Harold, 1873 : 103 Parachorius thomsoni Gillet 1911 : 54 Boucomont 1929 : 761 Arrow 1931 : 358 Paulian 1945 : 59 Balthasar 1963 : 276 Parachorius thomsoni Kryzhanovsky & Medvedev 1966 : 394 Parachorius thomsoni Masumoto 1987 : 127 Hanboonsong & Masumoto, 2001 : 138 A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data Tarasov, Sergei Zootaxa 2017 2017-10-04 4329 2 101 149 4CPV4 Harold, 1873 Harold 1873 [151,590,151,178] Insecta Scarabaeidae Parachorius Animalia Coleoptera 41 142 Arthropoda species thomsoni     Parachorius thomsoni  Harold, 1873: 103    Parachorius thomsoni;  Gillet 1911: 54     Parachorius thomsoni;  Boucomont 1929: 761    Parachorius thomsoni;  Arrow 1931: 358    Parachorius thomsoni;  Paulian 1945: 59    Parachorius thomsoni;  Balthasar 1963: 276     Parachorius thomsoni;  Kryzhanovsky & Medvedev 1966: 394  Parachorius thomsoni;  Masumoto 1987: 127    Parachorius lannathai  Hanboonsong & Masumoto, 2001: 138( new synonymy)     Typelocality. Indiaor. [ eastern India] (  Parachorius thomsoni); Maesa Valley Chiang Mai, Thailand (  Parachorius lannathai)  Taxonomic notes.  Parachorius lannathaidescribed from Thailandis synonymized with  P. thomsonibased on external morphology and the shape of aedeagus.   Distribution.Eastern India, Myanmar, Thailand, and Laos.   Diagnosis.The species is most similar to  P. fungorumand  P. hookeri. It differs from them by the following combination of characters (1) metafemoral posterior margin with two distinct denticles of which the apical denticle is spur-like, produced; (2) parameres simple, apex slightly rounded forming triangle.   Description. Body.Body black or dark brown, length 6.5–9.0 mm. Elytra uniformly colored.  Head.Clypeus with two indistinct teeth, outer margin of each tooth not notched basally.  Pronotum.Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–2 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.   FIGURE 24.Morphology of  Parachorius thomsoni. (A) aedeagus, lateral view (  P. thomsoniholotype, male) (B) aedeagus, lateral view (  P. lannathai, holotype, male). (C–D) aedeagus, lateral view. (E) left metafemur (  P. lannathai, holotype, male). (F) disc of left elytron (G) habitus (  P. lannathai, holotype, male). (H) habitus (  P. thomsoni, holotype, male).   FIGURE 25.Distribution maps of  Parachoriusspecies. (A)  Parachorius asymmetricus.(B)  Parachorius bolavensis.(C)  Parachorius fukiensis.(D)  Parachorius fungorum.(E)  Parachorius globosus.(F)  Parachorius gotoi. Type locality—type locality of species (holotype or lectotype); Type. Collection data—localities of type specimens associated with species (paratypes or paralectotypes); Collection data—localities of other specimens examined in this study.   FIGURE 26.Distribution maps of  Parachoriusspecies. (A)  Parachorius hookeri.(B)  Parachorius humeralis.(C)  Parachorius javanus.(D)  Parachorius longipenis.(E)  Parachorius maruyamai.(F)  Parachorius newthayerae. Type locality— type locality of species (holotype or lectotype); Type. Collection data—localities of type specimens associated with species (paratypes or paralectotypes); Collection data—localities of other specimens examined in this study.   FIGURE 27.Distribution maps of  Parachoriusspecies. (A)  Parachorius peninsularis.(B)  Parachorius pseudojavanus.(C)  Parachorius schuelkei.(D)  Parachorius semsanganus.(E)  Parachorius solodovnikovi.(F)  Parachorius thomsoni. Type locality—type locality of species (holotype or lectotype); Type. Collection data—localities of type specimens associated with species (paratypes or paralectotypes); Collection data—localities of other specimens examined in this study. Literature data— localities retrieved from literature.  Elytra.Elytra covered with sparse punctures. Wings.Wings normally developed.  Legs.Profemur with anterior margin medially produced in angle or with almost straight or sinuate anterior margin. Protibia with apical spur approximately reaching middle of protarsomere 3. First tooth almost not modified in males, approximately as wide as second tooth; first and second teeth distinctly separated from each other; first distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin not serrate, with two distinct denticles of which apical denticle is spur-like, produced. Metatibia slightly sinuate, inner margin not denticulate, excavated apically.  Metasternum.Transverse basal ridge of metasternum long, at least 1/5 of mesocoxa length.  Aedeagus.Parameres symmetrical, simple; apex slightly rounded, forming triangle.   Material examined. Holotype of  Parachorius thomsoni: India: 1 male, “India or. Parachorius Thomsoni Typ. Harold./ Ex Musaeo E.Harold / MUSEUM PARIS 1952 Coll. R. OBERTHÜR / TYPE / HOLOTYPE Parachorius thomsoni Horld, 1873det. S. Tarasov 2010” (MNHN). Holotype of  Parachorius lannathai: Thailand: 1 male, "Maesa Valley Chiang Mai THAILAND 3-6/V/1994K. MASUMOTO leg. / Coll Masumoto 200 / Holotype Parachorius lannathai Han. & Mas./ Parachorius thomsoni Harold, 1873det. S. Tarasov, 2015", 18°53.988'N, 98°52.524'E (NSMT). 1 male, collection J. Thomson 830 (ISNB). India: 1 male, India orientale (BMNH). Laos: 1 male, Khammouane Province, Ban Khun Kham (Nahin), 18°13.027'N, 104°30.88'E, 300 m, disturbed rainforest, on 3–5/vi/2008(A. Solodovnikov, J. Pedersen) (ZMUC). Myanmar: 1 male, Shan State, Shweudaung Wildlife Sanctuary (Moe Meik Township), Kyauk-maw village (91f), flight intercept trap, 23°5.129'N, 96°13.527'E, 360 m, on 1–15/viii/2002(Myint Hlaing, Aung Moe) (NMW). 1658411537 [151,1168,570,595] India eastern India 41 142 1 Maesa Valley Chiang Mai holotype