Taxonomic revision of the Australian Notoxinae (Coleoptera: Anthicidae) Kejval, Zbyněk Acta Entomologica Musei Nationalis Pragae 2013 suppl. 2013-11-15 53 1 98 5MFN 1272042 LaFerte-Senectere 1849 [402,571,219,245] Insecta Anthicidae Mecynotarsus Animalia Coleoptera 8 9 Arthropoda genus    1 (2) Elytra in female distinctly excavate apically ( Fig. 134); male protibiae with distinct pointed process subapically on inner margin ( Fig. 137); head vertex in male with sharp median incision ( Fig. 130); setation of elytra as in Fig. 174; male abdominal character as in Figs 44–46; species occurring in New Caledonia. ............................ ...............................................................................................   M. excavatus sp. nov.   2 (1) Elytra simple, rather conjointly rounded apically; male protibiae simple; head vertex in male at most shallowly impressed medially, lacking any incision; species occurring in Australia.   3 (4) Elytral disc uneven, with distinct omoplates and postbasal impression; elytral setation bicoloured and unevenly ordered, mostly brownish, with whitish setae pointing mostly laterad and forming two conspicuous transverse bands ( Fig. 203); elytral setae hairy and finely bifurcate apically; male characters as in Figs 76–79. ............................... ....................................................................................  M. mastersii MacLeay, 1872   4 (3) Elytral disc evenly curved, lacking omoplates and postbasal impression; elytral setation mostly scaly and simple, if bifurcate, then pointing evenly posteriad, forming no transverse bands.   5 (20) Pronotal disc granulate, with numerous, more or less distinct granules scattered dorsally and dorso-laterally (laterally sparser and coarser; Figs 152, 166).   6 (7) Elytral setae finely bifurcate apically ( Fig. 184); antennae rather short; male characters as in Figs 110, 111. ................................................................   M. speciosus sp. nov.   7 (6) Elytral setae simple, at most truncate apically; antennae conspicuously long.   8 (9) Antero-lateral margins of frons projecting into conspicuous, apically pointed, thornlike process ( Fig. 23); male characters as in Figs 24, 25. .........   M. armifer sp. nov.   9 (8) Antero-lateral margins of frons at most lobed, never with pointed process.   10 (13) Elytral setation longer and somewhat sparser, surface visible ( Fig. 183).   11 (12) Elytra with numerous long tactile setae, including apical half; appressed setae of pronotal disc very fine, nearly frayed apically; distal antennomeres more robust, wider, antennomere X 1.8 times as long as wide; male characters as in Figs 106, 107. ...... ....................................................................................................   M. regalis sp. nov.   12 (11) Elytra lacking tactile setae; appressed setae of pronotal disc coarser, well-shaped, truncate to shortly bifurcate apically; distal antennomeres slender, more elongate, antennomere X nearly twice as long as wide; male characters as in Figs 80, 81. ..... .....................................................................................................   M. mollis sp. nov.   13 (10) Elytral setation short and very dense, surface entirely obscured ( Fig. 175).   14 (15) Antero-lateral margins of frons distinctly raised and moderately lobed (lobule smooth and glossy subapically); male characters as in Figs 42, 43. ...................................... ..............................................................................................   M. dearmatus sp. nov.   15 (14) Antero-lateral margins of frons simple or at most slightly raised.   16 (17) Elytral setation somewhat heterogeneous, mostly appressed, with slightly raised setae of coarser punctures ( Fig. 178), and some tactile setae distinct at least near base; male characters as in Figs 82, 83. ..................................................   M. nobilis sp. n.   17 (16) Elytral setation quite uniformly short and appressed, tactile setae lacking, including basal portion.   18 (19) Larger, more robust species, body length 2.9–3.1 mm; darker elytral scales usually forming rather extensive and distinct brownish markings ( Fig. 196); male characters as in Figs 55–57. .................................................................   M. granulatus sp. nov.   19 (18) Smaller, slender species, body length 2.2–2.6 mm; darker elytral scales forming mostly rather vague brownish spots; male characters as in Figs 62, 63. ................... ..................................................................................................   M. imitator sp. nov.   20 (5) Pronotal disc at most with few minute granules near basal margin (  M. festivus sp. nov.) or antero-medially near base of pronotal horn.   21 (26) Antero-lateral margins of frons with distinct process ( Figs 26–27, 30, 128–129).   22 (23) Process of frontal margin conspicuously robust, long and curved, horn-like ( Figs 26, 128). .........................................................................................   M. bicornis sp. nov.   23 (22) Process of frontal margin much smaller, somewhat flattened.   