Heteromeringia spinulosa D.K. McAlpine, 1960:74
Heteromeringia pulla D.K. McAlpine, 1960: 73
The Heteromeringia (Diptera: Clusiidae: Clusiodinae) of Australia
Lonsdale, Owen
Records of the Australian Museum
2009
2009-11-25
61
3
229
262
6LSVJ
D. K. McAlpine, 1960
D. K. McAlpine
1960
[854,1452,1183,1209]
Insecta
Clusiidae
Heteromeringia
Animalia
Diptera
26
255
Arthropoda
species
spinulosa
Figs 1, 2, 5, 11, 73–80, 104, 105
Heteromeringia spinulosaD.K. McAlpine, 1960:74. Heteromeringia pullaD.K. McAlpine, 1960: 73, new synonym.
Holotype[ Heteromeringia spinulosa]: New South Wales: Below Govett’s Leap, Blue Mountains, 7.xii.1956, D.K. McAlpine( 1♂, AMS). Paratypes[ H. spinulosa]. New South Wales: Otford, 12.x.1957, D.K. McAlpine( 1♂, AMS), Sydney, ii.1925, Health Department ( 1♀, AMS) [ allotypeof H. australiae]. Holotype[ Heteromeringia pulla]. New South Wales: National Park, 13.iv.1954, D.K. McAlpine( 1♂, AMS). Paratypes[ H. pulla]. New South Wales: National Park, D.K. McAlpine, 13.iv.1954( 2♂♂, AMS), 20.iv.1954( 4♂♂, AMS), Otford, 26.i.1959, D.K. McAlpine( 1♂, AMS). Additional material examined. Australian Capital Territory. Black Mtn., Malaise, site 1, D.H. Colless, 20.ii.1980( 1♂ 1♀, ANIC), 18.ii.1980( 1♂, ANIC), 4–5.iii.1980( 1♂, ANIC), 29.xi.1979( 1♀, ANIC), 30.xi–2.xii.1979( 1♀, ANIC), 4.ii.1980( 1♀, ANIC), 14.ii.1980( 1♂, ANIC), 21.ii.1980( 1♂, ANIC), 25.ii.1980( 1♂, ANIC), 8–10.ii.1980( 1♀, ANIC), 2–3.ii.1980( 1♀, ANIC), 20.xi. 1980( 1♀, ANIC), 19.xi.1979( 2♂♂, ANIC), 28.ii.1980( 1♀, ANIC), 17–18.xi.1979( 1♀, ANIC), 29.ii.1980( 1♂ 1♀, ANIC), 15–17. iii.1980( 1♀, ANIC), 15–17.xii.1979( 2♂♂, ANIC), 29.ii–2.iii.1980( 1♂ 1♀, ANIC), Black Mtn., xii.1987, M. Irwin, Malaise trap( 2♂♂ 2♀♀, ANIC), Blundells Ck., ii.1987, D.H. Colless, Malaise trap( 2♀♀, ANIC). New South Wales: Katoomba, 19.xi.1963, G.H. Hardy( 1♂, AMS), 8.xii.1961( 2♀♀, AMS), Sassafras Gully, Springwood, 23.ix.1972, D.K. McAlpine( 1♂, AMS), Wright’s Lookout, New England National Park, 1.iv.1961, D.K. McAlpine( 1♀, AMS), Roberts Creeknear East Kurrajong, 1.i.1962, D.K. McAlpine( 1♀, AMS), Mooney Mooney Creeknear Gosford, 20.xi.1975, D.K.McAlpine( 1♂, AMS), 25.xi.1975( 1♂, AMS), 9.xi.1976, in copula ( 1♂ 1♀[same pin], AMS), 11.xii.1981, B.J. Day( 1♀, AMS), 16.xi.1978, D.K. McAlpine& B.J. Day( 1♂, AMS), 25.xi.1975, D.K. McAlpine& B.J. Day( 1♂, AMS), 6.xii.1977, D.K. McAlpine& B.J. Day( 1♂, AMS), 1.xii.1989, D.K. McAlpine& B.J. Day( 1♂, AMS), National Park, 19.. iii.1961, D.K. McAlpine( 1♀ 1♂[same pin], AMS), Brown Mtn.nr. Nimmitabel, 13–19.ii.1987, 1200 m, warm temp. rf., D. Bickel( 1♀, AMS), Royal NP, 12.ii.1985, Bela Ck., closed for., D. Bickel( 1♂, AMS), Royal National Park, 19.xii.1970, D.K. McAlpine( 1♂, AMS), Careel Bay, 14.iii.1963, Casuarinaswamp, D.K. McAlpine( 1♂ 1♀, AMS), ne Camden HeadSEPP 26 site 122, littoral r’for, G.