Motschulsky, 1863: 447
Pachysternum nigrovittatum : Knisch (1921: 88, morphological note) ; Knisch (1924: 159, catalogue) ; Orchymont (1926a: 222, 227, identification key, notes on variation) ; Orchymont (1928: 84, catalogue) ; Satô (1979: 50, faunistics) ; Biswas & Mukopadhyay (1995: 154, faunistics) ; Hebauer (2000: 7, faunistics) ; Mukhopadhyay & Sengupta (2003: 38, faunistics) ; Hebauer & Ryndevich (2005: 50, faunistics) ; Mukhopadhyay (2007: 408, faunistics) .
Megasternum nigrovittatum (Motschulsky)
Sharp (1890: 358)
A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)
Fikáček, Martin
Jia, Fenglong
Prokin, Alexander
Zootaxa
2012
2012-03-02
3219
1
53
4BV5D
Motschulsky, 1863
Motschulsky
1863
[151,735,786,812]
Insecta
Hydrophilidae
Pachysternum
GBIF
Animalia
Coleoptera
33
34
Arthropoda
species
nigrovittatum
( Figs. 1–3, 6–8, 12, 15, 20, 22–25, 28, 32, 46, 62)
Pachysternum nigrovittatum Motschulsky, 1863: 447. Pachysternum nigrovittatum: Knisch (1921: 88, morphological note); Knisch (1924: 159, catalogue); Orchymont (1926a: 222, 227, identification key, notes on variation); Orchymont (1928: 84, catalogue); Satô (1979: 50, faunistics); Biswas & Mukopadhyay (1995: 154, faunistics); Hebauer (2000: 7, faunistics); Mukhopadhyay & Sengupta (2003: 38, faunistics); Hebauer & Ryndevich (2005: 50, faunistics); Mukhopadhyay (2007: 408, faunistics). Megasternum nigrovittatum(Motschulsky): Sharp (1890: 358).
Typelocality. Sri Lanka, a mountain in the environs of Nuwara Eliya (“Montagne de Nura-Ellia”). Typematerial. Lectotype(hereby designated): 1 female( ZMUM): “[tiny red piece of paper] // [small rounded yellow label] // Type [small white label, in handwritten] // Pachysternum/ nigrovittatum/ Motsch. / Ind. or. Ceyl. M t. N. [yellow label, in handwritten]”. Paralectotype: 1 spec. [only head, prothorax and a mesothoracic leg preserved] (ZMUM): same label data as lectotype. On our request we have received two above mentioned specimens from the Motschulsky collection in ZMUM; both were glued on the same paper label. There is no indication of the number of syntypesin the original description, but the given typelocality (“Montagne de Nura-Ellia et sur le continent indien”) indicates that Motschulsky (1863)examined also the specimens from other localities than Sri Lankawhen describing the species. Therefore, some syntypesseem to be missing in his collection. In the present paper we are designating the complete specimen mentioned above as the lectotype, the remaining incomplete specimens as well as the remaining specimens not available to us have to be considered as paralectotypes. Additional material examined. CHINA: Fujian:48 spec. ( SYSU): Nanjing County, Hexitown [coordinates ca. 24°52'N 117°14'E], cow dung, 13.vii.2010, lgt. F. L. Jia. Guangdong: 1 spec.