24 (25) Paired rugules of clypeus very conspicuous ( Fig. 129, marked by an arrow); process of frontal margin somewhat larger, its posterior tip rather rounded and pointing dorsally ( Fig. 27); male metatarsomere III moderately enlarged in apical half; male characters as in Figs 28, 29. .......................................................  M. bidens sp. nov.   25 (24) Paired rugules of clypeus minute; process of frontal margin lobed anteriorly, its posterior tip sharp and pointing evenly posteriorly ( Fig. 30); male metatarsomere III strongly enlarged in apical half; male characters as in Figs 31, 32. ..................... ..............................................................................................  M. bidentatus sp. nov.   26 (21) Antero-lateral margins of frons simple or at most moderately raised / lobed, lacking any distinct process.   27 (30) Scales of elytra entirely covering surface, coalescent, with extremely fine, indistinct margins ( Fig. 173).   28 (29) Body somewhat paler coloured, elytral scales at least partly with pale reddish tinge (mainly apically; Fig. 192); parameres more robust and somewhat sinuously narrowing in apical third ( Fig. 41); male tergum VIII as in Figs 40. ............................. ............................................................................................  M. concolor King, 1869   29 (28) Body generally darker coloured, elytral scales dark brown and silvery, apex of elytra darker ( Fig. 197); parameres more slender and evenly narrowing in apical third ( Fig. 61); male tergum VIII as in Fig. 60. ...................................  M. hortensis Lea, 1922   30 (27) Scales of elytra well-spaced, if contiguous and entirely covering surface, then with margins clearly visible.   31 (34) Body setation generally scaly, scales short and wide, covering most of head surface, nearly all antennomeres, legs including tarsomeres, and dorsal side of pronotal horn.   32 (33) Eyes comparatively large; pronotal horn conspicuously slender and elongate, with extremely narrow crest ( Fig. 167), body scales greyish, white and black, forming conspicuous, sharply outlined markings on elytra ( Figs 185, 208); male characters as in Figs 112–115. .......................................................................   M. weiri sp. nov.   33 (32) Eyes very small; pronotal horn much wider and less elongate, its crest distinctly wider; body scales dark to pale brownish, partly mixed and thus markings of elytra less contrasting ( Fig. 189); male characters as in Figs 15–18. .................... ...................................................................................................   M. auceps sp. nov.   34 (31) Body setation at least partly hairy, pronotal horn sometimes with hairy or scaly setation, but never entirely covered dorsally (except for rugules) by contiguous, appressed scales.   35 (36) Horn crest at most slightly indicated, dorsal side of pronotal horn evenly vaulted and covered by small rugules ( Fig. 155); male characters as in Figs 72, 73. .................. ........................................................................................................  M. leai Pic, 1942   36 (35) Horn crest well-developed.   37 (40) Setation of elytra hairy, setae long and rather sharply pointed apically ( Fig. 170).   38 (39) Pronotum globose, evenly rounded laterally in dorsal view; pronotal horn wider, subtriangular, posterior angles distinct; horn crest short and compact, situated rather posteriorly; body setation sparser, silvery, pronotum (antero-laterally) and elytra with numerous erect tactile setae; male characters as in Figs 38, 39. ............................... .................................................................................................   M. centralis sp. nov.   39 (38) Pronotum somewhat more elongate, unevenly rounded laterally in dorsal view; pronotal horn narrower, posterior angles indistinct ( Fig. 145); horn crest longer, extending (evenly lowering) anteriorly; body setation denser, with goldish or cupreous shiny reflection, and forming brownish markings on elytra, pronotum (antero-laterally) and elytra lacking erect tactile setae; male characters as in Figs 19–21. .................. ..............................................................................................   M. auripilosus sp. nov.   40 (37) Setation of elytra scaly, setae mostly distinctly shorter and wider, rounded to truncate apically, rarely linear, but never sharply pointed apically.   41 (42) Setation of elytra clearly double, comprising shorter, appressed, whitish scales and distinctly longer, linear, more raised, apically truncate, cupreous shining setae ( Fig. 169); body length 3.7–4 mm; body coloration generally dark, at most with paler subapical spot on elytra ( Fig. 188); male characters as in Figs 13, 14. ..................... .........................................................................................  M. apicipennis Lea, 1895   42 (41) Setation of elytra quite uniform, appressed, if nearly double, comprising longer and more raised scales (  M. canthariphilus sp. nov.) then body length less then 2.5 mmand body generally paler coloured, mostly with darker markings of elytra as in Figs 187, 209.   