&T.Williams, Malaise trap, 24.xii.1999– 4.i.2000( 1♀, AMS), Sullivan Gap FR, nr. Buladelah, 3–16.i.1988, yellow pan trap, E. Tasker, 930 m, 24–30.iv.1998, sticky trapon E. cameronii, CS-FZ-048-5 ( 1♂, AMS), 11–16.i.1998, sticky trapon E. obliqua, CC-FK-018-5 ( 2♀♀, USNM), 11–16.i.1998, sticky trapon E. campanula, CC-CR-018-6 ( 1♀, AMS), 3–8.xii.1997, sticky trapon E. viminalis, CS-RO-127-1 ( 1♀, AMS), Deua NP, Wyanbene Caves Cpgd., 800 m, 35°45'30"S 149°39'30"E, sticky trap, trunk, wet sclera. for., D. Bickel, 16.xii.2000( 1♀, AMS), 3kmN Lansdownevia Taree, Lorien Wildlife Refuge, 50 m, 20–27.xii.1990, Malaise trap, D. Pollock& L. Reichert( 1♂, USNM), Royal Natl. Pk., Scientists’ Cabintrail, creek bed, sweep, 25.xii.2003, S.A. Marshal( 2♀♀, DEBU) [one with abdomen missing], Upp.Kangaroo Riv., nr. Yeola, 200 m, J.F. Lawrence, ex. rotting stick on rainforest floor, coll 3–4.iii.1983, Emg. 11.iii.1983( 1♂, ANIC), Emg. 9.iii.1983( 1♀, ANIC), Emg. 20–30.iii.1983( 1♂ 2♀♀, ANIC), Emg. 15–20.iii.1983( 1♀, ANIC), Monga State Forest, 24.i.1984, L. Masner( 1♀, CNC), 19–24.i.1984( 1♀, CNC). Queensland: Monto District, Bulburin( Austral), Site3, 24°34'S 151°29'E, 580 m, creek bed, 20.iii.1975, D.K. McAlpine( 1♂, AMS), Austral Forestvia Bulburin, Site3, 24°34'S 151°29'E, 22.iii.197, Malaise trap, D.K. McAlpine( 1♂, AMS), Great Sandy Nat. Pk., Cooloola Section, 25°56'49"S 153°05'30"E, 1–5.x.1996, D.K.Yeates, G. Lambkin& S. Winterton, Malaise trap( 2♂♂, UQIC), Cooloola NP, Freshwater Lake, 27.xi.1985, D. Bickel& G. Cassis( 1♂, AMS), Bunya Mtns NP. At Rd.to Nanaga, rainfor., 960 m, 26.xi.1992, yell. Pans, D.J. Bickel( 1♀, AMS), SE QLD, Mt.Glorious, Rainforest Circuittr, 24.x.2002, 27.20S/ 152.46E, rainforest, Merz, Földvari& McNeil( 2♂♂, MHNG), SE QLD, Brisbane, Griffith Uni., Nathan Campus, 23.x.2002, 27.33S/ 153.04E, dry&wet sclrophyll forest, ferns, B. Merz( 1♂, MHNG), Mt Glorious, 27°19'54"S 152°45'29"E, Malaise, 24–30.i.1998, T. Hiller( 2♂♂ 1♀, DEBU), 24–30.xi.1997( 1♂, DEBU), 3–9.i.1998( 1♂ 2♀♀, DEBU), Brisbane ForestPk., Scrub Creek Rd., 27°25'41"S 152°50'15"E, yellow pans, 29.ix.2002, C.R. Nelson( 1♂, DEBU), SE QLD, Tamborine Mts., Eagle Heights, Palm Grove, 26.x.2002, 27.56S/ 153.12E, rainforest, Merz& Földvari( 1♂, MHNG), N QLD, the Craternear Herberton, 16.xii.1961, D.K. McAlpine( 1♂, AMS), Birthday Crk, near Paluma, N Qld, 18.i.1967, D.K. McAlpine& G. Holloway( 1♂, AMS; 2♂♂, USNM), 14.i.1970, G.A. Holloway( 1♂, AMS), Skywindow Lookout, Eungella Nat. Pk., 8–9.v.1980, I.D. Naumann& J.C. Cardale, Malaise trap( 1♀, ANIC), N Qld, Julatten, A. Walford-Huggins, edge of rainforest along creek, ex. intercept trap, 30.xi–13.xii.1987( 2♂♂ 1♀, ANIC), 5.x.1987( 1♀, ANIC), 13.32S 143.19E, McIlwraith Range, vi–vii.1989, I. Naumann( 1♂, ANIC), Sirakan, ii.1924, G.H. Hardy( 1♀, UQIC), AthertonTablelands, Lake Eacham, sweep, 5–8.iv.1999, S.A. Marshall( 1♀, DEBU), Brisbane ForestPk, 27°25'5"S 152°50'13"E, Malaiseover creek, 13–19.vi.1998, N. Power( 1♂, DEBU), Mt Glorious, 27°19'54"S 152°45'29"E, Malaise, 24–30. xi.1998, T. Hiller( 3♀♀, DEBU), 24–30.i.1998( 1♂, DEBU), SE QLD, Brisbane, Griffith Uni., Nathan Campus, 23.x.2002, 27.33S/ 153.04E, dry&wet sclrophyll forest, ferns, B. Merz( 1♂, MHNG), N QLD, Mt. Spurgeon, wet sclera. For., 1250 m, 18–19.iv.1994, B.J. Sinclair, 16°26'S 145°12'E( 1♀, CNC), Monga State Forest, 19–24.i.1984, L. Masner( 1♀, CNC), Mt. Glorious, x.1978, A. Hiller, Malaise( 1♀, ANIC), Mt. Haig, 21kmNE by E of Atherton, 