( BMNH): Guangzhou, Baiyunshan[ 23°9'N 113°17'E], 27.ix.1980, lgt. P. M. Hammond; 17 spec., ( SYSU): Sanshuicounty, Lubaotown [ 23°21'13''N, 112°53'46''E], 28.v.2004, lgt. F. L. Jia; 14 spec.( SYSU): Gaomingcounty, Yangmeitown, 23– 26.iv.2006, lgt. F.L. Jia; 1 spec.( SYSU): Lianjiang County, Wuling, 29.ii.1985, lgt. Wu Wu; 2 spec.( SYSU): Xinyi County, 9.5.1986, lgt. Wu Wu; 13 spec.( SYSU): Xinyi Cattle Farm, 7–13.vi.1986, lgt. Wu Wu; 9 spec.( SYSU): Guangzhou, Shipai Cattle Farm, 25.viii.1985, lgt. Wu Wu; 4 spec.( SYSU): Guangzhou, Henan Cattle Farm, 10.x.1985, lgt. Wu Wu. Guangxi: 5 spec.( BMNH): Bama [ 24°10'14''N 107°14'56''E], i.1988, without collector; 5 spec.( SYSU): Wuzhou, viii.1986, lgt. Wu Wu. Hainan: 1 female( NHMW): 2 kmSMao’an, elev. 250 m[ 19°8'39.84''N 108°56'48.84''E], 20.i.1996, lgt. M. A. Jäch. Hong Kong: 4 spec.( BMNH, NHMW): Hong Kong[ca. 22°24'N 114°8'E], without date, lgt. J. J. Walker; 4 spec. ( NHMW): New Territories, Luk Keng marsh near Kai Kuk Shue Ha[ 22°31'28''N 114°13'10''E], 14.i.1996, lgt. M. A. Jäch; 3 spec.( NHMW): University Campus [= campus of the University of Hong Kong, ca. 22°17'N 114°8'E], 1996, lgt. G. de Rougemont. Zhejiang: 1 spec.( BMNH): Chusan Is.[= Zhoushan, ca. 30°2'N, 122°7'E], without date, lgt. J. J. Walker. INDIA: Arunachal Pradesh: 1 female( NMPC): 3.6 kmNW Bhalukpong, elev. 280 m, 27°1'57''N 92°36'24''E, 7.v.2008, lgt. Fikáček, Podskalská, Šípek(16). Assam: 2 spec.( BMNH): Sudiya[ 27°50'N 95°45'E], without date and collector. Goa:6 spec. ( NMPC): 30 kmS of Margao[= Madgaon], Palolemenv., elev. 0–30 m, 15°0'47''N 74°1'58''E, 4–12.viii.2002, lgt. P. Šípek& M. Fikáček. Karnataka: 1 spec.( BMNH): Belgaum[ 15°51'N 74°33'0''E], without date, lgt. Andrewes & Bequest. Kerala: 1 spec.( NHMW): 35 kmNNE Trivandrum, Ponmudi, elev. 600 m[ 8°46'N 77°7'E], 2.i.1999, lgt. D. Boukal; 1 spec.( BMNH): “ Malabar” [without specified locality], without date and collector. Puducherry:1 spec. ( FMNH): Karaikal, Kurumbagatam[ 10°55'N 79°49'E], ii.1947, lgt. Susai Nathan. Tamil Nadu: 3 spec.( FMNH): Negapatam[= Nagappattinam, 10°46'N 79°50'E], without date, lgt. F. Psota; 40 spec.( BMNH): Nilgiri Hills[without specified locality], without date, lgt. H. L. Andrewes; 2 spec.( IRSNB): Shempagonur[= Shenbaganur, 6 kmof Kodaikanal, ca. 10°14'N 77°30'E], without date and collector. Uttarakhand: 1 spec.