43 (66) Dorsal surface of pronotal horn nearly bare, or at most sparsely, inconspicuously setose, especially in apical half.   44 (57) Elytral scales shortly oval, rather rounded apically ( Figs 177, 179, 182), entirely appressed and very dense, mostly entirely covering surface; lacking scattered erect tactile setae of elytra (at most with some longer bristly setae on humeri and basal margin).   45 (46) Setation of genae rather coarse and modified, arranged as in Fig. 133; male characters as in Figs 108, 109. ................................................................   M. setulosus sp. nov.   46 (44) Setation of genae finer and evenly arranged (all setae pointing more or less anteriad).   47 (48) Head vertex sharply impressed along median margins of eyes ( Fig. 131); male characters as in Figs 47, 48. .................................................   M. exophthalmus sp. nov.   48 (47) Head vertex evenly curved, lacking any distinct impressions.   49 (50) Frontal portion of head prolonged; antero-lateral margins of frons subparallel and moderately raised / lobed ( Fig. 132); male characters as in Figs 67, 68. .................. ..........................................................................................  M. kingii MacLeay, 1872   50 (49) Frontal portion of head relatively short; antero-lateral margins of frons narrowing anteriorly and simple.   51 (52) Larger, robust species (body length 3.5–4.2 mm; Fig. 204); body scales generally distinctly glossy ( Fig. 179); male characters as in Figs 84, 85. ....   M. obesus sp. nov.   52 (51) Smaller species (body length 1.6–2.5 mm); body scales at most slightly glossy, if distinctly glossy (  M. pusillus sp. nov.; mainly scales on pronotal disc), then body length less than 2.0 mm.   53 (54) Horn crest longer, evenly narrowing and lowering anteriorly; median rugules largely fused and forming irregular sculpture, which may be connected with rugules of crest margins; male characters as in Figs 102, 103. ....................   M. pilbarensis sp. nov.   54 (53) Horn crest short and more compact, situated more posteriorly; rugules of crest margins usually fused and forming evenly shaped rim; single, rather evenly shaped median longitudinal rugule ( Figs 156–158, 165).   55 (56) Body scales at least partly whitish (always on pronotum); lobules of pronotal horn margins narrower and widely spaced, submarginal rugules rounded and rather unevenly scattered ( Figs 156–158); posterior collar of pronotum with distinct fringe of whitish scales; male characters as in Figs 74, 75. ...............   M. magelae sp. nov.   56 (55) Body scales mostly reddish to brownish, contiguous to nearly fused; lobules of pronotal horn margins wide, narrowly spaced to fused, and submarginal rugules elongate, contiguous to fused in dense rows ( Fig. 165); posterior collar of pronotum with inconspicuous scales few (not really fringed); male characters as in Figs 104, 105. ...........................................................................................   M. pusillus sp. nov.   57 (44) Elytral scales distinctly elongate (linear, nearly parallel-sided), truncate or at least subtruncate apically and mostly well-spaced ( Figs 172, 176); elytra with or without scattered erect tactile setae.   58 (59) Elytral scales brownish and rather contrastingly whitish, partly mixed and also forming wide transverse bands ( Fig. 199); pronotal horn rather long and slender; horn crest long, extending (evenly lowering) anteriorly; rugules of crest margins well-spaced; male characters as in Figs 69–71. ......................................  M. kreusleri King, 1869   59 (58) Elytral scales of various colours (whitish to greyish, reddish, brownish) but never forming wide transverse bands; pronotal horn usually short and wide (except  M. grandior sp. nov.); horn crest short and compact, situated rather posteriorly; rugules of crest margins contiguous to fused.   60 (61) Large species (body length 2.7–3.7 mm); body generally dark coloured and nearly opaque ( Fig. 195); pronotum narrower, moderately convex laterally; pronotal horn longer and more slender ( Fig. 151), rugules of crest margins mostly partly fused and forming semioval rim, interrupted anteriorly (see Variation); male characters as in Figs 53, 54. .............................................................................  M. grandior sp. nov.   61 (60) Mostly smaller species (body length 2.1–2.9 mm; except  M. bullatus sp. nov.); body generally paler coloured (pale reddish to reddish brown) and somewhat glossy; pronotum globose to moderately transverse, strongly convex laterally; pronotal horn short and wide, rather triangular, with distinct posterior angles ( Figs 147, 148, 162, 163); rugules of crest margins either separate or forming semicircular rim (complete, never interrupted anteriorly).   62 (63) Rugules of crest margins contiguous to nearly fused, but never forming evenly shaped rim ( Figs 162, 163); elytra lacking scattered tactile setae; male characters as in Figs 88–97. ............................................................................  