Cicada guttulosa Walker 1858b: 20 The cicadas (Hemiptera: Cicadidae) of Ecuador including the description of five new species, a new subtribe, four new synonymies, and fifteen new records Sanborn, Allen F. Zootaxa 2020 2020-11-17 4880 1 1 80 4ML9M (Walker, 1858) Walker 1858 [151,556,1770,1797] Insecta Cicadidae Proarna Animalia Hemiptera 22 23 Arthropoda species guttulosa      Cicada guttulosa Walker 1858b: 20. (unknown locality)  REMARKS.—This is the largest of the Ecuadorian  Proarnaspecies with some females reaching body lengths of 31 mmor more. The costal margin is unique for the Ecuadorian species of  Proarnain that it is curved from the base rather than being straight. The large body size combined with the curved costal margin makes the species fairly easy to distinguish from other Ecuadorian species of  Proarna. The lack of infuscation on the nodal line of the fore wing, the more elongate body, and the narrower pronotum are also useful in distinguishing it from  P. griseaand the small size and almost complete macular infuscation on the fore wing veins distinguish  P. insignisfrom this species.  DISTRIBUTION.—The species has been reported from Colombia, Ecuador, Peru, and South America ( Metcalf 1963a; Sanborn 2013; 2020b) with recent records from Bolivia( Sanborn 2019b). Goding (1925)reported the species from Macas, Morona Santiago province.   MATERIAL EXAMINED.— “ ECUADOR: Napo/ Prov.25km.e. Puerto/ Napo,   450m.  ix–1997// B.&B. Valentincoll. // Jatun Sacha / Bio/ Reserve / u-v light // Valentine / Coll’n / Rec. 2016” one male and two females ( FSCA); “ ECUADOR: Santo/ Domingode los / Colorados, / Hotel Tinlandia/  5–15 May 1988// Coll. Steve Passoa” one female ( FSCA); “ ECUADOR: Prov. Napo/ vic. Puerto Misahualli, / 1650– 1900’ ft., 6–19–IX– / 1998 J.E. Eger, coll. // 1º 2’ 4.2” Slat. / 77º 39’ 49.2” Wlon. /Mercury vapor & / Ultraviolet lights” two males ( AFSC); “ Ecuador, Orellana, Estacion/ Cientifica Yasuni; to UV/ & merc. vap. lights;  15–16Oct.2003/ D. Robacker, W. Warfield& M.H. Evans” one female ( AFSC); “ ECUADOR: Napo Prov./ Estación Cientifica Yasuní/ 00º 40’28” S, 76º 38’50” W/  IX–5–10–1999, UVlight / Coll. E.G. Riley,   215 m.” three males and three females ( AFSC); “ ECUADOR: Napo Province/ Misahualli Jungle Lodge area, jct. of / RioNapo & Rio Misahualli, 1650- / 1900’ elev. S 1º 2’ 4.2”, W 77º 39’/ 49.2”, 5–12:IX:1998. C. & K. / Messenger” seven males and seven females ( UNSM), two males and two females ( AFSC); “ Rio Palenque, ECUADOR/ Los Rios Prov.- D. Dodson/  XI–15–1972” three females ( AFSC); “ Ecuador, Orellana, Estacion/ Cientifica Yasuni; to UV/ & merc. vap. lights;  17–30Oct.2003/ D. Robacker, W. Warfield& M.H. Evans” one female ( AFSC); “ Ecuador, Orellana, Estacion/ Cientifica Yasuni; to UV/ & merc. vap. lights;  13–14Oct.2003/ D. Robacker, W. Warfield& M.H. Evans” one male ( AFSC); “2000 / Agua Santa, Ecuador/ Leg Bug Manic” one female ( AFSC); “ Ecuador/ Tinlandia/  4 MAR1981” one female ( BSNS), one male ( AFSC); “ Ecuador/ Tinlandia/  3 MAR1981” three males