New genera and species of Selenomorphini (Hemiptera: Cicadellidae: Iassinae), including redescription of Scaroidana Osborn, Pachyopsis Uhler and updated key to genera and species Domahovski, Alexandre Cruz Zootaxa 2019 2019-12-17 4711 3 517 544 8JXBZ Domahovski, 2019 Domahovski 2019 [151,390,1376,1403] Insecta Cicadellidae Pachyopsis Animalia Hemiptera 14 531 Arthropoda species pallidus sp. nov.  ( Figs 64−82)   Diagnosis.Pygofer (Fig. 68), with ventral process short, curved distally, reaching the apical third of pygofer; subgenital plate (Fig. 68) in lateral view, very long, slightly surpassing the pygofer apex, broadened apically with macrosetae only near apex, grouped in line; style (Fig. 72) with blade very long, length from lateral lobe to apex two times longer than length from lateral lobe to base.     Typelocality. São José dos Pinhais, Paraná, Southern Brazil.   Measurements(mm). Length 5.5−6.2 (male), 6.5−6.9 (female).  Coloration.Male and female: body ( Figs 64−66, 74−76) uniform pale yellow or yellowish-green (pale green in life), without markings. Eye and ocellus red.  Description.Head (Figs 66, 76), in ventral view, frons surface slightly rugose; maxillary plate produced ventrally as far as clypeus apex in both sexes. Forewing ( Figs 64, 65, 74, 75) translucent, with veins not elevated, veinlets inconspicuous, few and rare extra cross veins. Metatibia PD, AD and AV rows with 14−16, 10−11, 9−11 macrosetae respectively. Abdominal sternite II (Fig. 67) with internal apodemes narrow, long, slightly expanded apically and rounded. Other characteristics as in generic description.  Male terminalia.Pygofer (Fig. 68), with posteroventral margin slightly rounded; apex slightly tapered and rounded; ventral process short, curved distally, reaching the apical third of pygofer. Subgenital plate (Fig. 68) in lateral view, slightly surpassing the pygofer apex, broadened apically; in ventral view (Fig. 69) 5.2 times longer than wide; macrosetae only near apex, grouped in line. Connective (Fig. 70) stem with lateral margins strongly divergent apically; apex straight. Style (Fig. 72) with blade slightly curved dorsally, slightly constricted on mid-length and broadened apically; length from lateral lobe to apex two times longer than length from lateral lobe to base. Aedeagus (Fig. 73) with preatrium produced anterad; atrium wide; shaft apex tapered and rounded. Other characteristics as in generic description.   FIGURES 64─73.  Pachyopsis pallidus  sp. nov., male holotype. 64, habitus, dorsal view. 65, habitus, lateral view. 66, head, ventral view. 67, Abdominal sternites I & II and internal apodemes, ventral view. 68–73, male terminalia: 68, pygofer, valve, and subgenital plate, lateral view. 69, subgenital plate, ventral view. 70, connective, dorsal view. 71, style, dorsal view. 72, style, lateral view. 73, aedeagus, lateral view. Scale bars in mm.   FIGURES 74─82.  Pachyopsis pallidus  sp. nov., female paratype. 74, habitus, dorsal view. 75, habitus, lateral view. 76, head, ventral view. 77, pygofer, lateral view. 78, first valvifer and first valvula, lateral view. 79, apex of first valvula, lateral view. 80, second valvula, lateral view. 81, apex of second valvula, lateral view. 82, gonoplac, lateral view. Scale bars in mm.  Female terminalia.Sternite VII with posterior margin nearly straight. Pygofer (Fig. 77) 1.7 times longer than maximum height; apex subacute. First valvifer (Fig. 78) 1.1 times higher than long; posterior margin broadly round- ed. First valvula (Fig. 78) slightly higher near base than near apex, ca. 10 times longer than maximum height; dorsal sculpturing starting at mid-length; apical portion (Fig. 79) with ventral sculpture present only near apex, continuous with the dorsal sculpture, apex tapered and acute. Second valvula (Fig. 80) slightly higher apically than basally, ca. 10 times longer than high; apex (Fig. 81) with 3 subacute teeth. Second valvifer ca. three times higher than long. Gonoplac (Fig. 82) 5.2 times longer than high; dorsoapical margin long; apex tapered, subacute. Other characteristics as in generic description.   Material examined.   Holotypemale:“ Brasil, Paraná, S. [ São] J. [ José] dos\ Pinhais, 25º36’18”S\ 49º11’37”W  880m\  18.XI.2016Sweep\ A.C. Domahovskileg” ( DZUP).   Paratypes: 2♀: same data except  30.XI.2011( DZUP);   1♂: same data except  07.XII.2011( MZSP);   2♀: same data except  21.XII.2011( MZSP);   1♀: same data except  22.I.2012( DZUP);   1♀: same data except  20.II.2012( DZRJ);   1♀: same data except  27.XI.2013( DZUP);   1♀: same data except  12.III.2016( DZUP);   1♀: same data except  19.III.2016( DZUP);   1♀: same data except  01−30.X.2016( DZUP);   3♂,  12♀: same data except  17−31.XII.2016( DZUP);   1♀: same data except  07−21.I.2017( DZUP);   2♀: same data except  06.XII.2017( DZUP);   1♀: same data except  24.I.2018( DZUP);   3♂: same data except  03−17. XI.2018( DZUP);   2♂,  3♀: same data except  05−15.XII.2018( 1♂, 1♀: DZRJ; 1♂, 2♀: INHS);   1♀: same data except  01-28.II.2019 Malaise( DZUP);   1♂: “S. [ São] JoséPinhais—PR [ Paraná]\ Ser.