Hydroporus bodemeyeri Ganglbauer Bodemeyer, 1900: 139 Apfelbeck 1904: 374 Sahlberg 1913: 43 Bodemeyer 1927: 44 Zimmermann 1931: 9 H. tristis (Paykull, 1798)) Miller and Fery 1995: 409 Nilsson 2001: 159 Foster and Taylor 2006: 81 Darilmaz and Kiyak 2009: 1614 Hydroporus (s. str.) bodemeyeri : Zaitzev 1953: 155 Guéorguiev 1968: 34 Hydroporus ( Sternoporus ) bodemeyeri : Wewalka 1971: 211 Guéorguiev 1981: 407 Hydroporus ( Sternoporus ) collarti Guignot, 1949: 7 Wewalka 1971: 211 Taxonomic revision of the Hydroporus bodemeyeri species complex (Coleoptera: Dytiscidae) with a geometric morphometric analysis of body shape within the group Hájek, Jirí Fikácek, Martin Journal of Natural History 2010 2010-06-18 44 27 - 28 1631 1657 7JVWQ bodemeyeri Ganglbauer, 1900 bodemeyeri Ganglbauer 1900 [320,927,167,191] Insecta Dytiscidae Hydroporus Animalia Coleoptera 11 1640 Arthropoda subSpecies bodemeyeri bodemeyeri     Hydroporus bodemeyeriGanglbauerin  Bodemeyer, 1900: 139(original description);  Apfelbeck 1904: 374(notes);  Sahlberg 1913: 43(distribution);  Bodemeyer 1927: 44, 67 (distribution);  Zimmermann 1931: 9(as synonym of  H. tristis(Paykull, 1798));  Miller and Fery 1995: 409(description; partim);  Nilsson 2001: 159(catalogue); 2003: 60 (catalogue);  Foster and Taylor 2006: 81(distribution);  Darilmaz and Kiyak 2009: 1614(catalogue; partim).     Hydroporus(s. str.) bodemeyeri: Zaitzev 1953: 155(description);  Guéorguiev 1968: 34(description).     Hydroporus( Sternoporus) bodemeyeri: Wewalka 1971: 211(synonymy, distribution; partim.);  Guéorguiev 1981: 407(catalogue, distribution);     Hydroporus( Sternoporus) collarti Guignot, 1949: 7(original description) (synonymized by  Wewalka 1971: 211).    Typelocality. H. bodemeyeri: “Karakeuy” [ Turkey, Bilecik prov., Karaköy, ca. 39°57′N 29°59′E] (see Remarks under this species);  H. collarti: “Lesbos: Mytilini” [ Greece, Lesvos Isl., Mytilene, ca. 39°06′ N, 26°33′ E].  Type material. H. bodemeyeri:  Lectotypemale ( NHMW), designated by Wewalka (1971): “ Asia minor/ Karakeuy / v.Bodemeyer [printed] // Bodemeyeri / Gangl., Type [handwritten] // Coll. Mus. Vindob. [printed] // Hydroporus/ Bodemeyeri / Gnglb., Type. [handwritten] // Collect. / Hauser [printed] // HOLO- [sic!, handwritten] / TYPUS [red label, printed]”.  Paralectotypes( NHMW): two males, two females, same label data as lectotype.  H. collarti:  Holotypemale ( IRSNB): “ Coll. R. I. Sc. N. B./ Grèce/ I. Lesbos: Mytilini / N. W. 25-250., 4-V-’33 / A. d’Orchymont[printed] // Ƌ [printed] // Hydroporus/ ( Sternoporus) / Collarti Guign. [handwritten] // TYPE [printed, pink label with black frame]”.  Paratypes( IRSNB): one male, one female, “ Coll. R. I. Sc. N. B./ Grèce/ I. [ Island] Chios[ Khios]: Karyaes/ S. W.   140 m.,  2-V-1933/ A. d’Orchymont[printed] // Para- / type [red label, printed]”;  two females, “ Coll. R. I. Sc. N. B./ Grèce/ Samos: Platanos/ Velanidia  3-VI-1933/ A. d’Orchymont[printed] // Para- / type [red label, printed]”;  one male, three females, “ Coll. R. I. Sc. N. B./ Grèce/ I. [ Island] Ikaria: Xylocirti[ Xylosírtis] /   75 m.  25-V-1933/ A. d’Orchymont[printed] // Para- / type [red label, printed]”.  Additional material examined.  Greece: two males, Lésvos Island, Ayiásos-Megalokhóri, Mount Olympos,  55 m,  6 June 1996, M.A. Jächleg. ( NHMW);  one male, Lésvos Island, 10 kmnorth of Ayiásos, near Pigi Karini,  7 June 1996, M.A. Jächleg. ( NHMW);  nine males, 16 females, Lésvos Island, 4 kmsouth of Pétra, S Lafionas,  150 m,  8 June 1996, M.A. Jächleg. ( HFCB, NMPC, NHMW);  one male, two females, Lésvos Island, 2 kmwest of Plomari,  30 m,  9 June 1996, M.A. Jächleg. ( HFCB, NHMW).  