Heteromeringia spinulosa D.K. McAlpine, 1960:74 Heteromeringia pulla D.K. McAlpine, 1960: 73 The Heteromeringia (Diptera: Clusiidae: Clusiodinae) of Australia Lonsdale, Owen Records of the Australian Museum 2009 2009-11-25 61 3 229 262 6LSVJ D. K. McAlpine, 1960 D. K. McAlpine 1960 [854,1452,1183,1209] Insecta Clusiidae Heteromeringia Animalia Diptera 26 255 Arthropoda species spinulosa   Figs 1, 2, 5, 11, 73–80, 104, 105      Heteromeringia spinulosaD.K. McAlpine, 1960:74.   Heteromeringia pullaD.K. McAlpine, 1960: 73,  new synonym.     Holotype[  Heteromeringia spinulosa]:  New South Wales: Below Govett’s Leap, Blue Mountains,  7.xii.1956, D.K. McAlpine( 1♂, AMS).   Paratypes[  H. spinulosa].  New South Wales: Otford,  12.x.1957, D.K. McAlpine( 1♂, AMS), Sydney,  ii.1925, Health Department ( 1♀, AMS) [ allotypeof  H. australiae].    Holotype[  Heteromeringia pulla].  New South Wales: National Park,  13.iv.1954, D.K. McAlpine( 1♂, AMS).   Paratypes[  H. pulla].  New South Wales: National Park, D.K. McAlpine,  13.iv.1954( 2♂♂, AMS),   20.iv.1954( 4♂♂, AMS),  Otford,  26.i.1959, D.K. McAlpine( 1♂, AMS).  Additional material examined.   Australian Capital Territory. Black Mtn., Malaise, site 1, D.H. Colless,  20.ii.1980( 1♂ 1♀, ANIC),   18.ii.1980( 1♂, ANIC),   4–5.iii.1980( 1♂, ANIC),   29.xi.1979( 1♀, ANIC),   30.xi–2.xii.1979( 1♀, ANIC),   4.ii.1980( 1♀, ANIC),   14.ii.1980( 1♂, ANIC),   21.ii.1980( 1♂, ANIC),   25.ii.1980( 1♂, ANIC),   8–10.ii.1980( 1♀, ANIC),   2–3.ii.1980( 1♀, ANIC),   20.xi. 1980( 1♀, ANIC),   19.xi.1979( 2♂♂, ANIC),   28.ii.1980( 1♀, ANIC),   17–18.xi.1979( 1♀, ANIC),   29.ii.1980( 1♂ 1♀, ANIC),   15–17. iii.1980( 1♀, ANIC),   15–17.xii.1979( 2♂♂, ANIC),   29.ii–2.iii.1980( 1♂ 1♀, ANIC),  Black Mtn.,  xii.1987, M. Irwin, Malaise trap( 2♂♂ 2♀♀, ANIC),  Blundells Ck.,  ii.1987, D.H. Colless, Malaise trap( 2♀♀, ANIC).   New South Wales: Katoomba,  19.xi.1963, G.H. Hardy( 1♂, AMS),   8.xii.1961( 2♀♀, AMS),  Sassafras Gully, Springwood,  23.ix.1972, D.K. McAlpine( 1♂, AMS),  Wright’s Lookout, New England National Park,  1.iv.1961, D.K. McAlpine( 1♀, AMS),  Roberts Creeknear East Kurrajong,  1.i.1962, D.K. McAlpine( 1♀, AMS),  Mooney Mooney Creeknear Gosford,  20.xi.1975, D.K.McAlpine( 1♂, AMS),   25.xi.1975( 1♂, AMS),   9.xi.1976, in copula ( 1♂ 1♀[same pin], AMS),   11.xii.1981, B.J. Day( 1♀, AMS),   16.xi.1978, D.K. McAlpine& B.J. Day( 1♂, AMS),   25.xi.1975, D.K. McAlpine& B.J. Day( 1♂, AMS),   6.xii.1977, D.K. McAlpine& B.J. Day( 1♂, AMS),   1.xii.1989, D.K. McAlpine& B.J. Day( 1♂, AMS),  National Park, 19..  iii.1961, D.K. McAlpine( 1♀ 1♂[same pin], AMS),  Brown Mtn.nr. Nimmitabel,  13–19.ii.1987,  1200 m, warm temp. rf., D. Bickel( 1♀, AMS),  Royal NP,  12.ii.1985, Bela Ck., closed for., D. Bickel( 1♂, AMS),  Royal National Park,  19.xii.1970, D.K. McAlpine( 1♂, AMS),  Careel Bay,  14.iii.1963, Casuarinaswamp, D.K. McAlpine( 1♂ 1♀, AMS), ne Camden HeadSEPP 26 site 122, littoral r’for, G.