18.xi.1981, D.H. Colless, Malaise trap( 1♂, ANIC), 15.50S 145.20E, Gap Ck., 5kmESE Mt. Finnigan, 15.v.1981, D.H. Colless, Malaise trap( 1♀, ANIC), Great Sandy Nat. Pk., Cooloola Section, 25°56'49"S 153°05'30"E, 1–5.x.1996, D.K. Yeates, G. Lambkin& S. Winterton, Malaise trap( 1♂, UQIC), Noosa Natl. Pk., Palm Trail, sweeping, 30.iv.1999, S.A. Marshall( 1♂, DEBU). Tasmania: Claytons, i.1991, Nielsen& Edwards( 1♂ 1♀, ANIC). Victoria: Sherbrook Forestnear Melbourne Via., 21.i.1966, D.K. McAlpine( 1♀, AMS).
Redescription Male( Figs 1, 2, 5). Body length 2.7–5.2 mm. Anepisternal disc present. First flagellomere orbicular. Bristles black. Arista pubescent. Vibrissa relatively long and curved. Ocellar bristle thin and approximately as long as tubercle. Two dorsocentral bristles with small bristle in front of anterior dorsocentral. Gena relatively high and sometimes with inward bend at mid-height (not as distinct as in H. laticornis). Face and buccal cavity velvety and meeting at slightly rounded angle. Head yellow with posterior 2⁄ 3of frons (usually excluding lateral and posterior margins) dark brown (remainder orange) and back of head above foramen dark brown; first flagellomere sometimes with narrow infuscation along anterior margin or around base of arista; USNM specimen with distal 2⁄ 3of first flagellomere dark brown; occiput brown (venter often yellow in southern specimens) and gena yellow with posterior half sometimes brown (more frequently brown in northern specimens); clypeus brown; anterior half of frons pilose and dorsal margin of gena silvery tomentose. Thorax dark brown with proepisternum yellow and posterolateral margin of scutum to anterolateral margin of scutellum sometimes with variably yellow to orange tint (more common northwards). Legs yellow with fore coxa white, fore tarsi dark brown and laterally compressed, fore tibia brownish (lighter to base), and fore femur with long, illdefined spot on inner apex. Abdomen dark brown. M 1+2ratio 3.7–5.3. Wing with basal band confluent with second band through cells br and bm; second and third bands (confluent through first radial cell) separated evenly by thin clear strip. Halter white with base and side of stalk infuscated. Female. As described for male except as follows: pale spots on scutum faded or absent; anterior and inner-distal margins of first flagellomere infuscated, sometimes with distal 2⁄ 3of inner face dark; frons shiny or with small pilose anteromedial patch; only anterior and lateral margins of frons with orange tint; face and gena with piceous tint; fore tibia and tarsi evenly brown; terminalia yellow. Male terminalia( Figs 73–77). Length of epandrium slightly more than half width and height. Surstylus 2⁄ 3height of epandrium, very thin and pointed, with few inner-apical tubercle-like bristles and outer face bare. Cerci approximately as high as surstylus, heart-shaped and heavily setose. Hypandrium+pregonite bilobed (anterior lobe with two stout bristles and posterior lobe setulose at apex and base). Phallapodeme shorter than hypandrium. Distiphallus with membrane on basal half strongly spinulose; with small serrated process medially (vestigial, shallow, slightly longer than wide, or very long and blade-like); wide and heavily elaborated apically with serrated subapical ridge. Female terminalia(Figs 104, 105). Ventral receptacle rounded apically and flagellum very long, thin and coiled (coils tighter to apex). Spermatheca barrel-shaped (widest subapically) and long with several basal wrinkles.
