Records Of Mesoamerican Anacroneuria (Plecoptera: Perlidae), With Descriptions Of Four New Species Stark, Bill P. Illiesia 2014 10 2 6 16 DC6H Stark, 2014 Stark 2014 [321,587,1506,1530] Insecta Perlidae Anacroneuria Animalia Plecoptera 8 14 Arthropoda species totumas sp. nov.  ( Figs. 16-20)   Material examined.  Holotype ♂,  Panama, Chiriqui, Totumas Cloud Forest, lodge, 08 °53.060’ N, 82 °41.068’ W,  1920 m,  19-25 June 2011, E. Fuller( USNM).  Paratypes:  Costa Rica: Puntarenas, Rio Bellavista, ca.  1.5 kmNW Las Alturas,  1400 m,  10-11 August 1990, R. Holzenthal, R. Blahnik, F. Munoz, 1♂( BPSC).  San Jose, tributary Quebrada Caraiges,  3.6 kmSW La Legua,  1650 m, R. Holzenthal, K. Kjer, F. Quesada, 2♂( BPSC).   Adult habitus.General color brown patterned with yellow-brown. Head with dark lappets, antennae and a dark band extends from behind ocelli forward to M-line; dark band expanded laterally along M-line, and interrupted by a pale interocellar spot ( Fig. 16). Pronotum dark over most of disc, but with a slender pale median band. Wings pale brown, veins brown except for C and apical half of SC. Femora pale in proximal half and dark brown in distal half; fore and mid tibiae brown, hind tibiae with pale transverse median band.  Male.Forewing length 20 mm. Hammer a low, mound-like structure ( Fig. 17). Aedeagal apex simple ( Figs. 18-19); ventroapical aspect with a large pair of membranous lobes overlying darkly sclerotized lateral margins ( Fig. 18); dorsal keel absent, lateral margins slightly convergent to truncate apex ( Fig. 20). Hooks slender.  Female.Unknown.  Larva.Unknown.   Etymology.The species name, based on the typelocality at the Mount Totumas Cloud Forest, is used as a noun in apposition.   Diagnosis.This species keys to couplet 11 in Stark (1998)where  A. acutipennis Klapálek, 1923and  A. equisita, a pair of species with trilobed aedeagal apices are the options. Because the new species has a simple aedeagal apex without lateral lobes, neither of these choices are viable. The aedeagal apex of the new species is generally similar to that of  A. varilla, a much smaller species (male FWL = 9-10 mm) known from Costa Ricaand Panama( Stark 1998). 3111908358 2011-06-19 2011-06-25 2011-06-19 USNM E. Fuller Panama 1920 8.884334 Totumas Cloud Forest 1 -82.68446 8 14 1 1 Chiriqui holotype 3111908316 1990-08-10 1990-08-11 1990-08-10 BPSC R. Holzenthal & R. Blahnik & F. Munoz Costa Rica 1400 Rio Bellavista 8 14 1 1 Puntarenas paratype 3111908352 BPSC R. Holzenthal & K. Kjer & F. Quesada Costa Rica 1650 Quebrada Caraiges 8 14 2 2 San Jose paratype