&T.Williams, Malaise trap,  24.xii.1999–  4.i.2000( 1♀, AMS),  Sullivan Gap FR, nr. Buladelah,  3–16.i.1988, yellow pan trap, E. Tasker,  930 m,  24–30.iv.1998, sticky trapon E. cameronii, CS-FZ-048-5 ( 1♂, AMS),   11–16.i.1998, sticky trapon E. obliqua, CC-FK-018-5 ( 2♀♀, USNM),   11–16.i.1998, sticky trapon E. campanula, CC-CR-018-6 ( 1♀, AMS),   3–8.xii.1997, sticky trapon E. viminalis, CS-RO-127-1 ( 1♀, AMS),  Deua NP, Wyanbene Caves Cpgd.,  800 m, 35°45'30"S 149°39'30"E, sticky trap, trunk, wet sclera. for., D. Bickel,  16.xii.2000( 1♀, AMS),   3kmN Lansdownevia Taree, Lorien Wildlife Refuge,  50 m,  20–27.xii.1990, Malaise trap, D. Pollock& L. Reichert( 1♂, USNM),  Royal Natl. Pk., Scientists’ Cabintrail, creek bed, sweep,  25.xii.2003, S.A. Marshal( 2♀♀, DEBU) [one with abdomen missing], Upp.Kangaroo Riv., nr. Yeola,  200 m, J.F. Lawrence, ex. rotting stick on rainforest floor, coll  3–4.iii.1983, Emg.  11.iii.1983( 1♂, ANIC),  Emg.  9.iii.1983( 1♀, ANIC),  Emg.  20–30.iii.1983( 1♂ 2♀♀, ANIC),  Emg.  15–20.iii.1983( 1♀, ANIC),  Monga State Forest,  24.i.1984, L. Masner( 1♀, CNC),   19–24.i.1984( 1♀, CNC).   Queensland: Monto District, Bulburin( Austral), Site3, 24°34'S 151°29'E,  580 m, creek bed,  20.iii.1975, D.K. McAlpine( 1♂, AMS),  Austral Forestvia Bulburin, Site3, 24°34'S 151°29'E, 22.iii.197, Malaise trap, D.K. McAlpine( 1♂, AMS),  Great Sandy Nat. Pk., Cooloola Section, 25°56'49"S 153°05'30"E,  1–5.x.1996, D.K.Yeates, G. Lambkin& S. Winterton, Malaise trap( 2♂♂, UQIC),  Cooloola NP, Freshwater Lake,  27.xi.1985, D. Bickel& G. Cassis( 1♂, AMS),  Bunya Mtns NP. At Rd.to Nanaga, rainfor.,  960 m,  26.xi.1992, yell. Pans, D.J. Bickel( 1♀, AMS), SE QLD, Mt.Glorious, Rainforest Circuittr,  24.x.2002, 27.20S/ 152.46E, rainforest, Merz, Földvari& McNeil( 2♂♂, MHNG), SE QLD, Brisbane, Griffith Uni., Nathan Campus,  23.x.2002, 27.33S/ 153.04E, dry&wet sclrophyll forest, ferns, B. Merz( 1♂, MHNG),  Mt Glorious, 27°19'54"S 152°45'29"E, Malaise,  24–30.i.1998, T. Hiller( 2♂♂ 1♀, DEBU),   24–30.xi.1997( 1♂, DEBU),   3–9.i.1998( 1♂ 2♀♀, DEBU),  Brisbane ForestPk., Scrub Creek Rd., 27°25'41"S 152°50'15"E, yellow pans,  29.ix.2002, C.R. Nelson( 1♂, DEBU), SE QLD, Tamborine Mts., Eagle Heights, Palm Grove,  26.x.2002, 27.56S/ 153.12E, rainforest, Merz& Földvari( 1♂, MHNG), N QLD, the Craternear Herberton,  16.xii.1961, D.K. McAlpine( 1♂, AMS),  Birthday Crk, near Paluma, N Qld,  18.i.1967, D.K. McAlpine& G. Holloway( 1♂, AMS; 2♂♂, USNM),  14.i.1970, G.A. Holloway( 1♂, AMS),  Skywindow Lookout, Eungella Nat. Pk.,  8–9.v.1980, I.D. Naumann& J.C. Cardale, Malaise trap( 1♀, ANIC),  N Qld, Julatten, A. Walford-Huggins, edge of rainforest along creek, ex. intercept trap,  30.xi–13.xii.1987( 2♂♂ 1♀, ANIC),   5.x.1987( 1♀, ANIC),  13.32S 143.19E, McIlwraith Range,  vi–vii.1989, I. Naumann( 1♂, ANIC),  Sirakan,  ii.1924, G.H. Hardy( 1♀, UQIC),  AthertonTablelands, Lake Eacham, sweep,  5–8.iv.1999, S.A. Marshall( 1♀, DEBU),  Brisbane ForestPk, 27°25'5"S 152°50'13"E, Malaiseover creek,  13–19.vi.1998, N. Power( 1♂, DEBU),  Mt Glorious, 27°19'54"S 152°45'29"E, Malaise,  24–30. xi.1998, T. Hiller( 3♀♀, DEBU),   