Comments. Based on external and internal examination of male and female specimens of Heteromeringia spinulosaand H. pullafrom across both of their ranges, it now appears that these two taxa belong to the same species. Their colouration is nearly identical, as are the female genitalia, surstylus and most of the hypandrial complex. The absence of a serrated blade-like process on the phallus and its associated bell-shaped apical lobe ( Fig. 77) was previously considered diagnostic of H. pulla(although D.K. McAlpine (1960)noted some polymorphism), but these are actually present in a number of males, although the blade is rarely much longer than wide if present and the lobe is frequently underdeveloped ( Fig. 75). The other character previously considered diagnostic of either species was whether or not the venter of the gena was entirely ( H. spinulosa) or only partially ( H. pulla) yellow. While the presence of either character is generally indicative of whether or not the phallic blade is present, it is sometimes not, and specimens collected in and south of the Australian Capital Territoryalmost always have an entirely yellow gena (the occiput is also predominantly yellow) regardless of whether the blade is developed or not. The presence of a strong yellow patch extending from the side of the scutellum to the posterolateral corner of the notum is slightly more correlated with the presence of a well developed phallic blade, but the intensity of this patch often varies and is sometimes associated with the “opposite” genitalic state, particularly to the north. Since all of the specimens examined appear to belong to a single (albeit unusually polymorphic) species, H. pullais here treated as a junior synonym of H. spinulosaunder the first reviser principle (ICZN Article 24.2.1). Inaddition to the material listed above, there are several specimens that range from the Australian Capital Territoryto Queenslandthat vary as follows: first flagellomere entirely white; ventral margin of gena and occiput entirely yellow; notum with faint yellowish posterolateral tint; wing pattern relatively indistinct; surstylus much longer than epandrium ( Figs 78, 79); phallus branched at midpoint, with branch short, smooth and directed distally ( Fig. 80). Theelongate surstylus is the most striking feature of these males, but given the amount of genitalic variability already discussed for this species, they will be treated as H. spinulosafor the time being. Labeldata:ACT: 35.22S 148.50E, Blundells Ck., ii.1987, D.H. Colless, Malaise trap( 3♂♂, ANIC). NSW: Carrai SF, 30°54'35"S 152°16'26"E, 1090 m, E. Tasker, 11–16.i.1998, sticky trapon E. obliqua, CC-FK-018-5 ( 1♂, AMS). QLD: 18.55S 146.10E, S2 Mt. Spec, 880 m, 5.vi–3. vii.1995, M. Cermak, Fl trap, JCU (East) ( 1♂, ANIC).
