Noideattella and Tolegnaro, Two New Genera of Goblin Spiders from Madagascar, with Comments on the Gamasomorphoid and Silhouettelloid Oonopids (Araneae, Oonopidae) Álvarez-Padilla, Fernando Ubick, Darrell Griswold, Charles E. American Museum Novitates 2012 2012-05-25 2012 3745 1 76 73Z6X Álvarez-Padilla & Ubick & Griswold, 2012 Álvarez-Padilla & Ubick & Griswold 2012 [476,714,435,462] Arachnida Oonopidae Noideattella Animalia Araneae 52 53 Arthropoda species lakana sp. nov.    Figures 257–284, 375   TYPES: Male holotype and femaleallotype from Berlese and sifted leaf litter of a tropical dry forestat an elevation of  325 mat Reserve Speciale d’Ambre,  3.5 km235° SW Sakaramy, 12°28′8″S, 49°14′32″E, Antsiranana Province,  26–31 January 2001, Fisher-Griswold Arthropod Team, deposited in CASC( PBI_OON 00003275). The species epithet, a noun in apposition, is taken from the Malagasy word for a dugout canoeor any small boat .  DIAGNOSIS:  N. lakanacan be differentiated from other  Noideattellaspeciesby the following combination of characters: habitus color dark red-brown, body length 2.1 to 2.4 mm; carapace surface of elevated portion of pars cephalica strongly granulate, sides granulate (figs. 260, 275, 276); sternum furrows with row of large pits, surface coarsely reticulate, with microsculpture medially and in furrows (fig. 277); book lung covers large and elliptical (figs. 258, 259).  DESCRIPTION: Male(PBI_OON 0003462): Total length 2.1. Cephalothorax:Carapace dark red-brown, sternum longer than wide, dark red-brown, anterior margin with continuous transverse groove ( fig. 273). Mouthparts: Chelicerae, endites, and labium dark red-brown, paturon inner margin with scattered setae (fig. 263). Abdomen:Ovoid. Book lung covers large, elliptical. Dorsal scutum dark red-brown, middle surface smooth, sides smooth. Postepigastric scutum orange-brown, long, semicircular. Male genital opening slightly protruded, epigastric scutum apodemes small, separated from the anterior book lung opening and connected by a furrow. Posterior spiracles connected by a groove ( fig. 268). Legs:Orange-brown, without color pattern. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I, II p3-1-1; r2-1-1; IV p0-0-1; metatarsi: I, II p1-1-0; r1-1-0. Tarsal organ with two sensilla visible. Genitalia:Epigastric region with sperm pore situated at level of anterior spiracles ( fig. 268). Palp proximal segments pale orange; cymbium pale orange, with distal patch of setae; bulb pale orange (figs. 262, 264). Embolus dorsal and ventral lamellae with at least three distal apophyses each (figs. 281–284). Palp as in figures 279 and 280. Female(PBI_ OON 0003462): Total length 2.4. As in male except as noted. Legs:Metatarsi III and IV weak ventral scopula absent. Leg spination (only surfaces bearing spines listed, all spines longer than segment width): tibiae: I, II p3-1-1; r3-1-1; IV p0-0-1; metatarsi: I, II p1-1-0; r1-1-0. Trichobothria of metatarsus: I and IV one. Genitalia:Genital opening slit shaped ( fig. 267). Sperm sac secretory glands openings flat, extending all over the receptaculum posterior surface. GAp process shaped as a twisted tube ( figs. 269–271). Genitalia as in figures 265 and 266.  MATERIAL EXAMINED: N= 86.   Madagascar:  Antananarivo: 3 km41° NE Andranomay, 11.5 km147° SSE Anjozorobe, montane rainforest, 1300 m, S18.47333°, E47.96000°, Dec. 05–13, 2000, Fisher-Griswold Arthro- pod Team, 1♀(  CASCPBI_  OON3941).   