24–30.i.1998( 1♂, DEBU), SE QLD, Brisbane, Griffith Uni., Nathan Campus,  23.x.2002, 27.33S/ 153.04E, dry&wet sclrophyll forest, ferns, B. Merz( 1♂, MHNG), N QLD, Mt. Spurgeon, wet sclera. For.,  1250 m,  18–19.iv.1994, B.J. Sinclair, 16°26'S 145°12'E( 1♀, CNC),  Monga State Forest,  19–24.i.1984, L. Masner( 1♀, CNC),  Mt. Glorious,  x.1978, A. Hiller, Malaise( 1♀, ANIC),  Mt. Haig, 21kmNE by E of Atherton,  18.xi.1981, D.H. Colless, Malaise trap( 1♂, ANIC),  15.50S 145.20E, Gap Ck.,  5kmESE Mt. Finnigan,  15.v.1981, D.H. Colless, Malaise trap( 1♀, ANIC),  Great Sandy Nat. Pk., Cooloola Section, 25°56'49"S 153°05'30"E,  1–5.x.1996, D.K. Yeates, G. Lambkin& S. Winterton, Malaise trap( 1♂, UQIC),  Noosa Natl. Pk., Palm Trail, sweeping,  30.iv.1999, S.A. Marshall( 1♂, DEBU).   Tasmania: Claytons,  i.1991, Nielsen& Edwards( 1♂ 1♀, ANIC).   Victoria: Sherbrook Forestnear Melbourne Via.,  21.i.1966, D.K. McAlpine( 1♀, AMS).   Redescription  Male( Figs 1, 2, 5). Body length 2.7–5.2 mm. Anepisternal disc present. First flagellomere orbicular. Bristles black. Arista pubescent. Vibrissa relatively long and curved. Ocellar bristle thin and approximately as long as tubercle. Two dorsocentral bristles with small bristle in front of anterior dorsocentral. Gena relatively high and sometimes with inward bend at mid-height (not as distinct as in  H. laticornis). Face and buccal cavity velvety and meeting at slightly rounded angle. Head yellow with posterior  2⁄ 3of frons (usually excluding lateral and posterior margins) dark brown (remainder orange) and back of head above foramen dark brown; first flagellomere sometimes with narrow infuscation along anterior margin or around base of arista; USNM specimen with distal  2⁄  3of first flagellomere dark brown; occiput brown (venter often yellow in southern specimens) and gena yellow with posterior half sometimes brown (more frequently brown in northern specimens); clypeus brown; anterior half of frons pilose and dorsal margin of gena silvery tomentose. Thorax dark brown with proepisternum yellow and posterolateral margin of scutum to anterolateral margin of scutellum sometimes with variably yellow to orange tint (more common northwards). Legs yellow with fore coxa white, fore tarsi dark brown and laterally compressed, fore tibia brownish (lighter to base), and fore femur with long, illdefined spot on inner apex. Abdomen dark brown. M 1+2ratio 3.7–5.3. Wing with basal band confluent with second band through cells br and bm; second and third bands (confluent through first radial cell) separated evenly by thin clear strip. Halter white with base and side of stalk infuscated.  Female. As described for male except as follows: pale spots on scutum faded or absent; anterior and inner-distal margins of first flagellomere infuscated, sometimes with distal  2⁄  3of inner face dark; frons shiny or with small pilose anteromedial patch; only anterior and lateral margins of frons with orange tint; face and gena with piceous tint; fore tibia and tarsi evenly brown; terminalia yellow.  