The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae), part 3: species of the Pilbara bioregion of Western Australia Car, Catherine A. Harvey, Mark S. Hillyer, Mia J. Huey, Joel A. Zootaxa 2019 2019-06-17 4617 1 1 71 Car Car & Harvey & Hillyer & Huey 2019 [151,454,1853,1880] Diplopoda Paradoxosomatidae Antichiropus Animalia Polydesmida 60 61 Arthropoda species uvulus sp. nov.   ZooBank LSID: urn:lsid:zoobank.org:act: 4D3B4E88-272E-48F9-B33D-3AC6652552C9   Type material examined.   Australia:  Western Australia: holotypemale, Barlee Range Nature Reserve, site BR4, 23.04.46S, 115.47.27E,   15–18 June 1994, dry pitfall trap P.G. Kendrickand G.W. Kendrick( WAMT71872, Gen- Bankaccession number 12S,  MK735740; 28S,  MK735804; COI,  MK735848; COIII,  MK735926). Paratype: 1 male, Highway Tod Bore( West Robe), ca. 50 kmW. of Pannawonica, TOD04, 21°42’04.6”S, 115°50’58.0”E, 18 May 2015–   31 May 2015, targeted search, stony plain, M. Quinn( WAM T137144, GenBankaccession number 12S,  MK735771; COI,  MK735893; COIII,  MK735950; CytB,  MK735998).  Other material examined.   Australia:  Western Australia: 1 male, Pannawonica Rd, 21°38’50”S, 116°06’26”E, Biotasite SNZ08,  29 April 2005, R. Tealeand Z. Hamilton( WAM T66480, GenBankaccession number 12S,  MK735738; 28S,  MK735802; COI,  MK735845; COIII,  MK735924);  1 female, Pannawonica Rd, 21°35’34”S, 115°56’13”E,  1 May 2005, R. Tealeand Z. Hamilton( WAM T66481, GenBankaccession number COI,  MK735846);  1 male, Barlee Range Nature Reserve, site BR6, 23°23’21”S, 115°53’12”E,  11–4 June 1994, dry pitfall trap, P.G. Kendrickand G.W. Kendrick( WAMT71871);  1 male, 1 female(damaged), Barlee Range Nature Reserve, quadrat 6, 23°23’21”S, 115°53’12”E,  June 1994, wet pitfall trap, S. van Leeuwenand B. Bromilow( WAMT71873);  1 male, 9.5 kmS. of Mt Minnie, Pilbara Biological Surveysite WYW04, 22°11’19.1”S, 115°33’13.2”E,  May 2004, CALMstaff ( Pilbara Biological Survey) ( WAM T124604);  1 male, Mesa B C( West Robe), ca. 50 kmW. of Pannawonica, RVM02, 21°39’37”S, 115°56’54.9”E,  18 May 2015–  31 May 2015, targeted searches, M. Quinn( WAM T137140); 1 malecovered in fungus,  1 female(damaged), TOD01, Highway Tod Bore( West Robe), ca. 50 kmW. of Pannawonica, 21°41’37.0”S, 115°49’55.7”E,  18 May–31 May 2015, targeted search, stony plain, M. Quinn( WAM T137141);  1 male, TOD01, Highway Tod Bore( West Robe), ca. 50 kmW. of Pannawonica, 21°41’37.0”S, 115°49’55.7”E,  18 May–31 May 2015, targeted search, stony plain, M. Quinn( WAM T137143;  1 male, TOD04, Highway Tod Bore( West Robe), ca. 50 kmW. of Pannawonica, 21°42’04.6”S, 115°50’58.0”E,  18 May–31 May 2015, targeted search, stony plain, M. Quinn( WAM T137146);  3 males, 3 females(damaged), 9.5 kmS. of Mt Minnie, Pilbara Biological Surveysite WYW04, 22°11’19.1”S, 115°33’13.2”E,  May 2004, CALMstaff ( Pilbara Biological Survey) ( WAM T146727).   Diagnosis.Gonopod:  Antichiropus uvulusCar,  n. sp.is most similar in appearance to  A. georginaeCar,  n. sp.( Fig 17 )and both lack a gonopodal prolongation of the femorite. The latter has a much more robust main femoral process and a small, distinctive femoral process at the base of the main femoral process, lacking in  A. uvulus.   Description. Male holotype: Bodyca. 15 mmlong; midbody ring ca. 2 mmwide, with distinct, smooth waist, prozonite and metazonite of similar widths.  Colour(in alcohol) dark chestnut brown dorsally, cream ventrally ( Fig 38A); leg colour pale brown. No paranota ( Fig 38B).  Sterniteswithout obvious processes/tubercles, sternal lamella broad and relatively square. Leg coxal processes absent. Anterior spiracles at midbody ovoid, flat.  