Siphostoma starksii P. starksii Siphostoma starksi Jordan & Evermann, 1896a: 771 Rutter, 1896: 263 Meek, 1904: 162–163 Siphostoma ( Siphostoma ) starksii Jordan & Evermann, 1896b: 328 Dawson, 1982: 47 P. starksii Syngnathus starksi Starks, 1906: 783 Jordan et al., 1930: 242 De Buen, 1940: 46 Siphostoma elcapitanense Dawson, 1982: 47 P. starksii Syngnathus elcapitanense Meek & Hildebrand, 1923: 260 Dawson, 1982: 47 P. starksii Pseudophallus starksii : Gunter, 1942: 310 Fowler, 1944: 496 Gunter, 1956: 349 Follett, 1960: 213 Dawson, 1982: 45–49 Dawson, 1985: 161 Pseudophallus Ruiz-Campos et al., 2002: 148 Kuiter, 2009: 241 McMahan et al ., 2013: 448 Page et al., 2013: 110 González-Acosta et al., 2015: 72 Angulo et al., 2015: 7 Angulo et al ., 2020: 9 starksii Pseudophallus starksi Herald, 1940: 51 Pseudophallus Herald 1959: 473 Miller, 1966: 796 Bussing, 1966: 222 Whatley, 1969: 448 Villano & Balderas, 1987: 230 Castro-Aguirre et al ., 1995: 72 Castro-Aguirre & Balart, 1997: 144 Bussing 1998: 252–253 Castro-Aguirre et al ., 1999: 234 Angulo et al ., 2013: 998 Pseudophallus starksi starksi : Lüling, 1974: 201 P. starksi Pseudophallus lineatus Villar, 1970: 80 Dawson, 1982: 47 P. starksii Taxonomic review of the pipefish genus Pseudophallus Herald, with the description of a new species (Syngnathiformes: Syngnathidae) Dallevo-Gomes, Caio I. A. Mattox, George M. T. Toledo-Piza, Mônica Zootaxa 2020 2020-10-05 4859 1 81 112 4P6RH (Jordan & Culver 1895) [151,424,1266,1293] Actinopterygii Syngnathidae Pseudophallus Animalia Syngnathiformes 17 98 Chordata species starksii  ( Figs. 2–3, 5, 7; Table 2)    Siphostoma starksii Jordan& Culver ( in  Jordan, 1895): 381, 416, 514, pl. XXX (original description; “Río Presidio, about a mile below from village Presidio, Sinaloa, Mexico”; type specimens SU 2686);  Jordan& Evermann 1896b: 328(listed in North America);  Jordan& Evermann, 1898: 2838(redescription; type listed as SU 2686);  Jordan& Evermann, 1900: 3259, pl. CXX:3 25 (explanation of pl. CXX, drawing of SU 2686); Böhlke, 1953: 62( syntypeslisted); Dawson, 1982: 47(synonymic list of  P. starksii).    Siphostoma starksi(sic):   Jordan& Evermann, 1896a: 771(, redescription, typelisted as USNM 47425);  Rutter, 1896: 263(occurrence in San José del Cabo, Baja California, Mexico);  Meek, 1904: 162–163(occurrence in Río San Geronimo, Mexico, key of identification).    Siphostoma( Siphostoma) starksii:   Jordan& Evermann, 1896b: 328, 576 (new combination; listed in Central America);  Dawson, 1982: 47(synonymic list of  P. starksii).    Syngnathus starksi(sic):  Starks, 1906: 783(new combination, occurrence in Santa Rosa, Ecuador); Regan, 1907: 55 (listed);   Jordanet al., 1930: 242(listed in Central America);  De Buen, 1940: 46(listed in Mexico).    Siphostoma elcapitanense: (non Meek & Hildebrand, 1914): Meek & Hildebrand ( in Meek, 1914): 119 (misidentification; Jesús Maria and Turrubales, Costa Rica);  Dawson, 1982: 47(synonymic list of  P. starksii).    Syngnathus elcapitanense: (non Meek & Hildebrand, 1914):  Meek & Hildebrand, 1923: 260(in part: incorrect identification of specimens from Costa Ricaand Punta Chame, Panama);  Dawson, 1982: 47(synonymic list of  P. starksii).     