Male terminalia( Figs 73–77). Length of epandrium slightly more than half width and height. Surstylus  2⁄ 3height of epandrium, very thin and pointed, with few inner-apical tubercle-like bristles and outer face bare. Cerci approximately as high as surstylus, heart-shaped and heavily setose. Hypandrium+pregonite bilobed (anterior lobe with two stout bristles and posterior lobe setulose at apex and base). Phallapodeme shorter than hypandrium. Distiphallus with membrane on basal half strongly spinulose; with small serrated process medially (vestigial, shallow, slightly longer than wide, or very long and blade-like); wide and heavily elaborated apically with serrated subapical ridge.  Female terminalia(Figs 104, 105). Ventral receptacle rounded apically and flagellum very long, thin and coiled (coils tighter to apex). Spermatheca barrel-shaped (widest subapically) and long with several basal wrinkles.   Comments. Based on external and internal examination of male and female specimens of  Heteromeringia spinulosaand  H. pullafrom across both of their ranges, it now appears that these two taxa belong to the same species. Their colouration is nearly identical, as are the female genitalia, surstylus and most of the hypandrial complex. The absence of a serrated blade-like process on the phallus and its associated bell-shaped apical lobe ( Fig. 77) was previously considered diagnostic of  H. pulla(although D.K. McAlpine (1960)noted some polymorphism), but these are actually present in a number of males, although the blade is rarely much longer than wide if present and the lobe is frequently underdeveloped ( Fig. 75). The other character previously considered diagnostic of either species was whether or not the venter of the gena was entirely (  H. spinulosa) or only partially (  H. pulla) yellow. While the presence of either character is generally indicative of whether or not the phallic blade is present, it is sometimes not, and specimens collected in and south of the Australian Capital Territoryalmost always have an entirely yellow gena (the occiput is also predominantly yellow) regardless of whether the blade is developed or not. The presence of a strong yellow patch extending from the side of the scutellum to the posterolateral corner of the notum is slightly more correlated with the presence of a well developed phallic blade, but the intensity of this patch often varies and is sometimes associated with the “opposite” genitalic state, particularly to the north. Since all of the specimens examined appear to belong to a single (albeit unusually polymorphic) species,  H. pullais here treated as a junior synonym of  H. spinulosaunder the first reviser principle (ICZN Article 24.2.1).   