( BMNH): Akeswar, S Garhwal, Gauri Dutt. [ 29°48'N 78°37'E], 23.xi.1923, without collector; 2 spec.( IRSNB): Dehradun, Dobhalwala[ 30°20'N 78°2'37''E], without date and collector; 6 spec.( BMNH): W Almora, [ca. 29°36'N 79°34'E], without date, lgt. H. G. Champion. INDONESIA: East Nusa Tenggara:5 spec. ( NMPC): Sumba, 20 kmS of Waingapu, Wairinding[ 9°51'5''S 120°18'32''E], 30.i.–2.ii.2001, lgt. P. Votruba. North Sumatra: 1 spec.( NHMW): Prapat[= Parapat] env., elev. 1000 m[ 2°39'46''N 98°56'7''E], 17.ii.1990, lgt. S. Schödl. West Sumatra: 2 spec.( BMNH): Fort de Kock[= Bukittinggi, 0°18'20''S 100°22'9''E], elev. 920 m, x.1922, lgt. E. Jacobson; 5 spec.( IRSNB): same locality, 1925, lgt. E. Jacobson. LAOS: Bolikhamxai: 1 spec.( NHMB): 70 kmNEE Vientiane, elev. 150 m, 18°16'N, 103°11'E, 27– 30.iv.1997, lgt. V. Kubáň; 28 spec.( NHMB, NMPC): 8 kmNE Ban Nape, elev. 600 m, 18°21'N, 105°8'E, 1– 18.v.2001, lgt. V. Kubáň; 1 spec. ( FHGC): Ban Nape, Kaew Nua P. [ 18°18'7''N 105°4'24''E], 18.iv.–1.v.1998, lgt. E. Jendek & O. Šauša. Houaphan: 2 spec.( NMPC): Phou Pane Mt., elev. 1480–1510 m, 20°13'9''N 103°59'54''E, 1– 16.vi.2009, lgt. V. Kubáň. Louangphabang: 1 spec.( NHMB): Thong Khan, elev. 750 m, 19°35'N 101°58'E, 11– 21.v.2002, lgt. V. Kubáň. Xiangkhouang: 2 spec.( NMPC): Phou Sane Mt., 30 kmNE Phonsavan, elev. 1420 m, 19°38.2'N 103°20.2'E, 10–30.v.2009, lgt. V. Kubáň. MALAYSIA: Kedah: 3 spec. ( ASKC): Pulau Langkawi, environs of hotel Berjaya[= Pantai Kokvillage, 6°22'N, 99°40.5'E], 7–22.xi.2009, lgt. U. Schmidt. Melaka: 2 spec.( BMNH): Malacca[ 2°11'53''N 102°14'59''E], without date, lgt. Wallace. Pahang: 1 spec.( BMNH): Taman Negara[= Taman Negara National Park, ca. 4°41'N 102°34'E], 1–13.iii.1984, lgt. L. Jessop. Perak: 1 spec.( BMNH): “ Perak” [without specified locality], without date and collector. NEPAL: Gandaki:1 spec. ( FHGC): Sikles Mts., überh. Garlang, elev. 2000 m[ 28°19'N 84°0'E], 29.vii.1995, lgt. Fabrizi, Schmidt, Jäger. SINGAPORE: 8 spec.( BMNH, IRSNB): Singapore[ 1°21'N 103°50'E], without date and collector. SRI LANKA: 2 spec.( BMNH): “ Ceylon” [without specified locality], without date and collector; 1 spec.( IRSNB): same locality, without date, lgt. F. Chapuis. Sabaragamuwa: 1 spec.( BMNH): Kitulgalle[= Kitulgala, 6°59'30''N 80°26'54''E], without date, lgt. C. Lewis. Uva: 1 spec.( FHGC): 2 milesNW Haldummulla, elev. 1097 m[ 6°47'26''N 80°51'53''E], 2.iii.1962, lgt. Brinck, Anderson & Cederholm(111). Western Province:5 spec. ( BMNH): Colombo [ 6°55'N 79°50'30''E], 14–15.xi.1915, lgt. M. Cameron; 1 spec.