Headsmooth, with no sculpturing; frons smooth, with few setae; face narrow, maximum width ca. 4xthe distance between antennal sockets; sockets separated by ca. 1.5x width of socket.  Antennaeof moderate length, reaching to ring 2, antennomeres all similar shape, apical antennomere slightly stouter.  Collum0.75x length of head, reaching sternites (in lateral view) ( Fig 38A).  Gonopodof medium length, reaching posterior edge of ring 5; coxa (C) much shorter and stouter than femorite (F) with very slight ridge on anterior surface (not shown in Fig 38); prefemur (PF) much shorter and stouter than F, with pronounced prefemoral lip; femorite 2/3 length of acropodite, upright, small bulbous process at base; main femoral process (MFP) asymmetrically pointed, 1/3 solenomere length, second femoral process (fp1) present as an irregular saw-like process at the base of the MFP; prolongation of femorite (prof) absent, represented only by a slight rounding of femorite as it merges into solenomere; solenomere (S) relatively short, appearing L-shaped in anterior view, thickest at base, narrowing to apex; base with semi-transparent flange along one edge; solenomere tip, slender, rounded; solenomere process 1 (sp1) tiny, slender, pointed at solenomere tip; solenomere process 2 (sp2) relatively large, triangular, pointed, forming pincer-like structure with tip ( Figs 38C–F).  Female:Similar width to male, but slightly stouter, legs slightly shorter and more slender (WAM T124604).   Distribution.This species, like  A. anguinus, has been found in the Barlee Range National Park and in Pannawonica, about 200 kmSW. of the town of Karratha ( Fig 41).   Etymology.This species is named for the position of the first solenomere process (sp1) which has the appearance of a mammal’s uvula (Latin, noun, uvula, fleshy protuberance on the roof of the mouth). 2412528401 [455,1432,1996,2024] BR Australia Barlee Range Nature Reserve 60 61 BR4 1 Western Australia holotype 2412528392 1994-06-15 1994-06-18 1994-06-15 WAM, MK, COI, COIII P. G. Kendrick & G. W. Kendrick & Bank Pannawonica -21.701279 Highway Tod Bore 1 115.84945 West Robe 61 62 1 1 paratype 2412528395 2015-05-31 WAM, MK, COI, COIII M. Quinn & GenBank 61 62 T137144 1 paratype 2412528411 2005-04-29 WAM, MK, COI, COIII Biota & Teale & Hamilton & GenBank Australia -21.647223 Pannawonica Rd 21 116.10722 61 62 1 1 Western Australia 2412528351 2005-05-01 WAM, COI, MK Teale & Hamilton & GenBank Australia -21.59278 Pannawonica Rd 21 115.93694 61 62 1 1 Western Australia 2412528374 1994-06-04 1994-06-11 1994-06-04 WAM P. G. Kendrick & G. W. Kendrick Australia -23.389166 Barlee Range Nature Reserve 21 115.886665 61 62 BR6 1 1 Western Australia 2412528420 1994-06 WAM S. van Leeuwen & B. Bromilow Australia -23.389166 Barlee Range Nature Reserve 21 115.886665 61 62 2 1 1 Western Australia 2412528376 2004-05 CALM, WAM Pilbara Biological Survey Australia -22.188639 Pilbara Biological Survey 1 115.55367 Mt Minnie 61 62 T124604 1 1 Western Australia 2412528354 2015-05-18 2015-05-31 2015-05-18 WAM Mesa B C & Quinn Australia -21.660278 Pannawonica 15 115.948586 West Robe 61 62 T137140 1 1 Western Australia 2412528388 2015-05-18 2015-05-31 2015-05-18 WAM Quinn Australia Highway Tod Bore -21.693611 Pannawonica 1 115.83214 West Robe 61 62 T137141 1 1 Western Australia 2412528383 2015-05-18 2015-05-31 2015-05-18 WAM Quinn Australia Highway Tod Bore -21.693611 Pannawonica 1 115.83214 West Robe 61 62 T137143 1 1 Western Australia 2412528360 2015-05-18 2015-05-31 2015-05-18 WAM Quinn Australia Highway Tod Bore -21.701279 Pannawonica 1 115.84945 West Robe 61 62 T137146 1 1 Western Australia 2412528355 2004-05 CALM, WAM Pilbara Biological Survey Australia -22.188639 Pilbara Biological Survey 1 115.55367 Mt Minnie 61 62 T146727 6 3 3 Western Australia