and ten females ( BSNS), one male and two females ( AFSC); “ Ecuador/ Tinlandia” two females ( BSNS); “ ECUADOR: Pichincha/ Prov., Tinlandia, E. / Alluriquin,  4–8–V–1976/   700 m.elev. // Netted / T. E. Rogers” six females ( AFSC); “ ECUADOR: Pichincha/ Prov., Tinlandia, E. / Alluriquin,  26–28–V–1976/   700 m.elev. // Netted / T. E. Rogers” one male and one female ( AFSC); “ ECUADOR: Pichincha, / San Domingode / los Colorados, / Tinlandia/  4–V–1976, at light / G.B. Fairchild” two females ( AFSC); “ ECUA-DOR: Pichincha/ Province, Tinlandia, / at light /  27–28–V–1976/ G.B. Fairchild” two females ( AFSC); “ ECUA-DOR: Pichincha/ Province, Tinlandia, / at light /  6–V–1976/ G.B. Fairchild” two females ( AFSC); “ ECUADOR: Pichincha Prov./ Tinlandia; 12 km. E. Sto./ Domingode los Colorados./ ca. 2500 ft.,  11–17–V–1986. / J. E. Eger, coll.” one female ( AFSC); “ ECUADOR/ Santo Domingode los / Colorados, Pichincha/  6 February 1958/ R. W. Hodges, 2000’” two females ( AFSC); “ ECUADOR: Pichincha Prov./ Tinlandia; 12 km. E. Sto./ Domingode los Colorados./  11–17–V–1986. G.B. Edwards / forest, field, 2200–2500’” one female ( AFSC); “ ECUADOR: Pich. / nr Sta Domingo / 7–14 May ’88, 2000’ / Hanson, Bohart” two females ( AFSC); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 12, 1985/ Clausen & Cervenka” one female ( AFSC); “ ECUADOR: Oriente/ 00º24’S, 76º 36’W/ Limoncocha / XII.67 / CW & ME Rettenmeyer” one female ( AFSC); “ ECUADOR: Napo Province, / Limoncocha, on RioNapo /  28 – II – 1974/ Boyce A. Drummond, III / Blacklight trap  300 m” one male and three females ( AFSC); “ ECUADOR: Napo Province, / Limoncocha, on RioNapo /  13 – II – 1974/ Boyce A. Drummond, III / Blacklight trap  300 m” one female ( AFSC); “ ECUADOR/ Prov. Napo/ Limon Cocha/  16–31 Aug. 1964/ Bowerman & Dozier” one female ( AFSC); “ ECUADOR/ Prov. Napo/ Limon Cocha/ G. Dozier/  5 Sept. 1964” one female ( AUEM); “ ECUADOR: Tinlandia/ 8 mi.e. Santo/ Domingo2000’ /  5–12. V.1990// N.J. Smith / R. Kassabian / J. Halstead“ one male and one female ( AFSC); “ ECUADOR: Pichincha/ Prov., Tinlandia / El. 3400’ // Charles M. Stevens / XXIV–VI–1983” one female ( AFSC); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 14, 1985/ Clausen & Cervenka” four females ( UMSP); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 16, 1985/ Clausen & Cervenka” two females ( UMSP); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 12, 1985/ Clausen & Cervenka” nine females ( UMSP), one female ( AFSC); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 13, 1985/ Clausen & Cervenka” six females ( UMSP); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 21, 1985/ Clausen & Cervenka” one female ( UMSP); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 23, 1985/ Clausen & Cervenka” three females ( UMSP); “ Hotel Tinlandia, 12 km. E. of Santo/ Domingode los / Colorados, Ecuador/  May 11, 1985/ Clausen & Cervenka” six females ( UMSP); “ ECUADOR Napo Prov/ Misahualli~  350 m/  7 October 2001/ Mark Churchillcoll.” two females ( MSUC). 