Hydroporus bodemeyeri Ganglbauer
Bodemeyer, 1900: 139
Apfelbeck 1904: 374
Sahlberg 1913: 43
Bodemeyer 1927: 44
Zimmermann 1931: 9
H. tristis (Paykull, 1798))
Miller and Fery 1995: 409
Nilsson 2001: 159
Foster and Taylor 2006: 81
Darilmaz and Kiyak 2009: 1614
Hydroporus (s. str.) bodemeyeri : Zaitzev 1953: 155
Guéorguiev 1968: 34
Hydroporus ( Sternoporus ) bodemeyeri : Wewalka 1971: 211
Guéorguiev 1981: 407
Hydroporus ( Sternoporus ) collarti Guignot, 1949: 7
Wewalka 1971: 211
Taxonomic revision of the Hydroporus bodemeyeri species complex (Coleoptera: Dytiscidae) with a geometric morphometric analysis of body shape within the group
Hájek, Jirí
Fikácek, Martin
Journal of Natural History
2010
2010-06-18
44
27 - 28
1631
1657
7JVWQ
bodemeyeri Ganglbauer, 1900
bodemeyeri Ganglbauer
1900
[320,927,167,191]
Insecta
Dytiscidae
Hydroporus
Animalia
Coleoptera
11
1640
Arthropoda
subSpecies
bodemeyeri
bodemeyeri
Hydroporus bodemeyeriGanglbauerin Bodemeyer, 1900: 139(original description); Apfelbeck 1904: 374(notes); Sahlberg 1913: 43(distribution); Bodemeyer 1927: 44, 67 (distribution); Zimmermann 1931: 9(as synonym of H. tristis(Paykull, 1798)); Miller and Fery 1995: 409(description; partim); Nilsson 2001: 159(catalogue); 2003: 60 (catalogue); Foster and Taylor 2006: 81(distribution); Darilmaz and Kiyak 2009: 1614(catalogue; partim). Hydroporus(s. str.) bodemeyeri: Zaitzev 1953: 155(description); Guéorguiev 1968: 34(description). Hydroporus( Sternoporus) bodemeyeri: Wewalka 1971: 211(synonymy, distribution; partim.); Guéorguiev 1981: 407(catalogue, distribution); Hydroporus( Sternoporus) collarti Guignot, 1949: 7(original description) (synonymized by Wewalka 1971: 211).
Typelocality. H. bodemeyeri: “Karakeuy” [ Turkey, Bilecik prov., Karaköy, ca. 39°57′N 29°59′E] (see Remarks under this species); H. collarti: “Lesbos: Mytilini” [ Greece, Lesvos Isl., Mytilene, ca. 39°06′ N, 26°33′ E]. Type material. H. bodemeyeri: Lectotypemale ( NHMW), designated by Wewalka (1971): “ Asia minor/ Karakeuy / v.Bodemeyer [printed] // Bodemeyeri / Gangl., Type [handwritten] // Coll. Mus. Vindob. [printed] // Hydroporus/ Bodemeyeri / Gnglb., Type. [handwritten] // Collect. / Hauser [printed] // HOLO- [sic!, handwritten] / TYPUS [red label, printed]”. Paralectotypes( NHMW): two males, two females, same label data as lectotype. H. collarti: Holotypemale ( IRSNB): “ Coll. R. I. Sc. N. B./ Grèce/ I. Lesbos: Mytilini / N. W. 25-250., 4-V-’33 / A. d’Orchymont[printed] // Ƌ [printed] // Hydroporus/ ( Sternoporus) / Collarti Guign. [handwritten] // TYPE [printed, pink label with black frame]”. Paratypes( IRSNB): one male, one female, “ Coll. R. I. Sc. N. B./ Grèce/ I. [ Island] Chios[ Khios]: Karyaes/ S. W. 140 m., 2-V-1933/ A. d’Orchymont[printed] // Para- / type [red label, printed]”; two females, “ Coll. R. I. Sc. N. B./ Grèce/ Samos: Platanos/ Velanidia 3-VI-1933/ A. d’Orchymont[printed] // Para- / type [red label, printed]”; one male, three females, “ Coll. R. I. Sc. N. B./ Grèce/ I. [ Island] Ikaria: Xylocirti[ Xylosírtis] / 75 m. 25-V-1933/ A. d’Orchymont[printed] // Para- / type [red label, printed]”. Additional material examined. Greece: two males, Lésvos Island, Ayiásos-Megalokhóri, Mount Olympos, 55 m, 6 June 1996, M.A. Jächleg. ( NHMW); one male, Lésvos Island, 10 kmnorth of Ayiásos, near Pigi Karini, 7 June 1996, M.A. Jächleg. ( NHMW); nine males, 16 females, Lésvos Island, 4 kmsouth of Pétra, S Lafionas, 150 m, 8 June 1996, M.A. Jächleg. ( HFCB, NMPC, NHMW); one male, two females, Lésvos Island, 2 kmwest of Plomari, 30 m, 9 June 1996, M.A. Jächleg. ( HFCB, NHMW). Turkey: one female, Abant-Geb.