M. phanophilus Lea, 1922   63 (62) Rugules of crest margins fused and forming evenly shaped semicircular rim ( Figs 147, 148); elytra with scattered tactile setae.   64 (65) Larger, more robust species (body length 2.7–3.7 mm); scaly setation of pronotal disc double, with longer subdecumbent and shorter appressed scales; scales of elytra somewhat shorter and wider ( Fig. 172); elytra in females with apical margin rounded to straight, apical angle rather rounded; male characters as in Figs 33, 34. .............. ..................................................................................................   M. bullatus sp. nov.   65 (64) Smaller species (body length 2.2–2.6 mm); scaly setation of pronotal disc simple, uniformly short and appressed; scales of elytra somewhat longer and narrower ( Fig. 176); elytra in females with apical margin slightly concave and median apical angle thus more prominent; male characters as in Figs 58, 59. ...   M. hirtipennis sp. nov.   66 (43) Dorsal surface of pronotal horn rather densely, conspicuously setose.   67 (74) Dorsal setation of pronotal horn hairy, distinctly finer than scales of pronotal disc ( Figs 143, 150, 161, 164).   68 (69) Pronotal horn with single large, evenly shaped, longitudinal median rugule ( Fig. 161); scales of elytra whitish and brownish, mixed ( Fig. 180) and forming mostly distinct darker transverse bands /spots ( Fig. 205); male characters as in Figs 86, 87. .......... .................................................................................................   M. parvulus sp. nov.   69 (68) Pronotal horn with numerous median rugules, separate or largely fused and forming irregular sculpture; scales of elytra mostly paler coloured (pale reddish), setose dark markings different or indistinct.   70 (71) Dark markings of elytra as in Fig. 194(see Variation); metatarsi distinctly longer then metatibiae; male characters as in Figs 51, 52. ..................   M. festivus sp. nov.   71 (70) Dark markings of elytra as in Fig. 187(if well-developed); metatarsi at most slightly longer than metatibiae.   72 (73) Pronotum narrower, unevenly rounded laterally in dorsal view; pronotal horn more elongate ( Fig. 143); scales of elytra rather linear and subdecumbent ( Fig. 168); male sternum VII distinctly modified, subtruncate and moderately angled postero-laterally ( Fig. 10); male characters as in Figs 11, 12. ........................  M. amabilis Lea, 1895   73 (72) Pronotum wider, rather evenly rounded laterally in dorsal view; pronotal horn shorter and wider ( Fig. 164); scales of elytra shorter and wider, drop-shaped, and appressed; male sternum VII simple; male characters as in Figs 98–101. ............. ................................................................................................  M. pilicornis sp. nov.   74 (67) Dorsal setation of pronotal horn scaly, similar to that on pronotal disc ( Fig. 142; somewhat finer only in  M. canthariphilus sp. nov.).   75 (76) Elytral and pronotal scales at dark coloured places cupreous shiny; dorsal surface of pronotal horn rather scantily and finely setose in apical half, scales distinct only latero-basally, rather narrow to linear; male sternum VII with smooth, sharply bordered, subapical facet ( Fig. 35); male tergum VIII and aedeagus as in Figs 36, 37. .......................................................................................   M. canthariphilus sp. nov.   76 (75) Elytra and pronotal disc usually with brownish setose markings, but scales never cupreous; dorsal surface of pronotal horn largely distinctly setose, setae forming distinctly widened scales; male sternum VII simple, at most somewhat sparsely setose subapically (except  M. iuvenis sp. nov.).   77 (78) Male sternum VII with small bordered median impression near base ( Fig. 64); posterior setation of male tergum VIII rather hairy (setae pointed apically), long and raised ( Fig. 65); aedeagus as in Fig. 66. ....................................  M. iuvenis sp. nov.   78 (77) Male sternum VII simple, lacking bordered impression; posterior setation of male tergum VIII rather scaly (setae coarser, blunt apically), short and less raised ( Figs 7, 49, 116).   79 (80) Parameres of aedeagus moderately wide, subparallel and moderately curved in apical third, rather abruptly narrowed and slightly produced apically ( Fig. 50). ................ ......................................................................................................  M. fallax sp. nov.   80 (79) Parameres of aedeagus narrower and strongly curved in apical third ( Fig. 9), with simply rounded apex.   81(82) Crest of pronotal horn usually wider ( Fig. 142); parameres of aedeagus unevenly narrowing, with distinct preapical denticle / angulation on outer margin ( Fig. 8). ... ..........................................................................................  M. albellus Pascoe, 1866   82(81) Crest of pronotal horn usually narrower; parameres of aedeagus evenly narrowing, simple subapically ( Fig. 117). ...............................................  M. ziczac King, 1869