High level of endemism in Haiti’s last remaining forests: a revision of Modisimus (Araneae: Pholcidae) on Hispaniola, using morphology and molecules Huber, Bernhard A. Fischer, Nadine Astrin, Jonas J. Zoological Journal of the Linnean Society 2010 2010-01-25 158 2 244 299 43TVG HUBER & FISCHER Huber & Fischer & Astrin 2010 [844,1273,376,397] Arachnida Pholcidae Modisimus Animalia Araneae 45 289 Arthropoda species miri sp. nov.  ( Figs 61, 82, 190, 191, 201)  Type:  Male holotypefrom Reserva Científica Loma Quita Espuela, near summit ( 19°21.5 ′N, 70°09 ′W), Duarte Prov., Dominican Republic;  900 ma.s.l., underside of manacla (palm) leaves,  10 November 2005( B.A. Huber), in ZFMK( DR 23a).  Etymology:The species name honours the Dominicanpoet and writer Pedro Julio Mir Valentín (1913– 2000).  Diagnosis:Medium-sized, pale-greenish, long-legged species, with six eyes, distinguished from close relatives (  M. palvet sp. nov.and  M. berac sp. nov.) by epigynum shape ( Fig. 61) and pair of apophyses above fang joints ( Fig. 190); further distinguished from  M. palvet sp. nov.by pattern of club-shaped modified hairs on male chelicerae ( Fig. 190) and slightly wider procursus.  Male ( holotype):Total length, 1.9; carapace width, 0.80. Leg 1: 24.0 (5.8 + 0.3 + 5.8 + 9.8 + 2.3); tibia 2, 4.0; tibia 3, 2.8; tibia 4, 3.1. Tibia 1 L/d: 72. Habitus similar to  M. palvet sp. nov.(cf. Fig. 35), carapace pale ochre-yellow to whitish, clypeus with wide brown distal margin, sternum whitish; legs pale ochre-yellow, with darker patella area and tibia–metatarsus joints; abdomen pale greenish grey with purplish marks dorsally at heart area, monochromous ventrally. Ocular area slightly elevated; thoracic furrow distinct but shallow. PME–PME, 95 Mm; PME diameter, 70 Mm; PME–ALE, 80 Mm; no AME. Sternum slightly wider than long (0.60/0.55), unmodified. Chelicerae with about ten club-shaped modified hairs on each side, and distinctive apophyses above fang joints ( Fig. 190). Palps very similar to  M. palvet sp. nov.(cf. Fig. 185), procursus slightly wider, and bulbal apophysis slightly wider and shorter. Long spines on femora 1 (three or four) and femora 2 (two), many short vertical hairs on all femora; without curved hairs; retrolateral trichobothrium on tibia 1 at 7%; prolateral trichobothrium missing on tibia 1, present on all other tibiae; tarsus 1 with ~30 pseudosegments.  Variation:Other male with entire clypeus and anterior side of ocular area brown. Tibia 1: 5.4.   Figures 190–194.Diagnostic characters. 190, 191.   Modisimus miri sp. nov.190. Male chelicerae, frontal view. 191. Cleared epigynum, dorsal view. 192–194.   Modisimus paraiso sp. nov.192. Left palp, retrolateral view. 193. Male chelicerae, frontal view. 194. Cleared epigynum, dorsal view. Scale bars: 0.2 mm.   Figure 195.Known distributions of representatives of the  femoratusgroup, part 1 (see also Fig. 196). The question marks indicate specimens that are assigned tentatively to  Modisimus femoratus.   Figure 196.Known distributions of representatives of the  femoratusgroup, part 2 (see also Fig. 195).   Figure 197.Known distributions of  Modisimus vittatusand  Modisimus makandal sp. nov.   Figure 198.Known distributions of representatives of the  fuscusgroup.  Female:In general similar to male, without darkened clypeus. Tibia 1: 4.2. Epigynum, simple externally ( Fig. 61); dorsal view as in Figures 82, 191.  Distribution:Known from typelocality only ( Fig. 201).  Material examined:  Dominican Republic: Duarte Prov., Reserva Científica Loma Quita Espuela, 1♂, holotype above; same data, 1♂, 1♀and one juvenile (ZFMK, DR 23); same data, one juvenile, in pure ethanol ( ZFMK, DR 100-24). 2005-11-10 ZFMK B. A. Huber Dominican Republic 900 19.358334 Male 878 -70.15 Reserva Cientifica Loma Quita Espuela 45 289 DR 23 1 Duarte holotype Dominican Republic Reserva Cientifica Loma Quita Espuela 47 291 1 1 Duarte holotype [250,759,1880,1903] Dominican Republic Reserva Cientifica Loma Quita Espuela 47 291 3 1 1 1 Duarte Prov. holotype ZFMK 47 291 DR 100-24 1 1