( FHGC): lagoon 17 milesN of Colombo[ca. 6°45'N 79°54'E], 11.i.1962, lgt. Brinck, Anderson & Cederholm(8); 1 spec.( FHGC): Yakkala, 18 milesNE Colombo[ 7°5'25''N 80°2'2''E], 15–21.i.1962, lgt. Brinck, Anderson & Cederholm(11). THAILAND: Uthai Thani: 1 spec. ( NHMW): 25 kmNW Lan Sak, 240 kmN of Bangkok[ca. 15°34'N 99°31'E], iii.1989, lgt. Thielen. VIETNAM: Bac Thai: 1 spec.( NMPC): Tam Dao[ National Park, ca. 21°38'N, 105°30'E], without date, lgt. P. Marhoul. 1 spec.( IRSNB): “ Tonkin, Foret de Kep”, without date and collector. Gia Lai: 20 spec.( NHMW): 40 kmNW An Khe, Buon Luoi, elev. 620–750 m, 14°10'N 108°30'E, 28.iii.–12.iv.1995, lgt. Pacholátko & Dembický. Hoa Binh: 1 spec.( IRSNB): “ Chobo” [= Hoa Binhprovince, without precise locality], without date, lgt. H. Perrot; 1 spec.( BMNH): Hoa Binh[ca. 20°50'N 105°20'E], viii.1918, lgt. R. V. de Salvaza; 36 spec., 1 male( IRSNB): same locality, without date, lgt. A. de Cooman. Lam Dong: 1 spec.( NHMW): Da Latcity [ 11°56'N 108°27'E], 27–21.iv.1994, lgt. Pacholátko & Dembický. Nghe An: 3 spec.( BMNH): Lactho[= Lac Tho, 21°4'N 106°6'E], without date, lgt. de Cooman; 66 spec.( IRSNB): same locality, without date and collector. Published records (not examined). INDIA: Andhra Pradesh:Kurnool district, Betamcherla; Prakasan district, Matur [= Martur] (both Mukhopadhyay 2007). Manipur:Andro, Imphal ( Mukhopadhyay & Sengupta 2004). Sikkim:Mongan ( Mukhopadhyay & Sengupta 2003). West Bengal:without detailed localities ( Mukhopadhyay 2007). LAOS: Loung Namtha:Louang Namtha env. ( Hebauer & Ryndevich 2005). THAILAND: Chiang Mai: Chiang Mai ( Hebauer & Ryndevich 2005).
Diagnosis.The species may be very easily distinguished from all remaining Asian Pachysternumby the combination of pale pronotum with large M-shape spot, elytra pale with at least some of elytral intervals 2–10 completely dark ( Fig. 62), and deeply bisinuate outer margin of anterior tibia ( Fig. 46). By the coloration of pronotum it resembles P. cardoniwhich is much larger and wider and never bears the combination of entirely pale elytral interval 2 and largely dark interval 4, and the apical portion of its median lobe is gradually narrowing apicad (narrow and nearly parallel-sided in P. nigrovittatum, Fig. 32b). The elytral coloration of P. nigrovittatummay resemble pale specimens of P. sulawesicum, but may be easily distinguished from it by deeply bisinuate outer margin of anterior tibiae (continually arcuate in P. sulawesicum) and distinctly narrowed apical portion of the median lobe (nearly parallel-sided throughout in P. sulawesicum).