[Abant Mts., Bolu prov.], 30 May 1964, H. Korgeleg. ( NHMB); one male, Ephesus[near Selçuk, Izmir prov., ca. 37°56′ N, 27°19′ E]; U. Sahlbergleg. ( NHMW); two males, one female, Izmir [prov.], Yamanlar Da g[mt.], 700 m, 18 May 1991, S. Schödlleg. ( NHMW); four males, two females, Izmir [prov.], south of Tire, Aydin Berge, 900 m, 20 May 1991, M.A. Jächleg. (14) ( NHMW); one female, K iz ilcahamam [ Ankara prov., ca. 40°29′ N, 32°38′ E], 13 June to 16 June 1968, C. Holzschuhleg. ( NHMW); two males, one female, same label data, but 15 June 1968, G. Wewalkaleg. ( GWCW, NHMB); one male, Kappadokien[Cappadocia, Nev sehir prov.], Ürgüp, 30 September to 2 October 1984, L. Dobbersteinleg. ( LHCB); one female, north of Antalya[ Antalya prov.], ca. 700 m, stream, 4 April 1985, H. Feryleg. ( HFCB); one male, Antalya prov., south of Kemer, 50 m, 25 May 1991, M.A. Jächleg. ( NHMW); two females, Adana prov., Büyüksofulu[ca. 37°33′ N 35°11′ E], spring, 1 July 1997, J. Hájekleg. ( NMPC). Measurements.Body length: 3.15–3.50 mm(mean value 3.36 mm), width: 1.45–1.60 mm(mean value 1.54 mm). Description.Body flat, elongate, rather parallel sided ( Figure 3E), sides a little rounded. Elytra only weakly tapering to apex, with maximum width near midlength. Pronoto-elytral angle quite distinct. Upper surface blackish-brown, sides of pronotum and head often paler. Mouth appendages and legs testaceous. Ventral part blackish, head with paler gula. Head relatively broad, clypeus broadly rounded. Head on surface with two shallow depressions near anterior margin, and a groove along inner side of each eye. Punctation comprising sparse, almost regularly distributed punctures (denser in depressions), slightly smaller on clypeus and bigger on frons and vertex. Distance between punctures about twice their diameter. Microreticulation consisting of polygonal meshes with diameter smaller than that of punctures. Pronotum of rather variable shape. Usually with maximum width near midlength, sides more or less evenly curved over entire length; but in some specimens sides rather subparallel in posterior half. Lateral beading distinct, regular. Pronotum with a line of coarse setigerous punctures along anterior margin, and with shallow coarsely punctured posterolateral depressions. Disc of pronotum with sparse and fine setigerous punctures similar to those on head, but their diameter slightly smaller, and distance between punctures nearly three times their diameter. Microreticulation similar to that of head, but slightly less impressed. Elytra with sides almost straight behind shoulders, widest near midlength, then weakly tapering to apex; marginal rim visible at shoulders in dorsal view. Elytral disc with two rather indistinct punctured lines and with quite regularly distributed moderately large setigerous punctures between these lines. Distance between punctures about twice the diameter. Ventral part microreticulated. Reticulation indistinct on meso- and metaventrite, but stronger on metacoxal plates and abdominal ventrites. Metaventrite, metacoxal plates and two basal abdominal ventrites with large setigerous punctures. Other abdominal ventrites with sparse fine punctures. Male: dorsal surface shiny. Claws of fore tarsi broadened, strongly hooked. Median lobe of aedeagus regularly tapering to apex in ventral view, apical part relatively short, slightly turned to right ( Figure 2E). In lateral view, median lobe strongly inflexed in middle, apex long and nearly straight. Paramere as in Figure 2F. Female: pronotum slightly broader than in male. Microreticulation of elytra more distinct than in males