1956-12-07
AMS
D. K. McAlpine
Blue Mountains
Below Govett's Leap
26
255
1
1
New South Wales
holotype
1925-02
1957-10-12
1925-02
AMS
D. K. McAlpine
Otford
26
255
2
1
1
New South Wales
paratype
1954-04-13
AMS
D. K. McAlpine
National Park
26
255
1
1
New South Wales
holotype
1954-04-13
AMS
National Park, D. K
New South Wales
26
255
2
2
New South Wales
paratype
1954-04-20
AMS
National Park, D. K
true
New South Wales
26
255
4
4
New South Wales
paratype
[962,1359,1637,1658]
1959-01-26
AMS
D. K. McAlpine
Otford
26
255
1
1
New South Wales
paratype
1980-02-20
ANIC
D. H. Colless
Malaise
Black Mtn.
26
255
2
1
1
Australian Capital Territory
1980-02-18
ANIC
D. H. Colless
true
Malaise
Black Mtn.
26
255
1
1
Australian Capital Territory
[907,1116,1738,1759]
1980-03-04
1980-03-05
1980-03-04
ANIC
26
255
1
1
Australian Capital Territory
[1123,1320,1738,1759]
1979-11-29
ANIC
26
255
1
1
Australian Capital Territory
1979-11-30
1979-12-02
1979-11-30
ANIC
26
255
1
1
Australian Capital Territory
[949,1131,1762,1783]
1980-02-04
ANIC
26
255
1
1
Australian Capital Territory
[1138,1331,1762,1783]
1980-02-14
ANIC
26
255
1
1
Australian Capital Territory
1980-02-21
ANIC
26
255
1
1
Australian Capital Territory
[907,1100,1786,1807]
1980-02-25
ANIC
26
255
1
1
Australian Capital Territory
[1107,1320,1786,1807]
1980-02-08
1980-02-10
1980-02-08
ANIC
26
255
1
1
Australian Capital Territory
1980-02-02
1980-02-03
1980-02-02
ANIC
26
255
1
1
Australian Capital Territory
[907,1109,1810,1831]
1980-11-20
ANIC
26
255
1
1
Australian Capital Territory
[1116,1330,1810,1831]
1979-11-19
ANIC
26
255
2
2
Australian Capital Territory
1980-02-28
ANIC
26
255
1
1
Australian Capital Territory
[911,1150,1834,1855]
1979-11-17
1979-11-18
1979-11-17
ANIC
26
255
1
1
Australian Capital Territory
[1161,1397,1834,1855]
1980-02-29
ANIC
26
255
2
1
1
Australian Capital Territory
1980-03-15
1980-03-17
1980-03-15
ANIC
26
255
1
1
Australian Capital Territory
[1014,1261,1858,1879]
1979-12-15
1979-12-17
1979-12-15
ANIC
26
255
2
2
Australian Capital Territory
1980-02-29
1980-03-02
1980-02-29
ANIC
26
255
2
1
1
Australian Capital Territory
[909,1463,1882,1903]
1987-12
ANIC
M. Irwin
Black Mtn.
26
255
4
2
2
Australian Capital Territory
[839,1415,1906,1927]
1987-02
ANIC
D. H. Colless
Blundells Ck.
26
255
2
2
Australian Capital Territory
1963-11-19
AMS
G. H. Hardy
Katoomba
26
255
1
1
New South Wales
1961-12-08
AMS
26
255
2
2
New South Wales
1972-09-23
AMS
D. K. McAlpine
Springwood
Sassafras Gully
26
255
1
1
New South Wales
1961-04-01
AMS
D. K. McAlpine
27
256
New England National Park
Wright's Lookout
26
255
1
1
New South Wales
1962-01-01
AMS
D. K. McAlpine
East Kurrajong
Roberts Creek
27
256
1
1
New South Wales
1975-11-20
AMS
D. K. McAlpine
Gosford
Mooney Mooney Creek
27
256
1
1
New South Wales
[355,542,1717,1738]
1975-11-25
AMS
27
256
1
1
New South Wales
1976-11-09
AMS
27
256
2
1
1
New South Wales
[314,598,1741,1762]
1981-12-11
AMS
Day
27
256
1
1
New South Wales
1978-11-16
AMS
McAlpine & Day
27
256
1
1
New South Wales
1975-11-25
AMS
McAlpine & Day
27
256
1
1
New South Wales
[225,650,1789,1810]
1977-12-06
AMS
McAlpine & Day
27
256
1
1
New South Wales
1989-12-01
AMS
McAlpine & Day
27
256
1
1
New South Wales
1961-03
AMS
D. K. McAlpine
National Park
27
256
2
1
1
New South Wales
1987-02-13
1987-02-19
1987-02-13
AMS
D. Bickel
1200
Nimmitabel
Brown Mtn.