Redescription(based on specimens from Sri Lanka). Body elongate oval, shape of elytra sexually dimorphic, female with very distinctly pronounced humeral portion of elytra. Body length 2.2–3.0 mm ( lectotype: 2.3 mm); body width 1.4–2.0 mm ( lectotype 1.6 mm). Coloration( Fig. 62). Head brown widely pale reddish median portion of frons and transverse ridge. General coloration of pronotum pale reddish, bearing M-shaped black spot of variable extent, leaving wide lateral portions, anteromedian part and a pair of large posterior submedian parts reddish; posterior margin black. Elytra pale reddish brown, with alternating pale and black intervals as follows: interval 1 entirely black, interval 2 entirely pale, interval 3 pale except of short black portion in anterior third, interval 4 black in anterior four fifths, pale apically, interval 5 entirely pale, interval 6 similar to 4, but with short pale area subbasally, intervals 7–9 pale subbasally (forming pale humeral spot), otherwise black except for apical-most portion, Interval 10 (lateralmost one) pale. Ventral surface of head and prothorax reddish brown, meso- and metathorax dark brown, abdominal ventrites 1–4 largely pale reddish posteriorly, ventrite 5 entirely pale reddish. Legs pale reddish. External morphology. Clypeus with fine and rather sparse punctation consisting of slightly semicircular punctures of two slightly different sizes separated by ca. 1–2× puncture width. Punctation of frons similar to that on clypeus, only with slightly larger punctures. Interstices on head without microsculpture. Larger punctures on pronotum semicircular, deeply impressed, much larger than small punctures; small punctures transverse, divided from each other by 2–3× puncture width. Pronotal interstices without microsculpture. Prosternum with distinct median carina. Elytral series consisting of large, shallow punctures; interval punctation with very fine semicircular punctures much smaller than serial ones, with sparsely intermixed slightly larger setiferous punctures of the same shape; interstices with weakly developed mesh-like microsculpture. Elytral series weakly impressed. Preepisternal elevation of mesothorax without posterolateral pits. Femoral lines on metaventrite indistinctly angulate, median portion of metaventrite bearing moderately coarse and dense setiferous punctation, interstices without microsculpture; lateral portions with very coarse and dense, densely pubescent punctation. Anterior tibia rather narrow, outer margin deeply bisinuate, outer series of spines interrupted at the place of the sinuation. Malegenitalia( Fig. 32). Tegmen 1.3–1.4 mmlong, median lobe 1.3–1.4 mmlong. Phallobase ca. 2× as long as parameres, basally slightly asymmetrical but without detached manubrium. Median lobe widest in basal third, weakly narrowing to apical third, narrow and nearly parallel-sided in apical third, apex rounded; gonopore subapical, rather indistinct; lateral pubescent lobes distinct only subapically. Sternite 9 narrow, without V-shaped apical emargination. Variation.Rather variable species, with the variation concerning mainly the coloration, dorsal microsculpture, body size and slightly also the extent of sexual dimorphism in body shape. Pale specimens ( Fig. 62c) have largely reddish elytra with dark T-shape spot (consisting of dark elytral interval 1 and bark base of elytra) and dark spots at midlength of intervals 4 and 6–9 and subbasally on intervals 6–7. The extent of dark area is increasing in other specimens through the most typical coloration mentioned in the redescription ( Figs. 62a–b), to dark specimens with largely black elytra and very apparent reddish spots at bases and in apical portions of intervals 2, 3, 5 and 8– 10 and narrowly pale lateral margin of elytron ( Fig. 62g). The coloration of pronotum varies in the same extent, bearing a median and a pair of submedian dark spots in palest specimens ( Fig. 62c), the dark M-shaped spot in most specimens ( Figs. 62a–b, d–e) and a being largely dark at pronotal disc with only a weakly developed pale spots submedially in dark specimens ( Figs. 62f–g). Dorsal microsculpture usually consists of rather weak meshlike microsculpture which is often rather inconspicuous, but in extremely dark and/or small specimens the microsculpture is often very distinct. The body shape of males is constant in all specimens examined, but the shape of females may vary from rather short and wide specimens with weakly pronounced shoulders to very distinctly elongate specimens with highly pronounced shoulders and elytra strongly narrowing apicad (the most elongate females are known from the island of Sumba). The shape of the anterior tibia as well as of the male genitalia (including the median lobe and sternite 9) is very constant in all examined specimens irrespectively to their variation in the above characters, which we therefore consider as intraspecific.
Biology.Coprophagous species inhabiting excrements of various herbivorous mammals.
Distribution.Widespread throughout the Oriental Region from Nepaland Sri Lankathrough India, whole continental SE Asia and some of the islands of Sundaland (Sumatra, Sumba). Records are surprisingly missing for whole lowland northern part of India(and P. nigrovittatumseems therefore be absent from semidesert areas) and from many islands of Sundaland (in this case most probably because of the collecting bias).