and surface subopaque. Claws of fore legs thin, only slightly curved. Remarks.The typelocality of Hydroporus bodemeyeriwas stated as “In Mehrzahl bei Karakeuy gesammelt.” [Most specimens collected at Karakeuy (= Karaköy village).] (Ganglbauer in Bodemeyer 1900: 140). According to the Microsoft Encarta World Atlas 2000, there are about 50 different villages named “Karaköy” in Turkey. Miller and Fery (1995: 410)assumed the typelocality to be the village lying ca. 80 kmnorth of Adanaand 60 kmeast of Bulghar Dagh [= Bolkar Daglari Mountains]. We believe that the assumption is erroneous for two reasons. First, Bodemeyer (1900: 98)used the following abbreviation for the locality: “Ka. = Karakeuy, Station der anatolischen Bahn.” [station of the Anatolian railway]. There is no railway around the place cited by Miller and Fery (1995: 410). Second, Bodemeyer (1900: 24)presented precise information about Karaköy within the chapter “Reise von Konstantinopel nach Konia längs der anatolischen Bahn” [Travel from Constantinople (= Istanbul) to Konyaalong the Anatolian railway], and placed Karaköy between Bilecikand Eski sehir “...Steigt die Bahn von Biledjik aus schon steil zur Hochebene hinan, so hat man in Karakeuy 620 Meter, in Eski- Chéhir schon 800 Meter erreicht...” [...the railway runs up steeply from Bilecikto the plateau, and therefore it reaches 620 meters in Karaköy, and 800 meters already in Eski sehir...]. Therefore we assume that the typelocality of H. bodemeyeriis the Karaköy villagein Bilecik Province, ca. 22 kmsouth of Bilecik, geographic co-ordinates 39°57′ N, 29°59′ E. Distribution.The occurrence of the nominotypical subspecies is here confined to Greek Islands (Ikaría, Khios, Lésvos, Samos) and to the western half of Asian Turkey( Adana, Ankara, Antalya, Bolu, Izmir and Nev sehir provinces) ( Figure 6).
NHMW
Turkey
Mus
11
1640
1
Mus
lectotype
NHMW
Turkey
Mus
11
1640
2
1
1
Mus
paralectotype
IRSNB
Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & I. Lesbos & A. d'Orchymont
Greece
Grece
11
1640
1
holotype
1933-05-02
IRSNB
Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & A. d'Orchymont
Greece
Karyaes
140
Chios
Khios
11
1640
142
1
1
paratype
1933-06-03
IRSNB
Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & A. d'Orchymont
Greece
Velanidia
140
true
Samos
Platanos
11
1640
1
1
paratype
1933-05-25
Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & A. d'Orchymont
Greece
Xylosirtis
75
Ikaria
Xylocirti
11
1640
77
1
1
paratype
1996-06-06
NHMW
M. A. Jach
Greece
Lesvos Island
55
Mount Olympos
Ayiasos-Megalokhori
11
1640
1
1
1996-06-07
NHMW
M. A. Jach
Greece
Lesvos Island
Pigi Karini
Ayiasos
11
1640
1
1
1996-06-08
HFCB, NMPC, NHMW
S Lafionas & M. A. Jach
Greece
150
Petra
Lesvos Island
11
1640
17
16
1
1996-06-09
HFCB, NHMW
M. A. Jach
Greece
30
Plomari
Lesvos Island
11
1640
2
1
1
1964-05-30
NHMB
H. Korge
Turkey
Abant-Geb.
11
1640
1
1
Bolu
NHMW
U. Sahlberg
Turkey
37.933334
Ephesus
1178
27.316668
11
1640
1
1
1991-05-18
NHMW
S. Schodl
Turkey
700
Yamanlar Da
11
1640
2
1
1
1991-05-20
NHMW
M. A. Jach
Turkey
900
Berge
Tire
11
1640
2
1
1
Aydin
1968-06-13
1968-06-16
1968-06-13
NHMW
C. Holzschuh
Turkey
12
1641
40.483334
Ankara prov.
1162
32.633335
11
1640
1
1
Ankara
1968-06-15
GWCW, NHMB
G. Wewalka
Turkey
12
1641
2
1
1
1984-09-30
1984-10-02
1984-09-30
LHCB
L. Dobberstein
Turkey
Urgup
Kappadokien
12
1641
1
1
1985-04-04
HFCB
H. Fery
Turkey
700
Antalya
12
1641
1
1
Antalya
1991-05-25
NHMW
M. A. Jach
Turkey
50
Kemer
12
1641
1
1
Antalya
1997-07-01
NMPC
J. Hajek
Turkey
37.55
Buyuksofulu
1180
35.183334
12
1641
1
1
Adana
39.95
Karakoy village
1165
29.983334
13
1642
1
Bilecik