27
256
1
1
New South Wales
1985-02-12
AMS
Royal NP & Bela Ck. & D. Bickel
27
256
1
1
New South Wales
1970-12-19
AMS
D. K. McAlpine
Royal National Park
27
256
1
1
New South Wales
1963-03-14
AMS
Casuarina & D. K. McAlpine
Careel Bay
27
256
2
1
1
New South Wales
1999-12-24
2000-01-04
1999-12-24
AMS
G. & T. Williams
Camden Head
27
256
1
1
New South Wales
1988-01-03
1998-04-30
1988-01-03
AMS
Sullivan Gap FR & E. Tasker
930
Buladelah
27
256
1
1
New South Wales
1998-01-11
1998-01-16
1998-01-11
USNM
27
256
2
2
New South Wales
1998-01-11
1998-01-16
1998-01-11
AMS
27
256
1
1
New South Wales
[856,1418,1644,1665]
1997-12-03
1997-12-08
1997-12-03
AMS
27
256
1
1
New South Wales
2000-12-16
AMS
Deua NP & D. Bickel
800
-35.758335
Wyanbene Caves Cpgd.
19
149.65833
27
256
1
1
New South Wales
1990-12-20
1990-12-27
1990-12-20
USNM
Pollock & Reichert
50
Lorien Wildlife Refuge
Taree
27
256
1
1
New South Wales
2003-12-25
DEBU
Royal Natl. Pk. & S. A. Marshal
Scientists' Cabin
27
256
2
2
New South Wales
1983-03-03
1983-03-11
1983-03-03
ANIC
J. F. Lawrence & Emg.
200
Yeola
Upp. Kangaroo Riv.
27
256
1
1
New South Wales
[1001,1240,1836,1857]
1983-03-09
ANIC
Emg.
27
256
1
1
New South Wales
1983-03-20
1983-03-30
1983-03-20
ANIC
Emg.
27
256
3
2
1
New South Wales
[955,1241,1860,1881]
1983-03-15
1983-03-20
1983-03-15
ANIC
Emg.
27
256
1
1
New South Wales
1984-01-24
CNC
L. Masner
Monga State Forest
27
256
1
1
New South Wales
[1084,1299,1884,1905]
1984-01-19
1984-01-24
1984-01-19
CNC
27
256
1
1
New South Wales
1975-03-20
AMS
McAlpine
Monto District
580
-24.566668
Austral
1250
151.48334
Bulburin
27
256
1
1
Queensland
AMS
D. K. McAlpine
Austral Forest
-24.566668
Site
1250
151.48334
Bulburin
27
256
1
1
Queensland
1996-10-01
1996-10-05
1996-10-01
UQIC
D. K. Yeates & G. Lambkin & Winterton
28
257
-25.946943
Cooloola Section
20
153.09166
Great Sandy Nat. Pk.
27
256
2
2
Queensland
1985-11-27
AMS
Cooloola NP & D. Bickel & Cassis
Freshwater Lake
28
257
1
1
Queensland
1992-11-26
AMS
Bunya Mtns NP & Pans & D. J. Bickel
960
Nanaga
At Rd.
28
257
1
1
Queensland
2002-10-24
MHNG
Merz & Foldvari & McNeil
-27.2
Rainforest Circuit
743
152.46
Mt. Glorious
28
257
2
2
Queensland
2002-10-23
MHNG
B. Merz
Brisbane
-27.33
Nathan Campus
742
153.04
Griffith Uni.
28
257
1
1
Queensland
1998-01-24
1998-01-30
1998-01-24
DEBU
Hiller
-27.331667
Malaise
20
152.75806
Mt Glorious
28
257
3
1
2
Queensland
[344,592,1984,2005]
1997-11-24
1997-11-30
1997-11-24
DEBU
28
257
1
1
Queensland
[603,906,1792,2005]
1998-01-03
1998-01-09
1998-01-03
DEBU
28
257
3
2
1
Queensland
2002-09-29
DEBU
C. R. Nelson
-27.428055
Scrub Creek Rd.