Inaddition to the material listed above, there are several specimens that range from the Australian Capital Territoryto Queenslandthat vary as follows: first flagellomere entirely white; ventral margin of gena and occiput entirely yellow; notum with faint yellowish posterolateral tint; wing pattern relatively indistinct; surstylus much longer than epandrium ( Figs 78, 79); phallus branched at midpoint, with branch short, smooth and directed distally ( Fig. 80). Theelongate surstylus is the most striking feature of these males, but given the amount of genitalic variability already discussed for this species, they will be treated as  H. spinulosafor the time being. Labeldata:ACT: 35.22S 148.50E, Blundells Ck.,  ii.1987, D.H. Colless, Malaise trap( 3♂♂, ANIC). NSW: Carrai SF, 30°54'35"S 152°16'26"E,  1090 m, E. Tasker,  11–16.i.1998, sticky trapon E. obliqua, CC-FK-018-5 ( 1♂, AMS). QLD: 18.55S 146.10E, S2 Mt. Spec,  880 m,  5.vi–3. vii.1995, M. Cermak, Fl trap, JCU (East) ( 1♂, ANIC). 1956-12-07 AMS D. K. McAlpine Blue Mountains Below Govett's Leap 26 255 1 1 New South Wales holotype 1925-02 1957-10-12 1925-02 AMS D. K. McAlpine Otford 26 255 2 1 1 New South Wales paratype 1954-04-13 AMS D. K. McAlpine National Park 26 255 1 1 New South Wales holotype 1954-04-13 AMS National Park, D. K New South Wales 26 255 2 2 New South Wales paratype 1954-04-20 AMS National Park, D. K true New South Wales 26 255 4 4 New South Wales paratype [962,1359,1637,1658] 1959-01-26 AMS D. K. McAlpine Otford 26 255 1 1 New South Wales paratype 1980-02-20 ANIC D. H. Colless Malaise Black Mtn. 26 255 2 1 1 Australian Capital Territory 1980-02-18 ANIC D. H. Colless true Malaise Black Mtn. 26 255 1 1 Australian Capital Territory [907,1116,1738,1759] 1980-03-04 1980-03-05 1980-03-04 ANIC 26 255 1 1 Australian Capital Territory [1123,1320,1738,1759] 1979-11-29 ANIC 26 255 1 1 Australian Capital Territory 1979-11-30 1979-12-02 1979-11-30 ANIC 26 255 1 1 Australian Capital Territory [949,1131,1762,1783] 1980-02-04 ANIC 26 255 1 1 Australian Capital Territory [1138,1331,1762,1783] 1980-02-14 ANIC 26 255 1 1 Australian Capital Territory 1980-02-21 ANIC 26 255 1 1 Australian Capital Territory [907,1100,1786,1807] 1980-02-25 ANIC 26 255 1 1 Australian Capital Territory [1107,1320,1786,1807] 1980-02-08 1980-02-10 1980-02-08 ANIC 26 255 1 1 Australian Capital Territory 1980-02-02 1980-02-03 1980-02-02 ANIC 26 255 1 1 Australian Capital Territory [907,1109,1810,1831] 1980-11-20 ANIC 26 255 1 1 Australian Capital Territory [1116,1330,1810,1831] 1979-11-19 ANIC 26 255 2 2 Australian Capital Territory 1980-02-28 ANIC 26 255 1 1 Australian Capital Territory [911,1150,1834,1855] 1979-11-17 1979-11-18 1979-11-17 ANIC 26 255 1 1 Australian Capital Territory [1161,1397,1834,1855] 1980-02-29 ANIC 26 255 2 1 1 Australian Capital Territory 1980-03-15 1980-03-17 1980-03-15 ANIC 26 255 1 1 Australian Capital Territory [1014,1261,1858,1879] 1979-12-15 1979-12-17 1979-12-15 ANIC 26 255 2 2 Australian Capital Territory 1980-02-29 1980-03-02 1980-02-29 ANIC 26 255 2 1 1 Australian Capital Territory [909,1463,1882,1903] 1987-12 ANIC M. Irwin Black Mtn. 26 255 4 2 2 Australian Capital Territory [839,1415,1906,1927] 1987-02 ANIC D. H. Colless Blundells Ck. 26 255 2 2 Australian Capital Territory 1963-11-19 AMS G. H. Hardy Katoomba 26 255 1 1 New South Wales 1961-12-08 AMS 26 255 2 2 New South Wales 