Hydroporus guignoti Gschwendtner, 1935: 205 Hydroporus ( Hydroporinus ) guignoti : Zaitzev 1953: 177 Hydroporus ( Heterosternus ) bulgaricus Hlisnikovský, 1955: 100 Hydroporus ( Sternoporus ) bodemeyeri : Guéorguiev 1965: 102 Wewalka 1971: 211 Guéorguiev 1987: 74 Hydroporus bodemeyeri : Miller and Fery 1995: 409 Taxonomic revision of the Hydroporus bodemeyeri species complex (Coleoptera: Dytiscidae) with a geometric morphometric analysis of body shape within the group Hájek, Jirí Fikácek, Martin Journal of Natural History 2010 2010-06-18 44 27 - 28 1631 1657 5HZWW Gschwendtner, 1935 Gschwendtner 1935 [338,938,1271,1295] Insecta Dytiscidae Hydroporus Animalia Coleoptera 14 1643 Arthropoda subSpecies bodemeyeri stat. nov. guignoti      Hydroporus guignoti Gschwendtner, 1935: 205(original description).     Hydroporus( Hydroporinus) guignoti: Zaitzev 1953: 177(description).     Hydroporus( Heterosternus) bulgaricus Hlisnikovský, 1955: 100(original description) syn. revid.     Hydroporus( Sternoporus) bodemeyeri: Guéorguiev 1965: 102(notes);  Wewalka 1971: 211(distribution; partim); 1974: 106 (synonymy);  Guéorguiev 1987: 74(description).     Hydroporus bodemeyeri: Miller and Fery 1995: 409(partim).    Typelocality. H. guignoti: “ Albanien( Tirana)” [ Albania, ca. 7 kmnorth-east of Tirana, Mal i Dajtit, ca. 41°21′ N, 19°55′ E];  H. bulgaricus: “Paril, Alibotuš planina,   Bulgariaoccid.” [nowadays called Slavyanka Mountains, ca. 20 kmsouth-west of Gotse Delchev, ca. 41°25′ N, 23°37′ E].  Type material. H. guignoti:  Holotypefemale [sic!] ( OLML): “MAL-I-DAJTIT/ TIRANE[handwritten] // ALBANIA/ LEG. BISCHOFF / 1931 [handwritten] // Type Ƌ / Gsch. [= Gschwendtner] [handwritten, pink label] // Coll. / Gschwendtner [printed] // Hydroporus/ Guignoti Gsch [handwritten] / det. Gschwendt.[printed] // Type [handwritten, red label] // Hydroporus/ guignoti Gschw./ = bodemeyeri Ganglb.[handwritten] / det. Wewalka70 [printed]”.  Paratypes( OLML): one female, same label data as holotype, but “ Paratype” instead of “Type”; one female, “Parnass / Graecia [printed] // Type ♀/ Gsch. [handwritten, pink label] // Coll. / Gschwendtner [printed] // Hydr. ♀II / Guignoti Gsch. [handwritten] / det. Gschwendt.[printed] // Type [handwritten, red label] // Hydroporus/ guignoti Gschw./ = bodemeyeri Ganglb.[handwritten] / det. Wewalka70 [printed]”; one male, one female, same label data as holotype, but “ Paratype” instead of “Type”.  H. bulgaricus:  Holotypefemale [sic!] ( NMPC): “Alibotuš pl. [planina, = mountains] / Bulg. [Bulgaria], 15.6.[19]38 / Hlisnikowski [printed, date handwritten] // Type [printed, red label with black frame] // Coll. Hlisnikovský/ Mus. Nat. Pragae [printed] // HYDROPORUS/ BULHARICUS[sic!] HLIS. [handwritten] / det. Hlisnikovský1952 [printed]” ( NMPC).  Additional material examined.  Albania: one female, Borske, 1940, Bischoffleg. ( OLML).  Bulgaria: five males, two females, Blagoevgrad distr., Sandanskienv., valley of Lebnica river, stream,  26 May 1998, J. Hájekleg. ( HFCB, NMPC);  one male, Kurdzhali distr., Kaloyantsienv., stream,  28 June 1999, J. Hájekleg. ( NMPC);  three males, one female, Smolyan distr., 3 kmwest Smolyan, Gerzovicastream valley,  4 July 1999, J. Hájekleg. ( HFCB, NMPC).  