[ Serrado] Mar Br277 Km54\ Brasil  29.XI.1986\ Lev. [Levantamento] Ent. [Entomológico] PROFAUPAR\ Lâmpada” ( DZUP);   3♂,  1♀: same data except  1.I.87( DZUP);   2♂: “S. [ São] JoséPinhais—PR [ Paraná]\ Brasil( Br277− Km54)\  30.VII−06.VIII.1984\ C.I.I.F. (Luminosa)” ( 1♂: DZRJ; 1♂: INHS);   1♂: same data except  16−23.VII.1984( DZUP);   2♂: same data except  01−08.X.1984( DZUP);   1♂: “ Brasil, Paraná, Tibagi,\ P. [Parque] E. [Estadual] do Guartelá,  1000m\ 24º33’47”S 50º15’26”W\  21−24.XI.2016Sweep\ A. C. Domahovskileg.” ( DZUP);   1♂: “ Brasil, Paraná, Piraquara, Ma-\ nanciais da Serra, 25º29’47”S\ 48º58’54”W  1021m, Sweep,\  21−28.XI.2018, A.P. Pinto, B. R.\ Araujo & A.C. Doma- hovski” ( DZUP);   2♂: “Campo Largo—PR [ Paraná] \ Est. do Cerne, km 45.5\ Brasil  II/1999\ Chiamolera, L.col.” ( DZUP);   1♂: “Ponta Grossa ( V. [ Vila] Velha) PR[ Paraná]\ Reserva IAPARBr376\ Brasil  18.I.88\ Lev. [Levantamento] Ent. [Entomológico] PROFAUPAR\ Lâmpada” ( DZUP);   1♂: Brasil, Minas Gerais,\ Extrema, Morro do Lopo, 22.881º S 46.308ºW\  1520m,  09.xii.2012, G. Melo\ & P. Grossi, arm. Luminosa” ( DZUP);   2♂,  2♀: “Encru- zilhada—BA [ Bahia]\ Brasil  IX−1975\ M. Alvarenga Leg” ( DZUP).   Etymology.The new species name comes from the Latin word “  pallidus” that means pale. It refers to the general coloration pale-yellow (pale-green in life).  Notes.  P. pallidus  sp. nov.is similar to  P. similisKramerand  P. wappesiFreytagin having the ventral process of the pygofer short and the aedeagus with ventral margin rounded. However, the new species can be easily separated by the uniform pale green or pale yellow coloration (yellowish-brown to pale brown with forewing often darker than head and thorax in  P. similis; and yellowish-brown in  P. wappesi); the pygofer with apex rounded (acute in  P. similis); the style with blade comparatively longer and slightly curved dorsally (straight in  P. wappesi); and the aedeagus with apex comparatively narrower. 2521262937 [199,883,1628,1655] Sao Jose dos Pinhais Brazil Parana 14 531 1 Parana holotype 2521262871 2016-11-18 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 Parana holotype 2521262878 [820,1432,476,503] 2011-11-30 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 2 2 Parana paratype 2521262938 [151,654,512,539] 2011-12-07 MZSP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262874 [668,1171,512,539] 2011-12-21 MZSP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 2 2 Parana paratype 2521262880 2012-01-22 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262863 [371,831,548,575] 2012-02-20 DZRJ A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262884 [842,1318,548,575] 2013-11-27 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262868 2016-03-12 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262934 [528,1010,584,611] 2016-03-19 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262862 2016-10-01 2016-10-30 2016-10-01 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262881 [259,861,620,647] 2016-12-17 2016-12-31 2016-12-17 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 15 12 3 Parana paratype 2521262885 [873,1381,620,647] 2017-01-07 2017-01-21 2017-01-07 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262883 2017-12-06 2017-01-21 2017-01-07 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 2 2 Parana paratype 2521262879 [613,1085,656,683] 2018-01-24 2017-01-21 2017-01-07 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262922 2018-11-03 2018-11-17 2018-11-03 DZUP A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 3 3 Parana paratype 2521262926 [359,1218,692,719] 2018-12-05 2018-12-15 2018-12-05 DZRJ, INHS A. C. Domahovski Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 10 6 4 Parana paratype 2521262908 2019-02-01 2019-02-28 2019-02-01 DZUP Malaise Brazil Sao 880 -25.605 Pinhais 21 -49.19361 Jose 17 534 1 1 Parana paratype 2521262861 1986-11-29 DZUP Brazil Km Sao Mar Br 17 534 1 1 Parana paratype 2521262865 1987-01-01 DZUP Brazil Km Sao Mar Br 17 534 4 1 3 Parana paratype 2521262873 1984-07-30 1984-08-06 1984-07-30 DZRJ, INHS Brazil Km Sao Br 17 534 4 4 Parana paratype 2521262864 [661,1188,836,863] 1984-07-16 1984-07-23 1984-07-16 DZUP Brazil Km Sao Br 17 534 1 1 Parana paratype 2521262876 1984-10-01 1984-10-08 1984-10-01 DZUP Brazil Km Sao Br 17 534 2 2 Parana paratype 2571180948 2016-11-21 DZUP Brazil -24.563055 -50.25722 17 534 Parana 2571180947 2018-11-21 R Brazil -25.49639 -48.981667 17 534 Parana 2571180950 1999-02 DZUP Brazil 17 534 Parana 2571180949 1988-01-18 V Brazil 17 534 Pr 2571180951 2012-12-09 DZUP Brazil -22.881 -46.308 17 534 Minas Gerais 2521262901 1975-09 DZUP M. Alvarenga Leg Brazil Bahia 17 534 4 2 2 Bahia paratype