape 19 18.426666 Western 10 163 NCA 2013 2 2 Western Cape 4526325303 2012-12-11 NCA Kirstenbosch Botanical Gardens South Africa Town 214 -33.917778 Western Cape 19 18.426666 Western 10 163 NCA 2013 2 2 Western Cape 4526325304 2012-12-11 NCA Kirstenbosch Botanical Gardens South Africa Town 214 -33.917778 Western Cape 19 18.426666 Western 10 163 NCA 2013 2 2 Western Cape