Antsiranana:  7.2 km142° SE Antsiranana(= Diego Suarez), Montagne des Francais,  180 m, S12.53796°, E49.56361°,  Feb. 23–25, 2001, L.J. Boutin, 1♀(  CASCPBI_  OON3921);  Forêt de Binara,  9.4 km235° SW Daraina, montane rainforest,  1100 m, S13.26333°, E49.60000°,  Dec. 05, 2003, B.L. Fisher, 5 mixed (  CASCPBI_  OON3153);  P.N. Montagne d’Ambre, 2.79 air km NE of park entrance forest,  1000 m, S12.53333°, E49.16666°,  Nov. 21–30, 1993, J. Coddington, S. Larcher, C. Griswold, R. Andriamasimanana, N. Scharff, 1♀(  CASCPBI_  OON3460); 12 mixed ( ZMUC PBI_  OON3462); 5 mixed (  CASCPBI_  OON3463); 5 mixed (  CASCPBI_  OON3465);  2♂(  CASCPBI_  OON3468);  2♀(  CASCPBI_  OON3473);  1♂(  CASCPBI_  OON3474);  1♂(  CASCPBI_  OON3475); 15 mixed (  CASCPBI_  OON3477); 2 mixed (  CASCPBI_ OON36378); 5 mixed (  CASCPBI_ OON36380); 13 mixed (  CASCPBI_ OON36389); 6 mixed (  CASCPBI_ OON36392);  Parc National Montagne d’Ambre,  3.6 km235° SW Joffreville, montane rainforest,  925 m, S12.53444°, E49.17944°,  Jan. 20–26, 2001, Fisher-Griswold Arthropod Team, 4 mixed (  CASCPBI_ OON36376);  Reserve Speciale d’Ambre,  3.5 km235° SW Sakaramy, tropical dry forest,  325 m, S12.46888°, E49.24222°,  Jan. 26–31, 2001, Fisher-Griswold Arthropod Team, 1♀(  CASCPBI_  OON3359);  Reserve Speciale de l’Ankarana,  22.9 km224° SW Anivorano Nord, tropical dry forest,  80 m, S12.90888°, E49.10972°,  Feb. 10–16, 2001, Fisher-Griswold Arthropod Team, 1♂(  CASCPBI_  OON3172);  Reserve Speciale l’Ankarana,  13.6 km192° SSW Anivorano Nord, tropical dry forest,  210 m, S12.86361°, E49.22583°,  Feb. 16–20, 2001, Fisher-Griswold Arthropod Team, 3 mixed (  CASCPBI_  OON3721). 3443623516 [168,426,548,574] Berlese and sifted leaf litter of a tropical dry forest 2001-01-26 2001-01-31 2001-01-26 CASC 325 -12.468888 Reserve Speciale d'Ambre 21 49.24222 Berlese 52 53 PBI_OON 00003275 1 1 Antsiranana Province holotype 3443623603 Berlese and sifted leaf litter of a tropical dry forest 2001-01-26 2001-01-31 2001-01-26 CASC 325 -12.468888 Reserve Speciale d'Ambre 21 49.24222 Berlese 52 53 PBI_OON 00003275 1 1 Antsiranana Province allotype 3443623652 CASC, PBI 56 57 Team 52 53 OON 3941 1 1 3443623574 2001-02-23 2001-02-25 2001-02-23 CASC, PBI L. J. Boutin 180 -12.53796 Montagne des Francais 1 49.56361 Diego Suarez 56 57 OON 3921 1 1 Antsiranana 3443623458 2003-12-05 CASC, PBI B. L. Fisher 1100 -13.26333 Foret de Binara 1 49.6 56 57 OON 3153 1 Antsiranana 3443623387 1993-11-21 1993-11-30 1993-11-21 CASC, PBI P. N. Montagne d'Ambre & J. Coddington & S. Larcher & Griswold & Andriamasimanana & Scharff 1000 -12.53333 1 49.16666 56 57 OON 3460 1 1 Antsiranana 3443623644 [631,995,371,395] ZMUC, PBI 56 57 OON 3462 1 Antsiranana 3443623319 CASC, PBI 56 57 OON 3463 1 Antsiranana 3443623355 [256,603,400,424] CASC, PBI 56 57 OON 3465 1 Antsiranana 3443623577 [615,912,400,424] CASC, PBI 56 57 OON 3468 2 2 Antsiranana 3443623480 [924,1220,400,424] CASC, PBI 56 57 OON 3473 2 2 Antsiranana 3443623553 [120,406,430,454] CASC, PBI 56 57 OON 3474 1 1 Antsiranana 3443623486 [415,702,429,454] CASC, PBI 56 57 OON 3475 1 1 Antsiranana 3443623507 [711,1057,429,454] CASC, PBI 56 57 OON 3477 1 Antsiranana 3443623383 OON 36378 CASC, PBI 56 57 1 Antsiranana 3443623450 OON 36380 [317,673,459,483] CASC, PBI 56 57 1 Antsiranana 3443623555 OON 36389 [683,1051,459,483] CASC, PBI 56 57 1 Antsiranana 3443623373 OON 36392 CASC, PBI 56 57 1 Antsiranana 3443623576 OON 36376 2001-01-20 2001-01-26 2001-01-20 CASC, PBI Fisher-Griswold Arthropod Team 925 -12.53444 Parc National Montagne d'Ambre 1 49.17944 56 57 1 Antsiranana 3443623405 2001-01-26 2001-01-31 2001-01-26 CASC, PBI Fisher-Griswold Arthropod Team 325 -12.46888 Reserve Speciale d'Ambre 1 49.24222 56 57 OON 3359 1 1 Antsiranana 3443623566 2001-02-10 2001-02-16 2001-02-10 CASC, PBI Fisher-Griswold Arthropod Team 80 -12.90888 Reserve Speciale de l'Ankarana 1 49.10972 56 57 OON 3172 1 1 Antsiranana 3443623393 2001-02-16 2001-02-20 2001-02-16 CASC, PBI Fisher-Griswold Arthropod Team 210 -12.86361 Reserve Speciale l'Ankarana 1 49.22583 56 57 OON 3721 1 Antsiranana