A review of the genus Cyana Walker, 1854 from India, with descriptions of five new species and three new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini)
Singh, Navneet
Volynkin, Anton V.
Kirti, Jagbir Singh
Datta, Harvinder Singh
Ivanova, Maria S.
Zootaxa
2020
2020-02-18
4738
1
1
93
Volynkin & N. Singh
Volynkin & N. Singh
2020
[151,686,153,180]
Insecta
Erebidae
Cyana
GBIF
Animalia
Lepidoptera
37
38
Arthropoda
subSpecies
dohertyi
subsp. nov.
eirene
( Figs 102, 103, 205, 269)
Type material. Holotype( Figs 102, 205): ♂, “ Indien, [ Uttarakhand] U. P., Kumaon-Himalaya, Bhimtal, 1500 m, 12.IV.1976, leg. F. Smetaсek”, slide MWM 31804 Volynkin (Coll. MWM/ ZSM). Paratypes: INDIA. HIMACHAL PRADESH: 1 ♂, N.W. Himalaya( Coll. NHMUK); 1 ♂, India, H.P., Kan- daghat, 1500m, 9–10.VI.1975, leg. W. Thomas, slide MWM 35885 ( ♂) Volynkin( Coll. MWM/ ZSM); UTTARA- KHAND: 6 ♂, Indiasept., Kumaon-Himalaya, Distr. NainiTal, BhimTal, 1500 m, leg. F. Smetacek, 10.VII.1979( 1 ♂); 18.IX.1979( 2 ♂); 22.IX.1979( 1 ♂); 24.IX.1979( 2 ♂) ( Coll. MWM/ ZSM); 2 ♂, 3 ♀, Indiasept., Kumaon- Himalaya, Distr. NainiTal, BhimTal, 1500 m, Smetacek/ Bauer, 31.VIII.1976( 1 ♂, 1 ♀), slide MWM 35881 ( ♂) Volynkin; 1.IX.1976( 1 ♂, 1 ♀), slide MWM 35880 ( ♂) Volynkin; 11.IX.1976( 1 ♀) ( Coll. MWM/ ZSM); 3 ♂, 1 ♀, India, Kumaon-Himalaya, Bhimtal, 1500 m, leg. F. Smetaсek 3.V.1975( 1 ♂); 12.VI.1976( 1 ♂), slide MWM 31804 ( ♂) Volynkim; 5.IX.1976( 1 ♂); 24.IX.1975( 1 ♀); 1.X.1977( 1 ♂) ( Coll. MWM/ ZSM); 1 ♂, India, Bhim- tal, 1500 m, 11–20.VI.1975, leg. W. Thomas( Coll. MWM/ ZSM); 1 ♂, India, Kumaon-Himalaya, Distr. NainiTal, BhimTal, 20–23.V.1981, 1500 m, leg. C. Holzschuh( Coll. MWM/ ZSM); 2 ♂, 1 ♀, N-India, Bhimtal, 1500 m, 14–30.VII.1979, coll. Dr. Liedgens, slides MWM 31802 ( ♂), MWM 35884 ( ♀) Volynkin( Coll. MWM/ ZSM); 11 ♂, 2 ♀, India, Kumaon-Himalaya, Distr. NainiTal, BhimTal, 1500 m, leg. de Freina, Himalaya Expedition, 16.V.1971( 1 ♂); 26.V.1971( 2 ♂); 10.V.1971( 4 ♂, 1 ♀); 10.VI.1971( 1 ♂), slide MWM 35883 ( ♂) Volynkin; 12.VI.1971( 2 ♂); 18.VI.1971( 1 ♂, 1 ♀) ( Coll. MWM/ ZSM); 1 ♂, India, Kumaon-Himalaya, Distr. NainiTal, BhimTal, 1500 m, 21.VI.1981, leg. Smetaсek ( Coll. MWM/ ZSM); 3 ♂, Kumaon-Himalaya, Nainital-distr., Bhim- tal, 1500 m, leg. F. Smetaсek, 30.V.1978( 1 ♂), 1.VII.1978( 2 ♂) ( Coll. MWM/ ZSM); 1 ♂, 5 ♀, Bhimtal, N.- India, 1600 m, 24.VIII.–3.IX.80, Hauenstein, slide MWM 35882 ( ♀) Volynkin( Coll. MWM/ ZSM ) . PAKISTAN. AZAD JAMMU & KASHMIR: 1 ♂, Pakistan, Kashmir, Himalaya Mts., 20 kmВ Muzaffarabad, Nara village, 750m, 73°29’E 34°01’N, 20.V.1998, leg. Gy.M. László& G. Ronkay, coll. Museum Witt, slide MWM 31801 ( ♂) Volynkin( Coll. MWM/ ZSM); 8 ♂, 3 ♀, Pakistan, Kashmir, Himalaya Mts., 30 kmN Murree, near Nathia Ghali, Ayubiavil- lage, 2600m, 23.V.1998, leg. Gy.M. László& G. Ronkay, coll. Museum Witt, slides ZSM Arct.71/2017 ♂, ZSM Arct.72/2017 ♀ Volynkin( Coll. MWM/ ZSM).