4161951376 1997-09 FSCA B. & B. Valentin Ecuador 450 Puerto Prov. 23 24 3 Napo 3044196557 1988-05-05 1988-05-15 1988-05-05 FSCA Coll. Steve Passoa Ecuador Santo Colorados Domingo 23 24 1 3044196556 AFSC Ecuador Prov. -1.0345 Eger 1 -77.663666 Puerto Misahualli 23 24 1 Napo 3044196649 2003-10-15 2003-10-16 2003-10-15 UV, AFSC D. Robacker & W. Warfield Ecuador Cientifica Yasuni 23 24 1 3044196413 1999-09-05 1999-09-10 1999-09-05 UV, AFSC Coll. E. G. Riley Ecuador 215 -0.67444444 Estacion Cientifica Yasuni 21 -76.64722 23 24 215 Napo 4161951401 UNSM Ecuador Misahualli Jungle Lodge area -1.0345 Rio Misahualli 925 -77.65 Rio 23 24 1 Napo 3044196347 [351,1432,726,753] 1972-11-15 AFSC D. Dodson Ecuador Rio Palenque 23 24 1 Los Rios 3044196385 2003-10-17 2003-10-30 2003-10-17 UV, AFSC D. Robacker & W. Warfield Ecuador Cientifica Yasuni 23 24 1 3044196382 2003-10-13 2003-10-14 2003-10-13 UV, AFSC D. Robacker & W. Warfield Ecuador Cientifica Yasuni 23 24 1 3044196361 AFSC Ecuador Ecuador 23 24 1 4161951448 [579,1436,871,897] 1981-03-04 MAR, BSNS Ecuador Tinlandia 23 24 1 4161951462 [151,1378,907,933] 1981-03-03 MAR, BSNS Ecuador Tinlandia 23 24 1 3044196471 BSNS Ecuador Ecuador 23 24 1 3044196337 1976-05-04 1976-05-08 1976-05-04 T, AFSC E. Rogers Ecuador 700 Alluriquin Prov. 23 24 700 Pichincha 3044196482 1976-05-26 1976-05-28 1976-05-26 T, AFSC E. Rogers Ecuador 700 Alluriquin Prov. 23 24 700 Pichincha 3044196366 1976-05-04 1976-05-28 1976-05-04 AFSC, ECUA- G. B. Fairchild Ecuador San Domingo Tinlandia Colorados 23 24 1 Pichincha 3044196622 1986-05-11 1986-05-17 1986-05-11 AFSC E. Sto. Ecuador Tinlandia 762 Colorados. Domingo 23 24 1 Pichincha 3044196636 1958-02-06 R, AFSC W. Hodges Ecuador Colorados Santo Domingo 23 24 1 Pichincha 3044196642 1986-05-11 1986-05-17 1986-05-11 AFSC E. Sto. Ecuador Tinlandia Colorados. Domingo 23 24 1 Pichincha 3044196464 AFSC Ecuador ECUADOR 23 24 1 3044196660 1985-05-12 AFSC Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196468 AFSC Ecuador -0.4 Oriente 1308 -76.6 23 24 1 3044196521 1974-02-28 AFSC Ecuador 300 Rio Limoncocha 23 24 1 Napo 3044196503 1974-02-13 AFSC Ecuador 300 Rio Limoncocha 23 24 1 Napo 3044196423 1964-08-16 1964-08-31 1964-08-16 AFSC Ecuador Limon Cocha Prov. 23 24 1 Napo 3044196386 1964-09-05 AUEM Ecuador Prov. Dozier Limon Cocha 23 24 1 Napo 3044196329 1990-05-05 1990-05-12 1990-05-05 V, R, AFSC Ecuador Tinlandia Domingo Santo 23 24 1 3044196323 AFSC Ecuador Pichincha 23 24 1 Pichincha 3044196547 1985-05-14 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196411 1985-05-16 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 4161951465 1985-05-12 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196501 1985-05-13 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196607 1985-05-21 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196346 1985-05-23 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196545 1985-05-11 UMSP Ecuador Hotel Tinlandia Domingo Santo 23 24 1 3044196604 2001-10-07 MSUC Mark Churchill Ecuador 350 Misahualli Prov 23 24 1 Napo