Pseudophallus starksii: Gunter, 1942: 310(euryhaline);  Fowler, 1944: 496(listed in Central America);  Gunter, 1956: 349(listed in Central America);  Follett, 1960: 213, 214, 224 (mention of affinities with congeners);  Dawson, 1982: 45–49, 165, figs. 17, 20, 70 (species account, comparisons with congeners, identification key; specimen CAS(SU) 67185 (ex SU 2686) as lectotype);  Dawson, 1985: 161, 162, 209, figs. 240, 241 (map) (characters, distribution, distribution, comparisons with other species of  Pseudophallus,identification key); Ferraris Jr., 2003: 591 (listed, distribution; SU 67185 (ex SU 2686) as lectotype);  Ruiz-Campos et al.,2002: 148, 149 (listed in Baja California Sur, Mexico); Nelson et al.,2004: 112 (listed for Central America); Robles & Vega, 2004: 82 (listed in Santa Maria river, Veraguas, Panama);  Kuiter, 2009: 241(distribution, freshwater, figure from Dawson, 1982); Mol, 2012: 51 (citation of literature regarding positive rheotropism);  McMahan et al., 2013: 448(listed in El Salvador);  Page et al.,2013: 110(listed in Central America);  González-Acosta et al.,2015: 72(listed in Baja California, Mexico);  Angulo et al.,2015: 7, 9, 10; fig. 5K (listed, distribution in Costa Rica, identification key);  Angulo et al., 2020: 9; fig. 8N (as starski), 19, 28 (as  starksii, listed in Costa Rica, identification key).    Pseudophallus starksi(sic):  Herald, 1940: 51, 59, 63 (new combination; designated as typespecies of  Pseudophallus; species account; identification key);  Herald 1959: 473(general occurrence in the Pacific coast);  Miller, 1966: 796(coastal streams, San José del Cabo to Ecuador);  Bussing, 1966: 222, 236 (occurrence in Costa Rica. listed);  Whatley, 1969: 448(cited as euryhaline fish); Dawson, 1974:409, 410 (meristic characters in tables and identification key); Lagler et al., 1977: 39 (figure from Jordan& Evermann 1896);  Villano & Balderas, 1987: 230(listed in Mexico);  Castro-Aguirre et al., 1995: 72(listed in Gulf of California, Mexico, biogeography);  Castro-Aguirre & Balart, 1997: 144(listed, La Paz Bay, Baja California Sur, Mexico);  Bussing 1998: 252–253, 257–259, figures and map 24 (identification key, species account, distribution in Costa Rica);  Castro-Aguirre et al., 1999: 234(synonymy list, distribution in Mexico); Umaña in Lobo & Bolaños, 2005:76(listed in Costa Rica); Miller et al., 2006:148(listed, short diagnosis, distribution in Mexico); Vega et al., 2006: 92 (listed in Panamá); Matamoros et al., 2009:16, 35 (in part, listed in Honduras);  Angulo et al., 2013: 998(listed, distribution in Costa Rica).     Pseudophallus starksi starksi: Lüling, 1974: 201(as a geographic race of  P. starksi).    Pseudophallus lineatus: (non Kaup): Alvarez del  Villar, 1970: 80(new combination, incorrect identification, characters and distribution in Mexico);  Dawson, 1982: 47(synonymic list of  P. starksii).   Diagnosis.  Pseudophallus starksiidiffers from  P. brasiliensis,  P. mindiiand  P. galadrielaein having 95.3% of specimens with 14 trunk rings ( vs. most usually with 13 trunk rings).   