1972-09-23 AMS D. K. McAlpine Springwood Sassafras Gully 26 255 1 1 New South Wales 1961-04-01 AMS D. K. McAlpine 27 256 New England National Park Wright's Lookout 26 255 1 1 New South Wales 1962-01-01 AMS D. K. McAlpine East Kurrajong Roberts Creek 27 256 1 1 New South Wales 1975-11-20 AMS D. K. McAlpine Gosford Mooney Mooney Creek 27 256 1 1 New South Wales [355,542,1717,1738] 1975-11-25 AMS 27 256 1 1 New South Wales 1976-11-09 AMS 27 256 2 1 1 New South Wales [314,598,1741,1762] 1981-12-11 AMS Day 27 256 1 1 New South Wales 1978-11-16 AMS McAlpine & Day 27 256 1 1 New South Wales 1975-11-25 AMS McAlpine & Day 27 256 1 1 New South Wales [225,650,1789,1810] 1977-12-06 AMS McAlpine & Day 27 256 1 1 New South Wales 1989-12-01 AMS McAlpine & Day 27 256 1 1 New South Wales 1961-03 AMS D. K. McAlpine National Park 27 256 2 1 1 New South Wales 1987-02-13 1987-02-19 1987-02-13 AMS D. Bickel 1200 Nimmitabel Brown Mtn. 27 256 1 1 New South Wales 1985-02-12 AMS Royal NP & Bela Ck. & D. Bickel 27 256 1 1 New South Wales 1970-12-19 AMS D. K. McAlpine Royal National Park 27 256 1 1 New South Wales 1963-03-14 AMS Casuarina & D. K. McAlpine Careel Bay 27 256 2 1 1 New South Wales 1999-12-24 2000-01-04 1999-12-24 AMS G. & T. Williams Camden Head 27 256 1 1 New South Wales 1988-01-03 1998-04-30 1988-01-03 AMS Sullivan Gap FR & E. Tasker 930 Buladelah 27 256 1 1 New South Wales 1998-01-11 1998-01-16 1998-01-11 USNM 27 256 2 2 New South Wales 1998-01-11 1998-01-16 1998-01-11 AMS 27 256 1 1 New South Wales [856,1418,1644,1665] 1997-12-03 1997-12-08 1997-12-03 AMS 27 256 1 1 New South Wales 2000-12-16 AMS Deua NP & D. Bickel 800 -35.758335 Wyanbene Caves Cpgd. 19 149.65833 27 256 1 1 New South Wales 1990-12-20 1990-12-27 1990-12-20 USNM Pollock & Reichert 50 Lorien Wildlife Refuge Taree 27 256 1 1 New South Wales 2003-12-25 DEBU Royal Natl. Pk. & S. A. Marshal Scientists' Cabin 27 256 2 2 New South Wales 1983-03-03 1983-03-11 1983-03-03 ANIC J. F. Lawrence & Emg. 200 Yeola Upp. Kangaroo Riv. 27 256 1 1 New South Wales [1001,1240,1836,1857] 1983-03-09 ANIC Emg. 27 256 1 1 New South Wales 1983-03-20 1983-03-30 1983-03-20 ANIC Emg. 27 256 3 2 1 New South Wales [955,1241,1860,1881] 1983-03-15 1983-03-20 1983-03-15 ANIC Emg. 27 256 1 1 New South Wales 1984-01-24 CNC L. Masner Monga State Forest 27 256 1 1 New South Wales [1084,1299,1884,1905] 1984-01-19 1984-01-24 1984-01-19 CNC 27 256 1 1 New South Wales 1975-03-20 AMS McAlpine Monto District 580 -24.566668 Austral 1250 151.48334 Bulburin 27 256 1 1 Queensland AMS D. K. McAlpine Austral Forest -24.566668 Site 1250 151.48334 Bulburin 27 256 1 1 Queensland 1996-10-01 1996-10-05 1996-10-01 UQIC D. K. Yeates & G. Lambkin & Winterton 28 257 -25.946943 Cooloola Section 20 153.09166 Great Sandy Nat. Pk. 27 256 2 2 Queensland 1985-11-27 AMS Cooloola NP & D. Bickel & Cassis Freshwater Lake 28 257 1 1 Queensland 1992-11-26 AMS Bunya Mtns NP & Pans & D. J. Bickel 960 Nanaga At Rd. 28 257 1 1 Queensland 2002-10-24 MHNG Merz & Foldvari & McNeil -27.2 Rainforest Circuit 743 152.46 Mt. Glorious 