Etymology. In the Greek mythology, Eirene was one of the Horae, the personification of peace.
Diagnosis. Forewing length is 15–16.5 mmin males and 17–18 mmin females. Cyana dohertyi eirenediffers from the nominate subspecies and C. d. mertsana(illustrated by Volynkin et al. (2019)) by its narrower and paler yellow transverse lines. The male and female genitalia of the new subspecies have no significant differences from those of the nominate subspecies. Compared to those of C. d. mertsana(illustrated by Volynkin et al. (2019)), the male genitalia of C. d. eirenehave the slightly narrower sacculus with slightly weaker distal process, and the slightly shorter band-like cluster of spinules on the largest diverticulum of vesica. The female genitalia of C. d. eirenediffer from those of C. d. mertsana(illustrated by Volynkin et al. (2019)) by the slightly shorter sclerotized posterior section of cospus bursae.
Distribution. Western Himalaya: Pakistan(Azad Jammu & Kashmir) and North India( Himachal Pradeshand Uttarakhand) ( Smetacek 2008, as C. dohertyi).
2565941157
1976-04-12
MWM, ZSM
India
1500
Bhimtal
Kumaon-Himalaya
37
38
1
1
Uttarakhand
holotype
2565941253
[199,1178,333,360]
NHMUK
India
Coll.
Himalaya
37
38
1
1
Himachal Pradesh
paratype
2565941350
1975-06-09
1975-06-10
1975-06-09
MWM, ZSM
India, H. P. & W. Thomas
India
1500
Coll.
Volynkin
37
38
2
2
HIMACHAL PRADESH
paratype
2565941440
1979-07-10
1979-09-24
1979-07-10
MWM, ZSM
F. Smetacek
India
India
1500
Distr. NainiTal
Kumaon-Himalaya
37
38
12
12
KHAND
paratype
2565941281
1976-08-31
1976-09-11
1976-08-31
MWM, ZSM
Smetacek & Bauer & Volynkin & Coll.
India
India
1500
Distr. NainiTal
Himalaya
37
38
12
6
6
KHAND
paratype
2565941381
1975-05-03
1977-10-01
1975-05-03
MWM, ZSM
India
India
1500
Bhimtal
Kumaon-Himalaya
37
38
10
2
8
KHAND
paratype
2565941294
1975-06-11
1975-06-20
1975-06-11
MWM, ZSM
W. Thomas & Coll.
India
1500
India
37
38
1
1
KHAND
paratype
2565941151
1981-05-20
1981-05-23
1981-05-20
MWM, ZSM
C. Holzschuh & Coll.
India
India
1500
Distr. NainiTal
Kumaon-Himalaya
37
38
1
1
KHAND
paratype
2565941378
1979-07-14
1979-07-30
1979-07-14
MWM, ZSM
Dr. Liedgens
India
Bhimtal
1500
Coll.
Volynkin
37
38
5
2
3
KHAND
paratype
2565941254
1971-05-10
1971-06-18
1971-05-10
MWM, ZSM
de Freina & Himalaya Expedition
India
India
1500
Distr. NainiTal
Kumaon-Himalaya
37
38
27
4
23
KHAND
paratype
2565941376
1981-06-21
MWM, ZSM
Coll.
India
India
1500
Distr. NainiTal
Kumaon-Himalaya
37
38
1
1
KHAND
paratype
2565941313
1978-05-30
1978-07-01
1978-05-30
MWM, ZSM
India
Kumaon-Himalaya
1500
Coll.
Nainital-distr.
37
38
6
6
KHAND
paratype
2565941127
MWM, ZSM
Hauenstein
India
Bhimtal
1600
Volynkin
India
37
38
7
6
1
KHAND
paratype
2565941342
1998-05-20
MWM, ZSM
Gy. M. Laszlo & G. Ronkay & Museum Witt
Pakistan
Pakistan
750
34.016666
Himalaya Mts.
1202
73.48333
Kashmir
37
38
2
2
KASHMIR
paratype
2565941249
1998-05-23
Gy. M. Laszlo & G. Ronkay & Museum Witt & Arct.
Pakistan
Pakistan
2600
Himalaya Mts.
Kashmir
37
38
11
3
8
KASHMIR
paratype
2565941315
Pakistan
Arct.
37
38
1
1
KASHMIR
paratype
2565941211
[314,672,1089,1116]
MWM, ZSM
Pakistan
Coll.
Volynkin
37
38
1
1
KASHMIR
paratype