Pseudophallus starksiican be distinguished from  P. elcapitanensisby the following combination of characters: 36–44 dorsal-fin rays (89.5% of specimens with 38–41), 49–53 body rings (91.9% of specimens with 50–51), 19–21 brood-pouch rings (64.9% of adult male specimens with 21) and 8.5–10.5 subdorsal tail rings (86.7% of specimens with 9.25–10.0) vs. 30–35 dorsal-fin rays (82.1% of specimens with 31–33), 47–51 body rings (89.8% of specimens with 48–49), 17–19 brood-pouch rings (90.9% of adult males specimens with 17–18) and 6.75–7.5 subdorsal tail rings (97.4% of specimens with 6.75–7.25) in  P. elcapitanensis.   Description.Meristic and morphometric characters detailed in Table 2. General appearance and coloration in Figs. 2Dand 3D. Body without scales, covered with bony plates forming rings along its axis, bony plates horizontally oval, with an elevation in midline forming bony ridges along body. Trunk rings 13–15 (95.9% of specimens with 14). Tail rings 35–39 (91.1% of specimens with 36–37). Body rings 49–53 (91.9% of specimens with 50–51). Subdorsal tail rings 8.5–10.5 (87.8% of specimens with 9.25–10.0). Subdorsal trunk rings from 0–0.5. Subdorsal tail rings 8.5–10.5 (87.0% of specimens with 9.25–10.0). Dorsal-fin origin at vertical between posterior half of posteriormost trunk ring and anteriormost caudal ring. Posterior tip of superior trunk ridge at vertical through superior tail ridge, between 6.0 and 7.5 anteriormost tail rings. Lateral trunk ridge continuous with superior tail ridge with origin on first caudal ring. Inferior trunk ridge continuous with inferior tail ridge. Body slender and cylindrical, transversal section of trunk heptagonal, with transition to quadrangular in tail at end of superior trunk ridge. Maximum body depth of females and juveniles at middle of trunk; in males at anterior region of the brood pouch. Body depth decreasing posteriorly along superior trunk ridge to minimum depth on caudal peduncle. Dorsal profile of head straight or slightly inclined at snout region, more inclined at orbital region and straight at posterior region, with two nuchal plates between occipital region and first trunk ring. Ventral profile of head straight. Dorsal fin not elevated, curved on first rays, straight along median portion with pointed end. Dorsal-fin rays of similar length, except the first two rays. Dorsal-fin rays 36–44 (89.5% of specimens with 38–41), posteriormost two rays closer to each other, difficult to distinguish in some specimens. Pectoral fin small, middle rays longer than outer and inner rays. Pectoral-fin origin on anteriormost trunk ring, and distal margin rarely extending beyond third trunk ring. Pectoral-fin rays 11–15 (68.5% of specimens with 14). Caudal fin small and fan like, with 10 rays (5 rays in each hypural plate). Pelvic and anal fins absent. Mouth terminal, slightly inclined dorsally, jaws edentulous. Lower jaw slightly prognathous. Snout short, tubular, without dorsal or lateral ridges. Dorsal profile of snout with slight inclination following dorsal margin of eyes. Eyes located longitudinally at middle of head. Opercle occupying most of lateral surface of head posterior to eye, opercular opening reduced to pore on posterodorsal margin of opercle. Opercular ridge generally indistinct, except for some specimens with small protuberance on anterior region of opercle, following superior margin of bone. Lateral line and lateral line canals absent.  