Greece: one male, two females, Aitoloakarnania distr., 30 kmsouth of Agrénion,  4 May 1971, G. Wewalkaleg. ( HFCB, NMPC);  four males, one female, Levkás Island,  11 September 1988, M.A. Jächleg. ( NHMW);  one male, Thásos Island[Kavala distr.], Megálos Prínos,  11 September 1991, M.A. Jächleg. ( NHMW);  two males, one female, Thásos Isl., N Theológos,  12.ix.1991, M.A. Jächleg. ( NHMW);  1 female, Thásos Island, Potamiás,  13 September 1991, M.A. Jächleg. ( NHMW);  three females, Euboea Island[Evvoia distr.], Steni-Stroponas,  700-900 m,  30 September 1992, M.A. Jächleg. ( NHMW);  three females, Samothráki Island, [Évros distr.], 1 kmwest of Kremniotsa,  15 June 1993, M.A. Jächleg. ( NHMW);  one male, one female, Khalkidhikí, Athos[ Ágion ÓrosPeninsula], 2 kmnorth-west of Megistis Lavras, ca.  30 m,  3 July 2002, M.A. Jächleg. ( NHMW);  four males, six females, Xanthi distr., ca. 10 kmeast of Stavroúpolis, 41°12.4′ N, 24°47.7′ E, ca.  360 m, stream,  25–26 April 2007, J. Hájekleg. ( NMPC).  Measurements.Body length: 3.10–3.40 mm(mean value = 3.27 mm), width: 1.40–1.55 mm(mean 1.48 mm).  Description.Similar to nominotypical subspecies in general habitus. Usually slightly smaller; sides of elytra more curved, so pronoto-elytral angle more distinct than in nominotypical subspecies ( Figure 3D). Punctation on pronotum and elytra coarser and deeper. Distance between punctures about that of their diameter, in particular in basal half of elytra. Male: dorsal surface shiny. Claws of fore tarsi broadened, strongly hooked. Median lobe of aedeagus in ventral view ( Figure 2H) more intensively tapering in distal half, apical part longer than in nominotypical subspecies, distinctly turned to right, apex very acute. Female: microreticulation of elytra usually more distinct than in males so surface subopaque. Claws of fore legs thin, only slightly curved.  Habitat.Collected in gravel and under submerged leaves in small temporary streams.  Distribution.A subspecies which occurs in the Balkan Peninsula. Known from Albania, southern Bulgariaand Greece, including the islands of Euboea, Levkás, Samothráki and Thásos ( Figure 6). Bulgaria 41.416668 Slavyanka Mountains 1156 23.616667 Gotse Delchev 15 1644 1 OLML Albania Gschwendt. & Wewalka TIRANE 15 1644 1 Tirane holotype [661,887,423,448] OLML Albania 15 1644 1 Tirane paratype OLML Albania Gschwendt. & Wewalka TIRANE 15 1644 1 Tirane paratype Albania Gschwendt. & Wewalka 15 1644 2 2 Tirane paratype OLML Albania Gschwendt. & Wewalka TIRANE 15 1644 1 Tirane paratype NMPC Coll. Hlisnikovsky Albania Hlisnikovsky BULHARICUS 15 1644 1 BULHARICUS holotype OLML Bischoff Albania Borske 15 1644 1 1 1998-05-26 HFCB, NMPC J. Hajek Bulgaria Lebnica river Sandanski 15 1644 2 1 1 Blagoevgrad 1999-06-28 NMPC J. Hajek Bulgaria Kaloyantsi 15 1644 1 1 Kurdzhali 1999-07-04 HFCB, NMPC J. Hajek Bulgaria Gerzovica 15 1644 2 1 1 Smolyan 1971-05-04 HFCB, NMPC G. Wewalka Greece Agrenion Aitoloakarnania distr. 15 1644 2 1 1 1988-09-11 NHMW M. A. Jach Greece Levkas Island 15 1644 2 1 1 1991-09-11 NHMW M. A. Jach Greece Megalos Prinos Thasos Island 15 1644 1 1 1991-09-12 NHMW N Theologos & M. A. Jach Greece Thasos Isl. 15 1644 2 1 1 1991-09-13 NHMW M. A. Jach Greece Potamias Thasos Island 15 1644 1 1 1992-09-30 NHMW M. A. Jach Greece 800 Steni-Stroponas Euboea Island 15 1644 1 1 1993-06-15 NHMW M. A. Jach Greece Kremniotsa Samothraki Island 15 1644 1 1 2002-07-03 NHMW M. A. Jach Greece 30 Megistis Lavras Khalkidhiki 15 1644 2 1 1 Agion Oros 2007-04-25 2007-04-26 2007-04-25 NMPC J. Hajek Greece 360 41.206665 Stavroupolis 115 24.795 Xanthi distr. 15 1644 2 1 1