Tricorythus australis Banks, 1913 : 85 Leptohyphodes australis New species of Macunahyphes Dias, Salles & Molineri (Ephemeroptera: Leptohyphidae), with taxonomic notes Paula Malaquias Souto Frederico Falcão Salles European Journal of Taxonomy 2016 254 1 15 6R3WJ Banks, 1913 Banks 1913 [559,1029,267,294] Insecta Leptohyphidae Macunahyphes Animalia Ephemeroptera 3 4 Arthropoda species australis   Fig. 2C    Tricorythus australis Banks, 1913: 85.   Leptohyphodes australis– Ulmer 1920: 50.    Tricorythodes australis– Traver 1958: 501–503. — Hubbard 1982: 274. — Molineri 2002: 278.  Macunahyphes australis– Dias, Salles & Molineri 2005: 199.    Material examined   BRAZIL: Espírito Santo: 113 ♁♁, imagines, Colatina, Itapina, Doce River, 19°31′18.1″ S, 40°50′11.1″ W,  7–8 Sep. 2014, E.M. Rozárioand F.F. Sallesleg. ( CZNCEp-6440);  14 ♀♀, imagines, same data ( CZNCEp-6441); 72 ♁♁, imagines, same data ( CZNCEp-6443); 73 ♁♁, imagines, same data ( CZNCEp- 6444); 64 ♁♁, imagines, same data ( CZNCEp-6445); 100 ♁♁, imagines, same data ( CZNCEp-6447); 3 ♁♁, imagines, Marilândia, Doce River, 19°31′5.3″ S, 40°34′49.9″ W, 8–9 Sep. 2014, E.M. Rozário and F.F. Salles leg. (CZNC Ep-6439); 7 ♁♁, imagines, same data ( CZNCEp-6442);  1 ♀, imago, same data ( CZNCEp-6446);  IFES Itapina, Doce River, 19°31′15.1″ S, 40°46′53.6″ W, E.M. Rozário and F.F. Salles leg. (CZNC Ep-6448); 2 ♁♁, imagines, same data ( CZNCEp-6449); 5 ♁♁, imagines, Nova Venécia, city gate, Cricaré River, 18°42′54.91″ S, 40°22′33.33″ W,  23–24 May 2012,  F.F. Sallesand K.B. Angelileg. ( CZNCEp-4523); 1 ♁, 1 ♀, imagines, São Mateus, Cotaxé River, Japirawaterfall, 18°34′39.1″ S, 40°16′58″ W,   7–8 Sep. 2012, K.B. Angeliand E.A. Raimundileg. ( CZNCEp-4524); 1 ♁, imago, São Mateus River, 18°39′17.1″ S, 39°59′36″ W,   7–8 Sep. 2012, K.B. Angeliand E.A. Raimundileg. ( CZNCEp-4527); 1 ♁ imago, Pedra d’Água, 18°43′15.2″ S, 39°48′50.8″ W,   19–20 Apr. 2012, K.B. Angeliand E.A. Raimundileg. ( CZNCEp-4528); 4 ♁♁, imagines, Cotaxé River, Japirawaterfall, 18°34′39.1″ S, 40°16′58″ W,   6–7 Feb. 2013( CZNCEp-6159); 8 ♁♁, 6 ♀♀, imagines, Roraima, Caracaraí, Balneário Bem Querer, 01°55′46.3″ N, 61°00′06.9″ W, 13 Mar. 2014, F.F. Salles,  R. Boldrini, E. Dominguezleg. ( CZNCEp-6560); 3 ♁♁, Balneário Bem Querer, 18–21 Nov. 2002, A.M.O. Pes leg. (CZNC Ep-6561).    Diagnosis 1) Longitudinal vein CuP poorly developed in both sexes or absent; 2) membranous filaments of mesoscutellum absent; 3) forceps tri-segmented with first segment, distomedially projecting, 4) penis with very wide base, becoming thinner towards a subapical constriction, and then slightly widening again, penis lobes of each side almost completely fused except apical incision; 5) penis with a ventral projection covered with spines. As  M. australisis the only species for which nymphs have been described, the generic diagnostic features of the nymphal stage should also be used for specific diagnosis.   Distribution( Fig. 1)   Argentina( Misiones),  Brazil( Amazonas, Espírito Santo, Mato Grosso, Pará, Paraná, Roraima) and Guyana. 1420231162 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.521694 Espirito Santo 1 -40.836414 3 4 1 Espirito Santo 1420231159 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.521694 Espirito Santo 1 -40.836414 3 4 14 14 Espirito Santo 1420231171 [541,1109,684,713] 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.521694 Espirito Santo 1 -40.836414 3 4 1 Espirito Santo 1420231161 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.521694 Espirito Santo 1 -40.836414 3 4 1 Espirito Santo 1420231177 [487,1048,720,748] 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.521694 Espirito Santo 1 -40.836414 3 4 1 Espirito Santo 1420231167 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.518139 Doce River 1 -40.58053 Marilandia 3 4 1 Espirito Santo 1420231174 [786,1121,792,819] 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.518139 Doce River 1 -40.58053 Marilandia 3 4 1 Espirito Santo 1420231160 2014-09-07 2014-09-08 2014-09-07 CZNC E. M. Rozario & F. F. Salles Brazil -19.518139 Doce River 1 -40.58053 Marilandia 3 4 1 1 Espirito Santo 1420231169 [410,1054,825,854] IFES Brazil -19.52086 Doce River 1 -40.781555 Itapina 3 4 1 Espirito Santo 1420231170 2012-05-23 2012-05-24 2012-05-23 CZNC Brazil -18.715254 Cricare River 1 -40.375923 Nova Venecia 3 4 1 Espirito Santo 1420231181 CZNC F. F. Salles & K. B. Angeli Brazil Sao Mateus 4 5 -18.577528 Japira 14 -40.282776 Cotaxe River 3 4 1 1 Espirito Santo 1420231180 2012-09-07 2012-09-08 2012-09-07 CZNC K. B. Angeli & E. A. Raimundi Brazil -18.654749 Sao Mateus River 14 -39.993332 4 5 1 Espirito Santo 1420231164 2012-09-07 2012-09-08 2012-09-07 CZNC K. B. Angeli & E. A. Raimundi Brazil -18.72089 Pedra d'Agua 1 -39.81411 4 5 1 Espirito Santo 1420231178 2012-04-19 2012-04-20 2012-04-19 CZNC K. B. Angeli & E. A. Raimundi Brazil -18.577528 Japira 14 -40.282776 Cotaxe River 4 5 1 Espirito Santo 1420231172 2013-02-06 2013-02-07 2013-02-06 CZNC Brazil 1.9295279 Balneario Bem Querer 1 -61.001915 Caracarai 4 5 6 6 Roraima 1420231183 CZNC R. Boldrini & E. Dominguez Brazil Balneario Bem Querer 4 5 1 Roraima 1420231166 [189,447,944,971] Argentina Misiones 4 5 1 Misiones 1420231182 Brazil Amazonas, Espirito Santo, Mato Grosso, Para, Parana, Roraima 4 5 1 Amazonas, Espirito Santo, Mato Grosso, Para, Parana, Roraima