Sexual dimorphism.Males with brood pouch. Origin of brood pouch on posteriormost trunk rings, brood pouch rings extending along 19–22 rings (64.8% of specimens 21 brood pouch rings). Bony plates of the brood pouch well developed and T-shaped encasing entire lateral portion of pouch, decreasing in size posteriorly and parallel with the inferior tail ridge. Ventral region of pouch with contralateral dermal folds extending towards the midline, dividing brood pouch longitudinally. Adult females with anal papilla on anal ring with small, barely visible spicules.  Coloration in alcohol.Ground color of head and body light beige to brown, sometimes with horizontal stripe more evident on the head and anterior portion of the body formed by different tones of brown. Smaller stripes on dorsolateral part of head and dorsal part of opercle. Thicker bony regions at body ridges with less or no chromatophores, forming nearly white longitudinal lines. Some specimens with series of darker arch-shaped blotches in lateral view, at every two or four rings. Bony rings with one or two white spots underneath bony plates; shape of spot more circular when single spot present and oval shaped due to proximity of the spots when two spots present. Dorsolateral part of snout with layer of skin and mesethmoid forming light stripe. Dorsal-fin rays with scattered brown chromatophores. Pectoral fin hyaline. Caudal fin with same general coloration of body, sometimes in slightly different tone. ( Figs. 2Dand 3D).   Distribution.Freshwater coastal drainages of the Pacific, from Río Santa Rosa, Ecuador, to San Jose del Cabo, Baja California, Mexico( Fig. 5).   Remarks.A holotypewas not designated in the original description of  Siphostoma starksii, but  Jordan(1895: 381)listed as 2686 the catalog number of the type specimens of the species at the Museum of Stanford University (SU, currently at CAS). Three specimens from this original lot were sent to three different institutions: the United StatesNational Museum of Natural History (USNM 47425), Zoologisches Museum Berlin (ZMB Pisces 14058) and the British Museum, London (BMNH 1895.5.27.321), with five specimens remaining as SU 2686 (now CAS 102686). Later, one additional specimen from lot SU 2686 was dismembered into CAS-SU 67185 (now CAS 167185). Therefore, those eight specimens are part of the original type series (i.e., syntypes) of the species (ICZN, 1999: Art. 72.4.1.1 and 73.2). In the first part of a series of four volumes of “The fishes of North and Middle America”,  Jordan& Evermann (1896a: 771)indicated the lot USNM 47425 as “ type” of  Siphostoma starksi(sic). On the third part of the work by  Jordan& Evermann (1898: 2838), SU 2686 is indicated as “ type” of  Siphostoma starksiiand in the fourth part of the work by  Jordan& Evermann (1900: 3259, pl. CXX) one specimen from lot SU 2686 is illustrated.  