3499776350
ZMUM
33
34
1
1
lectotype
3499776404
2010-07-13
SYSU
China
Nanjing County
24.866667
Hexi
1249
117.23333
33
34
1
3499776478
1980-09-27
BMNH
P. M. Hammond
23.15
Guangzhou
1257
113.28333
Guangdong
33
34
1
3499776325
2004-05-28
SYSU
F. L. Jia
23.353611
Lubao
21
112.89611
Sanshui
33
34
17
3499776384
2006-04-23
2006-04-26
2006-04-23
SYSU
F. L. Jia
Yangmei
Gaoming
33
34
14
3499776373
[422,1177,1647,1671]
1985-02-29
SYSU
Lianjiang County
Wu Wu
Wuling
33
34
1
3499776424
1986-05-09
SYSU
Xinyi County
Wu Wu
Wu Wu
33
34
2
3499776450
[508,1241,1685,1709]
1986-06-07
1986-06-13
1986-06-07
SYSU
Wu Wu
33
34
13
3499776368
1985-08-25
SYSU
Guangzhou
33
34
9
3499776441
1985-10-10
SYSU
Guangzhou
33
34
4
3499776385
1986-08
1988-01
1986-08
BMNH, SYSU
24.170555
Wu Wu
21
107.248886
33
34
10
3499776443
1996-01-20
NHMW
M. A. Jach
250
19.1444
Hainan
1
108.9469
33
34
1
1
3499776364
BMNH, NHMW
J. J. Walker
Hong Kong
22.4
Hong Kong
1260
114.13333
33
34
4
3499776313
1996-01-14
NHMW
M. A. Jach
Cook Islands
22.524445
Shue Ha
21
114.219444
33
34
1
3499776369
NHMW
G. de Rougemont
Hong Kong
22.283333
Hong Kong
1260
114.13333
33
34
3
3499776431
BMNH
J. J. Walker
30.033333
Chusan Is.
1224
122.11667
Zhejiang
33
34
1
3499776301
2008-05-07
NMPC
India
280
34
35
27.0325
Bhalukpong
20
92.60667
Arunachal Pradesh
33
34
1
1
3499776370
[414,1291,189,214]
BMNH
27.833334
Sudiya
1235
95.75
Assam
34
35
2
3499776352
2002-08-04
2002-08-12
2002-08-04
NMPC
India
15
15.013056
Madgaon
21
74.03278
Margao
34
35
1
3499776432
BMNH
15.85
Belgaum
925
74.55
Karnataka
34
35
1
3499776470
1999-01-02
NHMW
D. Boukal
600
8.766666
Trivandrum
1300
77.11667
Kerala
34
35
1
3499776315
[247,1191,338,362]
BMNH
Malabar
34
35
1
3499776359
1947-02
FMNH
India
10.916667
Kurumbagatam
1296
79.816666
34
35
1
3499776330
FMNH
F. Psota
10.766666
Negapatam
1297
79.833336
Tamil Nadu
34
35
3
3499776401
BMNH
H. L. Andrewes
Nilgiri Hills
34
35
40
3499776475
IRSNB
10.233334
Shenbaganur
1298
77.5
Shempagonur
34
35
2
3499776366
1923-11-23
BMNH
29.8
Akeswar
1225
78.61667
Uttarakhand
34
35
1
3499776416
IRSNB
30.333334
Dobhalwala
925
78.04361
Dehradun
34
35
2
3499776390
BMNH
H. G. Champion
29.6
Almora
1226
79.566666
34
35
6
3499776338
2001-01-30
2001-02-02
2001-01-30
NMPC
P. Votruba
Indonesia
-9.851389
Sumba
21
120.30889
East Nusa Tenggara
34
35
1
3499776434
1990-02-17
NHMW
S. Schodl
1000
2.662778
Prapat
21
98.93528
North Sumatra
34
35
1
3499776354
1922-10
BMNH
E. Jacobson
920
-0.30555558
Fort de Kock
21
100.36917
West Sumatra
34
35
2
3499776411
IRSNB
E. Jacobson
34
35
5
3499776462
1997-04-27
1997-04-30
1997-04-27
NHMB
V. Kuban
Laos
150
18.266666
Bolikhamxai
1275
103.183334
Bolikhamxai
34
35
1
3499776314
2001-05-01
2001-05-18
2001-05-01
NHMB, NMPC
V. Kuban
600
18.35
1275
105.13333
34
35
28
3499776429
1998-04-18
1998-05-01
1998-04-18
FHGC
Iceland
18.301943
Sausa
21
105.07333
34
35
1
3499776311
2009-06-01
2009-06-16
2009-06-01
NMPC
V. Kuban
1495
20.219168
Phou Pane Mt.