28 257 2 2 Queensland 2002-10-23 MHNG B. Merz Brisbane -27.33 Nathan Campus 742 153.04 Griffith Uni. 28 257 1 1 Queensland 1998-01-24 1998-01-30 1998-01-24 DEBU Hiller -27.331667 Malaise 20 152.75806 Mt Glorious 28 257 3 1 2 Queensland [344,592,1984,2005] 1997-11-24 1997-11-30 1997-11-24 DEBU 28 257 1 1 Queensland [603,906,1792,2005] 1998-01-03 1998-01-09 1998-01-03 DEBU 28 257 3 2 1 Queensland 2002-09-29 DEBU C. R. Nelson -27.428055 Scrub Creek Rd. 20 152.8375 Brisbane Forest 28 257 1 1 Queensland 2002-10-26 MHNG Merz & Foldvari Tamborine Mts. -27.56 Palm Grove 742 153.12 Eagle Heights 28 257 1 1 Queensland 1961-12-16 AMS D. K. McAlpine Herberton Crater 28 257 1 1 Queensland 1967-01-18 1970-01-14 1967-01-18 AMS, USNM D. K. McAlpine & Holloway & G. A. Holloway Birthday Crk Qld Paluma 28 257 4 4 Queensland 1980-05-08 1980-05-09 1980-05-08 ANIC I. D. Naumann & J. C. Cardale Eungella Nat. Pk. Skywindow Lookout 28 257 1 1 Queensland 1987-11-30 1987-12-13 1987-11-30 ANIC N Qld & Julatten, A 29 258 Walford-Huggins 28 257 3 1 2 Queensland [561,745,154,175] 1987-10-05 1987-12-13 1987-11-30 ANIC N Qld & Julatten, A true Walford-Huggins 29 258 1 1 Queensland [121,745,178,199] 1989-06 1989-07-31 1989-06 ANIC I. Naumann -13.32 McIlwraith Range 774 143.19 29 258 1 1 Queensland [121,501,202,223] 1924-02 UQIC G. H. Hardy Sirakan 29 258 1 1 Queensland 1999-04-05 1999-04-08 1999-04-05 DEBU S. A. Marshall Lake Eacham Atherton 29 258 1 1 Queensland 1998-06-13 1998-06-19 1998-06-13 DEBU N. Power -27.418055 Malaise 20 152.83694 Brisbane Forest 29 258 1 1 Queensland 1998-11-24 1998-11-30 1998-11-24 DEBU Hiller -27.331667 Malaise 20 152.75806 Mt Glorious 29 258 3 3 Queensland [421,656,298,319] 1998-01-24 1998-01-30 1998-01-24 DEBU 29 258 1 1 Queensland 2002-10-23 MHNG B. Merz Brisbane -27.33 Nathan Campus 742 153.04 Griffith Uni. 29 258 1 1 Queensland 1994-04-18 1994-04-19 1994-04-18 CNC Sinclair 1250 -16.433332 For. 1282 145.2 Mt. Spurgeon 29 258 1 1 Queensland 1984-01-19 1984-01-24 1984-01-19 CNC L. Masner Monga State Forest 29 258 1 1 Queensland [185,653,418,439] 1978-10 ANIC A. Hiller & Malaise Mt. Glorious 29 258 1 1 Queensland 1981-11-18 ANIC D. H. Colless Mt. Haig 29 258 1 1 Queensland 1981-05-15 ANIC Gap Ck. & D. H. Colless -15.5 5 km ESE Mt. Finnigan 771 145.2 29 258 1 1 Queensland 1996-10-01 1996-10-05 1996-10-01 UQIC D. K. Yeates & G. Lambkin & Winterton -25.946943 Cooloola Section 20 153.09166 Great Sandy Nat. Pk. 29 258 1 1 Queensland 1999-04-30 DEBU Noosa Natl. Pk. & S. A. Marshall Palm Trail 29 258 1 1 Queensland 1991-01 ANIC Nielsen & Edwards Claytons 29 258 2 1 1 Tasmania 1966-01-21 AMS D. K. McAlpine Melbourne Via. Sherbrook Forest 29 258 1 1 Victoria 1987-02 ANIC Blundells Ck. & D. H. Colless In -35.22 Label 716 148.5 The 29 258 3 3 Australian Capital Territory 1998-01-11 1998-01-16 1998-01-11 AMS Carrai SF & E. Tasker 1090 -30.909721 20 152.27388 29 258 1 1 Australian Capital Territory 1995-06-05 1995-07-03 1995-06-05 ANIC M. Cermak 880 -18.55 Mt. Spec 765 146.1 29 258 1 1 Australian Capital Territory