Dawson (1982: 45, 49) listed lot SU 67185 (now CAS 167185) as lectotypeof  Siphostoma starksii. Ferraris (2003: 591)indicated this specimen as being the same of that illustrated in  Jordan& Evermann (1900: 3259, pl. CXX) and therefore, credited the lectotypedesignation of the species to the latter authors. However, the previous mention of the specimen USNM 47425 as type of  S. starksii, by  Jordan& Evermann (1896a: 771)should be considered as the lectotypedesignation of the species (ICZN, 1999: Art. 74.5) and other lectotypedesignations (e.g.,  Jordan& Evermann, 1900or Dawson, 1982) are deemed invalid (ICZN 1999: Art. 74.1.1).   Type material:   Lectotype. USNM 47425, 1, Río Presidio, about a mile below village of Presidio, Sinaloa, Mexico, D. S. Jordan et al.,  24 Dec 1894–  25 Jan 1895(not examined).   Paralectotypes. CAS 102686, 4, 51.0– 123.3 mmSL,  CAS 167185, 1, 132.2 mmSL, same data of lectotype; ZMB 14058, 1, BMNH 1895.5.27.321, 1, same data of lectotype(not examined).  Non   typematerial. Ecuador: El Oro: CAS 109312, 1, 117.2 mmSL, Riverat Santa Rosa, P. O. Simmons 1898-1899.    Colombia: Valle del Cauca:AMNH 32971 2, 117.3– 124.4 mmSL, quebrada Guangui, 1/  2 kmabove Río Patia, upper Río Saijadrainage, J. W. Daly,  12 Feb 1973.    Panama: Darien:MCZ 52992, 1, 64.0 mm SL, Boca de Río Capeti, in fast riffle about  10mupstream of mouth at Río Tuira, 08º05`N 77º35`W, K. E. Hartel,  27 Mar 1978; MCZ 88624, 7, 112.4–  146.5 mmSL, Río Picuro, just above confluence with Río Tuira, 08º00`N 77º32`W, W. C. Starnes et al.,  16 Feb 1985; USNM 208365, 6, 134.9–  171.1 mmSL, upper tributary of Río Jaque, H. G. Loftin& Dean,  11 Feb 1968; USNM 293603, 9,  68.9–135.6 mmSL, Río San Antonioat Garachini, trib. Golfo de San Miguel, J. Lundberg, R. Bouchard& J. Louton,  07 Mar 1988;    Panama:UF 12983, 1, 127.8 mmSL, Creek  11 mi.S of Chepoon road to El Llano, Brockman& Child,  29 Apr 1965; USNM 293475, 8,  72.3–100.1 mmSL, Pan American Highwayca. 17 Km. Eof el Chepo( Bayano Drive), Río Terable, W. Starnes et al.,  25 Feb 1985;   Veraguas:ANSP 104211, 3, 106.0– 124.5 mmSL, Creek Quebrada Piedraat bridge about 8 mi.W. of El Maria on Sona-Remediosroad, H. G. Loftin, E. L. Tyson& M. Canipe,  11 Nov 1961; ANSP 104408, 4, 63.3– 116.3 SL, creek at bridge  3 mi.E of Sona-Santiagoroad, H. G. Loftin& E. L. Tyson  28 Oct 1961;   Chiriqui:ANSP 104181, 1, 121.9 mmSL, creek into Río Jacaquenear San Lorenzoat farm of Dr. Alvarez, H. G. Loftin& E. L. Tyson,  9 Dec 1961; ANSP 119934, 2,  54.4–81.6 mmSL, Creek  5 mi.W of Boca del Monteon old road to David, H. G. Loftin& E. L. Tyson  18 Nov 1961; ANSP 119935, 2,  63.7–91.1 mmSL, creek at bridge 5 miW. of San Felixbridge on Interamerican Highwaynew section, H. G. Loftin& E. L. Tyson  18 Nov 1961; ANSP 11936, 18,  50.6–124.7 mmSL, Río Tinta, creek at bridge  3 mi.W of Río Tabasara on Sona-Remediosroad, H. G. Loftin, E. L. Tyson& M. Canipe,  11 Nov 1961: ANSP 146754, 1, 98.8 mmSL, Río Las Vueltasat Interamerican Hwy, st. 51, D. Fromm& P. Fromm,  4 Mar 1981.   