Turkey: one female, Abant-Geb.[Abant Mts., Bolu prov.],  30 May 1964, H. Korgeleg. ( NHMB);  one male, Ephesus[near Selçuk, Izmir prov., ca. 37°56′ N, 27°19′ E]; U. Sahlbergleg. ( NHMW);  two males, one female, Izmir [prov.], Yamanlar Da g[mt.],  700 m,  18 May 1991, S. Schödlleg. ( NHMW);  four males, two females, Izmir [prov.], south of Tire, Aydin Berge,  900 m,  20 May 1991, M.A. Jächleg. (14) ( NHMW);  one female, K iz ilcahamam [ Ankara prov., ca. 40°29′ N, 32°38′ E],  13 June to 16 June 1968, C. Holzschuhleg. ( NHMW);  two males, one female, same label data, but  15 June 1968, G. Wewalkaleg. ( GWCW, NHMB);  one male, Kappadokien[Cappadocia, Nev sehir prov.], Ürgüp,  30 September to 2 October 1984, L. Dobbersteinleg. ( LHCB);  one female, north of Antalya[ Antalya prov.], ca.  700 m, stream,  4 April 1985, H. Feryleg. ( HFCB);  one male, Antalya prov., south of Kemer,  50 m,  25 May 1991, M.A. Jächleg. ( NHMW);  two females, Adana prov., Büyüksofulu[ca. 37°33′ N 35°11′ E], spring,  1 July 1997, J. Hájekleg. ( NMPC).  Measurements.Body length: 3.15–3.50 mm(mean value 3.36 mm), width: 1.45–1.60 mm(mean value 1.54 mm).  Description.Body flat, elongate, rather parallel sided ( Figure 3E), sides a little rounded. Elytra only weakly tapering to apex, with maximum width near midlength. Pronoto-elytral angle quite distinct. Upper surface blackish-brown, sides of pronotum and head often paler. Mouth appendages and legs testaceous. Ventral part blackish, head with paler gula. Head relatively broad, clypeus broadly rounded. Head on surface with two shallow depressions near anterior margin, and a groove along inner side of each eye. Punctation comprising sparse, almost regularly distributed punctures (denser in depressions), slightly smaller on clypeus and bigger on frons and vertex. Distance between punctures about twice their diameter. Microreticulation consisting of polygonal meshes with diameter smaller than that of punctures. Pronotum of rather variable shape. Usually with maximum width near midlength, sides more or less evenly curved over entire length; but in some specimens sides rather subparallel in posterior half. Lateral beading distinct, regular. Pronotum with a line of coarse setigerous punctures along anterior margin, and with shallow coarsely punctured posterolateral depressions. Disc of pronotum with sparse and fine setigerous punctures similar to those on head, but their diameter slightly smaller, and distance between punctures nearly three times their diameter. Microreticulation similar to that of head, but slightly less impressed. Elytra with sides almost straight behind shoulders, widest near midlength, then weakly tapering to apex; marginal rim visible at shoulders in dorsal view. Elytral disc with two rather indistinct punctured lines and with quite regularly distributed moderately large setigerous punctures between these lines. Distance between punctures about twice the diameter. Ventral part microreticulated. Reticulation indistinct on meso- and metaventrite, but stronger on metacoxal plates and abdominal ventrites. Metaventrite, metacoxal plates and two basal abdominal ventrites with large setigerous punctures. Other abdominal ventrites with sparse fine punctures. Male: dorsal surface shiny. Claws of fore tarsi broadened, strongly hooked. Median lobe of aedeagus regularly tapering to apex in ventral view, apical part relatively short, slightly turned to right ( Figure 2E). In lateral view, median lobe strongly inflexed in middle, apex long and nearly straight. Paramere as in Figure 2F. Female: pronotum slightly broader than in male. Microreticulation of elytra more distinct than in males and surface subopaque. Claws of fore legs thin, only slightly curved.  