21
103.99833
Houaphan
34
35
2
3499776460
2002-05-11
2002-05-21
2002-05-11
NHMB
V. Kuban
750
19.583334
Thong Khan
1271
101.96667
Louangphabang
34
35
1
3499776406
2009-05-10
2009-05-30
2009-05-10
NMPC
V. Kuban
1420
19.636667
Phou Sane Mt.
127
103.33667
Xiangkhouang
34
35
2
3499776383
2009-11-07
2009-11-22
2009-11-07
ASKC
U. Schmidt
Malaysia
6.366667
Langkawi
929
99.675
Kedah
34
35
1
3499776445
BMNH
Malaysia
2.1980555
Wallace
21
102.24972
34
35
2
3499776348
1984-03-01
1984-03-13
1984-03-01
BMNH
L. Jessop
4.6833334
Taman Negara
1306
102.566666
Pahang
34
35
1
3499776312
BMNH
Perak
Perak
34
35
1
3499776316
1995-07-29
FHGC
Nepal
2000
28.316668
Sikles Mts.
1232
84.0
Gandaki
34
35
1
3499776480
BMNH, IRSNB
Singapore
1.35
SINGAPORE
1308
103.833336
34
35
8
3499776469
BMNH
Sri Lanka
SRI LANKA
34
35
2
3499776339
[698,1363,1234,1258]
IRSNB
F. Chapuis
34
35
1
3499776335
BMNH
C. Lewis
6.991667
Kitulgalle
21
80.44833
Sabaragamuwa
34
35
1
3499776457
1962-03-02
FHGC
1097
6.7905555
Haldummulla
21
80.86472
Uva
34
35
1
3499776464
1915-11-14
1915-11-15
1915-11-14
BMNH
Cameroon
6.9166665
Cameron
925
79.84167
34
35
1
3499776345
1962-01-11
FHGC
6.75
Brinck
1304
79.9
Colombo
34
35
1
3499776451
1962-01-15
1962-01-21
1962-01-15
FHGC
7.090278
Brinck
21
80.03389
Colombo
34
35
1
3499776317
1989-03
NHMW
Thailand
15.566667
Lan Sak
1284
99.51667
Uthai Thani
34
35
1
3499776357
NMPC
P. Marhoul
Vietnam
21.633333
Tam
1263
105.5
Bac
34
35
1
3499776403
IRSNB
Vietnam
Foret de Kep
34
35
1
3499776437
1995-03-28
1995-04-12
1995-03-28
NHMW
685
14.166667
An Khe
1289
108.5
Gia Lai
34
35
20
3499776402
IRSNB
H. Perrot
Chobo
Hoa Binh
34
35
1
3499776340
1918-08
BMNH
R. V. de Salvaza
20.833334
Hoa Binh
1266
105.333336
34
35
37
3499776310
[385,1093,1682,1707]
IRSNB
A. de Cooman
34
35
1
1
3499776341
1994-04-21
1994-04-27
1994-04-21
NHMW
11.933333
Da Lat
1294
108.45
Lam Dong
34
35
1
3499776474
BMNH
21.066668
Lactho
1265
106.1
Nghe An
34
35
3
3499776323
IRSNB
34
35
66