20
152.8375
Brisbane Forest
28
257
1
1
Queensland
2002-10-26
MHNG
Merz & Foldvari
Tamborine Mts.
-27.56
Palm Grove
742
153.12
Eagle Heights
28
257
1
1
Queensland
1961-12-16
AMS
D. K. McAlpine
Herberton
Crater
28
257
1
1
Queensland
1967-01-18
1970-01-14
1967-01-18
AMS, USNM
D. K. McAlpine & Holloway & G. A. Holloway
Birthday Crk
Qld
Paluma
28
257
4
4
Queensland
1980-05-08
1980-05-09
1980-05-08
ANIC
I. D. Naumann & J. C. Cardale
Eungella Nat. Pk.
Skywindow Lookout
28
257
1
1
Queensland
1987-11-30
1987-12-13
1987-11-30
ANIC
N Qld & Julatten, A
29
258
Walford-Huggins
28
257
3
1
2
Queensland
[561,745,154,175]
1987-10-05
1987-12-13
1987-11-30
ANIC
N Qld & Julatten, A
true
Walford-Huggins
29
258
1
1
Queensland
[121,745,178,199]
1989-06
1989-07-31
1989-06
ANIC
I. Naumann
-13.32
McIlwraith Range
774
143.19
29
258
1
1
Queensland
[121,501,202,223]
1924-02
UQIC
G. H. Hardy
Sirakan
29
258
1
1
Queensland
1999-04-05
1999-04-08
1999-04-05
DEBU
S. A. Marshall
Lake Eacham
Atherton
29
258
1
1
Queensland
1998-06-13
1998-06-19
1998-06-13
DEBU
N. Power
-27.418055
Malaise
20
152.83694
Brisbane Forest
29
258
1
1
Queensland
1998-11-24
1998-11-30
1998-11-24
DEBU
Hiller
-27.331667
Malaise
20
152.75806
Mt Glorious
29
258
3
3
Queensland
[421,656,298,319]
1998-01-24
1998-01-30
1998-01-24
DEBU
29
258
1
1
Queensland
2002-10-23
MHNG
B. Merz
Brisbane
-27.33
Nathan Campus
742
153.04
Griffith Uni.
29
258
1
1
Queensland
1994-04-18
1994-04-19
1994-04-18
CNC
Sinclair
1250
-16.433332
For.
1282
145.2
Mt. Spurgeon
29
258
1
1
Queensland
1984-01-19
1984-01-24
1984-01-19
CNC
L. Masner
Monga State Forest
29
258
1
1
Queensland
[185,653,418,439]
1978-10
ANIC
A. Hiller & Malaise
Mt. Glorious
29
258
1
1
Queensland
1981-11-18
ANIC
D. H. Colless
Mt. Haig
29
258
1
1
Queensland
1981-05-15
ANIC
Gap Ck. & D. H. Colless
-15.5
5 km ESE Mt. Finnigan
771
145.2
29
258
1
1
Queensland
1996-10-01
1996-10-05
1996-10-01
UQIC
D. K. Yeates & G. Lambkin & Winterton
-25.946943
Cooloola Section
20
153.09166
Great Sandy Nat. Pk.
29
258
1
1
Queensland
1999-04-30
DEBU
Noosa Natl. Pk. & S. A. Marshall
Palm Trail
29
258
1
1
Queensland
1991-01
ANIC
Nielsen & Edwards
Claytons
29
258
2
1
1
Tasmania
1966-01-21
AMS
D. K. McAlpine
Melbourne Via.
Sherbrook Forest
29
258
1
1
Victoria
1987-02
ANIC
Blundells Ck. & D. H. Colless
In
-35.22
Label
716
148.5
The
29
258
3
3
Australian Capital Territory
1998-01-11
1998-01-16
1998-01-11
AMS
Carrai SF & E. Tasker
1090
-30.909721
20
152.27388
29
258
1
1
Australian Capital Territory
1995-06-05
1995-07-03
1995-06-05
ANIC
M. Cermak
880
-18.55
Mt. Spec
765
146.1
29
258
1
1
Australian Capital Territory