Remarks.The typelocality of  Hydroporus bodemeyeriwas stated as “In Mehrzahl bei Karakeuy gesammelt.” [Most specimens collected at Karakeuy (= Karaköy village).] (Ganglbauer in Bodemeyer 1900: 140). According to the Microsoft Encarta World Atlas 2000, there are about 50 different villages named “Karaköy” in Turkey. Miller and Fery (1995: 410)assumed the typelocality to be the village lying ca. 80 kmnorth of Adanaand 60 kmeast of Bulghar Dagh [= Bolkar Daglari Mountains]. We believe that the assumption is erroneous for two reasons. First, Bodemeyer (1900: 98)used the following abbreviation for the locality: “Ka. = Karakeuy, Station der anatolischen Bahn.” [station of the Anatolian railway]. There is no railway around the place cited by Miller and Fery (1995: 410). Second, Bodemeyer (1900: 24)presented precise information about Karaköy within the chapter “Reise von Konstantinopel nach Konia längs der anatolischen Bahn” [Travel from Constantinople (= Istanbul) to Konyaalong the Anatolian railway], and placed Karaköy between Bilecikand Eski sehir “...Steigt die Bahn von Biledjik aus schon steil zur Hochebene hinan, so hat man in Karakeuy 620 Meter, in Eski- Chéhir schon 800 Meter erreicht...” [...the railway runs up steeply from Bilecikto the plateau, and therefore it reaches 620 meters in Karaköy, and 800 meters already in Eski sehir...].  Therefore we assume that the typelocality of  H. bodemeyeriis the Karaköy villagein Bilecik Province, ca. 22 kmsouth of Bilecik, geographic co-ordinates 39°57′ N, 29°59′ E.  Distribution.The occurrence of the nominotypical subspecies is here confined to Greek Islands (Ikaría, Khios, Lésvos, Samos) and to the western half of Asian Turkey( Adana, Ankara, Antalya, Bolu, Izmir and Nev sehir provinces) ( Figure 6). NHMW Turkey Mus 11 1640 1 Mus lectotype NHMW Turkey Mus 11 1640 2 1 1 Mus paralectotype IRSNB Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & I. Lesbos & A. d'Orchymont Greece Grece 11 1640 1 holotype 1933-05-02 IRSNB Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & A. d'Orchymont Greece Karyaes 140 Chios Khios 11 1640 142 1 1 paratype 1933-06-03 IRSNB Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & A. d'Orchymont Greece Velanidia 140 true Samos Platanos 11 1640 1 1 paratype 1933-05-25 Coll. R. I. Sc. N. B. & Coll. R. I. Sc. & A. d'Orchymont Greece Xylosirtis 75 Ikaria Xylocirti 11 1640 77 1 1 paratype 1996-06-06 NHMW M. A. Jach Greece Lesvos Island 55 Mount Olympos Ayiasos-Megalokhori 11 1640 1 1 1996-06-07 NHMW M. A. Jach Greece Lesvos Island Pigi Karini Ayiasos 11 1640 1 1 1996-06-08 HFCB, NMPC, NHMW S Lafionas & M. A. Jach Greece 150 Petra Lesvos Island 11 1640 17 16 1 1996-06-09 HFCB, NHMW M. A. Jach Greece 30 Plomari Lesvos Island 11 1640 2 1 1 1964-05-30 NHMB H. Korge Turkey Abant-Geb. 11 1640 1 1 Bolu NHMW U. Sahlberg Turkey 37.933334 Ephesus 1178 27.316668 11 1640 1 1 1991-05-18 NHMW S. Schodl Turkey 700 Yamanlar Da 11 1640 2 1 1 1991-05-20 NHMW M. A. Jach Turkey 900 Berge Tire 11 1640 2 1 1 Aydin 1968-06-13 1968-06-16 1968-06-13 NHMW C. Holzschuh Turkey 12 1641 40.483334 Ankara prov. 1162 32.633335 11 1640 1 1 Ankara 1968-06-15 GWCW, NHMB G. Wewalka Turkey 12 1641 2 1 1 1984-09-30 1984-10-02 1984-09-30 LHCB L. Dobberstein Turkey Urgup Kappadokien 12 1641 1 1 1985-04-04 HFCB H. Fery Turkey 700 Antalya 12 1641 1 1 Antalya 1991-05-25 NHMW M. A. Jach Turkey 50 Kemer 12 1641 1 1 Antalya 1997-07-01 NMPC J. Hajek Turkey 37.55 Buyuksofulu 1180 35.183334 12 1641 1 1 Adana 39.95 Karakoy village 1165 29.983334 13 1642 1 Bilecik