TABLE 3. Frequency distribution of number of dorsal–fin rays in specimens of  Pseudophallusfrom Atlantic drainages. N=261. Regions are arranged following a South to North sequence. * indicates specimens of  P. galadrielaefrom Lago Izabal drainage. Values for  P. mindiiare marked in grey.    27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 Total  Brazil São Paulo 1 6 14 12 10 12 1 56  Rio de Janeiro 2 3 2 7  Espírito Santo 1 3 1 6 4 3 4 1 23  Bahia 1 3 2 3 1 1 11  Piauí 1 1  Pará 5 7 4 2 18  Suriname 2 2 2 6  Guiana 1 1  Venezuela—until Orinoco Delta 1 2 1 5 1 1 11  Venezuela—From Rio Cumboto 2 1 1 4  Panama 1 2 4 3 1 11  Costa Rica 2 2 3 6 2 1 16  Nicaragua 1 1 1 3  Honduras 1 1 1 3  South of Lago Izabal 2 1 2  Lago Izabal* 1 10 17 24 14 4 1 1 72  North of Lago Izabal 1 1 2  Belize 1 5 2 1 9  Caribbean 3 1 1 5  Total 1 18 30 34 26 15 19 25 31 20 26 8 4 2 2 1 261   TABLE 4. Frequency distribution of number of total body rings in specimens of  Pseudophallusfrom Atlantic drainages. N=256. * indicate specimens of  P. galadrielaefrom Lago Izabal drainage. Values for  P. mindiiare marked in grey.    40 41 42 43 44 45 46 47 48 49 50 51 Total  Brazil São Paulo 1 4 19 18 10 4 56  Rio de Janeiro 4 2 1 7  Espírito Santo 1 10 10 3 24  Bahia 5 1 3 9  Piauí 1 1  Pará 1 1 2 11 2 1 18  Suriname 3 2 5  Guiana 1 1  Venezuela— until Orinoco Delta 3 2 1 4 10  Venezuela— From Rio Cumboto 1 1 2  Panama 3 7 2 12  Costa Rica 1 3 8 4 16  Nicaragua 2 1 3  Honduras 2 1 3  South of Lago Izabal 2 2  Lago Izabal* 3 15 35 16 2 1 71  North of Lago Izabal 1 1 2  Belize 1 5 2 1 2 9  Caribe 1 2 2 5  Total 1 4 15 41 31 26 38 45 34 14 5 2 256    Costa Rica: Puntarenas:FMNH 7901, 1, 83.4 mmSL, Jesus Maria River, S. E. Meek, 1912; FMNH 76970, 1, 98.1 mmSL, Río Grande Terraba, R. K. Johnson,  7 Jul 1973; ROM 88191, 3, 81.3–138.0 mm SL, Quebrada Benjamin,  1 kmWest of Palmar Norte, Osa Peninsula, 08º57`54’’N 83º27`44”W, B. Wisenden,  1 May 1991; TNHC 15359, 9, 91.6–130.3 mmSL, Río Pavo, upstream from Los Patostrail, P. N. Corcovado, 08º30`29’’N 83º36`03’’W, K. Winemiller,  13 Jul 1983; UF 19558, 1, 125.1 mmSL, Río Chires, ca 2 kmN off Estarillos, ca 25 air mi. WNW of Quepos, C. Gilbert et al.,  2 Jul 1973;   San José:ANSP 164263, 3,76.1–138.0 mm SL, Río Barúca  7 kmabove Dominical, D. Fromm,  11 Mar 1989;   Guanacaste:UF 19569, 1, 158.8 mmSL, Río Corobia,  5 kmNW of Las Canas, near Pan American Hwy, at La Pacificamotel, Gilbert, McAllister& Balon,  4 Jul 1973.    Honduras: Valle:LACM 32295–11, 1, 135.8 mmSL, Río Nacaomeat Pan American Highway, Martin& Brame,  8 Feb 1969.    El Salvador: La Union:TU 202696, 7, 88.2–131.6 mmSL, trib. to Gulfo de Fonseca, 35.2 milesE of San Miguel, Pan American Hwy, R. D. Sutkus& Smalley,  2 Jun 1962.    Guatemala: Santa Rosa:AMNH 25100, 1, 87.3 mmSL, Río Margarita, under bridge of  Carretera Internacional Del Pacifico, D.E. Rosen, K. Kallman & R. Dorion,  24 Feb 1963.    Mexico: Oaxaca:FMNH 4717, 1, 104.3 mmSL, San Geronimo, S. E. Meek,  29 Feb 1903;   Guerrero:UMMZ 178462, 8, 69.2–103.0 mm SL, Río Coyaquilla, 60 road miles SE of Zihuatenejo, R. R. Miller& M. Miller,  21 Mar 1957;   Nayarit:UMMZ 172107, 8, 87.3–120.1 mmSL, Río La Limaat La Lima, 7.5 miby road SSW of Las Varas, R. R. Miller& J. T. Greenbank,  24 Feb 1955; UMMZ 178588, 10, 85.0– 100.4 mmSL, Río Rosarioat Hwy.15 bridge near Rosario, Pacific Dr, R. R. Miller& M. Miller,  08 Apr 1957;   Baja California Sur: CAS( SU) 5943, 4, 67.1–124.9 mmSL, San Jose del Cabo, Eisen& Vaslit. 3034656372 1894-12-24 1895-01-25 1894-12-24 USNM D.J. Stewart Mexico Rio Presidio 19 100 USNM 47425, 1 1 Sinaloa lectotype 3034656383 CAS Mexico 19 100 CAS 102686, 4 1 Sinaloa paralectotype 3034656438 ZMB 14058 CAS Mexico 19 100 CAS 167185 2 Sinaloa lectotype 3034656434 CAS River & Santa Rosa, P. O Ecuador El Oro 19 100 CAS 109312 1 El Oro holotype 3034656431 1973-02-12 J. W. Daly Colombia upper Rio Saija 19 100 1 Valle del Cauca 3034656439 MCZ 52992, MCZ 88624 1978-03-27 MCZ K. E. Hartel Panama 10 8.083333 Rio Tuira 1302 -77.583336 Boca de Rio Capeti 19 100 7, 112.4 1 Darien 3034656441 1985-02-16 USNM W. C. Starnes Panama 8.0 Rio Tuira 1302 -77.53333 Rio Picuro 19 100 USNM 208365, 6, 134.9 1 Darien 3034656416 [151,1248,1770,1797] 1968-02-11 USNM H. G. Loftin & Dean Panama Rio Jaque 19 100 USNM 293603, 9 1 Darien 3034656366 1988-03-07 Lundberg, R & Bouchard & Louton Panama Garachini Rio San Antonio 19 100 1 Darien 3034656353 1965-04-29 USNM El Llano & Brockman & Child Panama Creek 19 100 USNM 293475, 8 1 3034656356 1985-02-25 Km. E & Bayano Drive & W. Starnes Panama Pan American Highway Rio Terable Chepo 19 100 1 3034656329 1961-11-11 ANSP H. G. Loftin & E. L. Tyson & Canipe Panama El Maria on Sona-Remedios Creek Quebrada Piedra 19 100 ANSP 104408, 4, 63.3, 116.3 1 Veraguas 3034656345 1961-10-28 H. G. Loftin & Tyson Panama 21 102 3 mi. E of Sona-Santiago 19 100 1 3034656364 1961-12-09 ANSP Dr. Alvarez, H. G & Loftin & E. L. Tyson Panama San Lorenzo Rio Jacaque 21 102 ANSP 119934, 2 1 Chiriqui 3034656377 1961-11-18 ANSP David, H. G & Loftin & E. L. Tyson Panama Creek 21 102 ANSP 119935, 2 1 3034656425 1961-11-18 ANSP H. G. Loftin & E. L. Tyson Panama Interamerican Highway San Felix 21 102 ANSP 11936, 18 1 3034656404 1961-11-11 H. G. Loftin & E. L. Tyson & Canipe Panama Rio Tinta 21 102 1 3034656338 1981-03-04 D. Fromm & P. Fromm Panama Rio Las Vueltas 21 102 1 3034656359 ROM 88191 1973-07-07 S. E. Meek & R. K. Johnson Costa Rica Rio Grande Terraba Jesus Maria River 21 102 1 Puntarenas 3034656388 1991-05-01 B. Wisenden Costa Rica 8.965 Osa Peninsula 21 -83.46222 Quebrada Benjamin 21 102 1 Puntarenas 3034656365 1983-07-13 K. Winemiller Costa Rica Rio Pavo 8.508056 Corcovado 21 -83.60083 Los Patos 21 102 1 Puntarenas 3034656397 1973-07-02 C. Gilbert Costa Rica Estarillos Rio Chires 21 102 1 Puntarenas 3034656363 1989-03-11 D. Fromm Costa Rica Rio Baru 21 102 1 San Jose 3034656340 1973-07-04 Gilbert & McAllister & Balon Costa Rica Rio Corobia 21 102 1 Guanacaste 3034656436 1969-02-08 Martin & Brame Honduras Rio Nacaome 21 102 1 Valle 3034656308 TU 202696 1962-06-02 de Fonseca & Pan American Hwy, R. D & Sutkus & Smalley El Salvador Gulfo de Fonseca 21 102 1 La Union 3034656440 1963-02-24 Carretera Internacional Del Pacifico, D.E & Rosen, K & R. Dorion Guatemala Carretera Internacional Del Pacifico Rio Margarita 21 102 1 Santa Rosa 3034656421 [199,1210,1969,1996] 1903-02-29 San Geronimo, S. E & Meek Mexico Oaxaca 21 102 1 Oaxaca 3034656328 1957-03-21 R. R. Miller & M. Miller Mexico 22 103 Rio Coyaquilla 21 102 1 Guerrero 3034656407 1955-02-24 R. R. Miller & J. T. Greenbank Mexico La Lima Rio La Lima 22 103 1 Nayarit 3034656306 1957-04-08 Miller & M. Miller Mexico Rio Rosario Rosario Hwy. 22 103 1 Nayarit 3034656408 CAS, SU Vaslit Mexico Eisen San Jose del Cabo 22 103 1 Baja California Sur