Motschulsky, 1863: 447 Pachysternum nigrovittatum : Knisch (1921: 88, morphological note) ; Knisch (1924: 159, catalogue) ; Orchymont (1926a: 222, 227, identification key, notes on variation) ; Orchymont (1928: 84, catalogue) ; Satô (1979: 50, faunistics) ; Biswas & Mukopadhyay (1995: 154, faunistics) ; Hebauer (2000: 7, faunistics) ; Mukhopadhyay & Sengupta (2003: 38, faunistics) ; Hebauer & Ryndevich (2005: 50, faunistics) ; Mukhopadhyay (2007: 408, faunistics) . Megasternum nigrovittatum (Motschulsky) Sharp (1890: 358) A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae) Fikáček, Martin Jia, Fenglong Prokin, Alexander Zootaxa 2012 2012-03-02 3219 1 53 4BV5D Motschulsky, 1863 Motschulsky 1863 [151,735,786,812] Insecta Hydrophilidae Pachysternum GBIF Animalia Coleoptera 33 34 Arthropoda species nigrovittatum  ( Figs. 1–3, 6–8, 12, 15, 20, 22–25, 28, 32, 46, 62)     Pachysternum nigrovittatum  Motschulsky, 1863: 447.     Pachysternum nigrovittatum: Knisch (1921: 88, morphological note); Knisch (1924: 159, catalogue); Orchymont (1926a: 222, 227, identification key, notes on variation); Orchymont (1928: 84, catalogue); Satô (1979: 50, faunistics); Biswas & Mukopadhyay (1995: 154, faunistics); Hebauer (2000: 7, faunistics); Mukhopadhyay & Sengupta (2003: 38, faunistics); Hebauer & Ryndevich (2005: 50, faunistics); Mukhopadhyay (2007: 408, faunistics).    Megasternum nigrovittatum(Motschulsky):  Sharp (1890: 358).    Typelocality. Sri Lanka, a mountain in the environs of Nuwara Eliya (“Montagne de Nura-Ellia”).   Typematerial.  Lectotype(hereby designated): 1 female( ZMUM): “[tiny red piece of paper] // [small rounded yellow label] // Type [small white label, in handwritten] // Pachysternum/ nigrovittatum/ Motsch. / Ind. or. Ceyl. M t. N. [yellow label, in handwritten]”. Paralectotype: 1 spec. [only head, prothorax and a mesothoracic leg preserved] (ZMUM): same label data as lectotype. On our request we have received two above mentioned specimens from the Motschulsky collection in ZMUM; both were glued on the same paper label. There is no indication of the number of syntypesin the original description, but the given typelocality (“Montagne de Nura-Ellia et sur le continent indien”) indicates that Motschulsky (1863)examined also the specimens from other localities than Sri Lankawhen describing the species. Therefore, some syntypesseem to be missing in his collection. In the present paper we are designating the complete specimen mentioned above as the lectotype, the remaining incomplete specimens as well as the remaining specimens not available to us have to be considered as paralectotypes.  Additional material examined.   CHINA: Fujian:48 spec. ( SYSU): Nanjing County, Hexitown [coordinates ca. 24°52'N 117°14'E], cow dung,  13.vii.2010, lgt. F. L. Jia.   Guangdong: 1 spec.( BMNH): Guangzhou, Baiyunshan[ 23°9'N 113°17'E],  27.ix.1980, lgt. P. M. Hammond;  17 spec., ( SYSU): Sanshuicounty, Lubaotown [ 23°21'13''N, 112°53'46''E],  28.v.2004, lgt. F. L. Jia;  14 spec.( SYSU): Gaomingcounty, Yangmeitown,  23– 26.iv.2006, lgt. F.L. Jia;  1 spec.( SYSU): Lianjiang County, Wuling,  29.ii.1985, lgt. Wu Wu;  2 spec.( SYSU): Xinyi County,  9.5.1986, lgt. Wu Wu;  13 spec.( SYSU): Xinyi Cattle Farm,  7–13.vi.1986, lgt. Wu Wu;  9 spec.( SYSU): Guangzhou, Shipai Cattle Farm,  25.viii.1985, lgt. Wu Wu;  4 spec.( SYSU): Guangzhou, Henan Cattle Farm,  10.x.1985, lgt. Wu Wu.  Guangxi: 5 spec.( BMNH): Bama [ 24°10'14''N 107°14'56''E],  i.1988, without collector; 5 spec.( SYSU): Wuzhou,  viii.1986, lgt. Wu Wu.   Hainan: 1 female( NHMW):  2 kmSMao’an, elev.  250 m[ 19°8'39.84''N 108°56'48.84''E],  20.i.1996, lgt. M. A. Jäch.   Hong Kong: 4 spec.( BMNH, NHMW): Hong Kong[ca. 22°24'N 114°8'E], without date, lgt. J. J. Walker; 4 spec. ( NHMW): New Territories, Luk Keng marsh near Kai Kuk Shue Ha[ 22°31'28''N 114°13'10''E],  14.i.1996, lgt. M. A. Jäch;  3 spec.( NHMW): University Campus [= campus of the University of Hong Kong, ca. 22°17'N 114°8'E], 1996, lgt. G. de Rougemont.   Zhejiang: 1 spec.( BMNH): Chusan Is.[= Zhoushan, ca. 30°2'N, 122°7'E], without date, lgt. J. J. Walker.   INDIA: Arunachal Pradesh: 1 female( NMPC):  3.6 kmNW Bhalukpong, elev.  280 m, 27°1'57''N 92°36'24''E,  7.v.2008, lgt. Fikáček, Podskalská, Šípek(16).   Assam: 2 spec.( BMNH): Sudiya[ 27°50'N 95°45'E], without date and collector.   Goa:6 spec. ( NMPC):  30 kmS of Margao[= Madgaon], Palolemenv., elev.  0–30 m, 15°0'47''N 74°1'58''E,  4–12.viii.2002, lgt. P. Šípek& M. Fikáček.   Karnataka: 1 spec.( BMNH): Belgaum[ 15°51'N 74°33'0''E], without date, lgt. Andrewes & Bequest.   Kerala: 1 spec.( NHMW):  35 kmNNE Trivandrum, Ponmudi, elev.  600 m[ 8°46'N 77°7'E],  2.i.1999, lgt. D. Boukal;  1 spec.( BMNH): “ Malabar” [without specified locality], without date and collector.   Puducherry:1 spec. ( FMNH): Karaikal, Kurumbagatam[ 10°55'N 79°49'E],  ii.1947, lgt. Susai Nathan.   Tamil Nadu: 3 spec.( FMNH): Negapatam[= Nagappattinam, 10°46'N 79°50'E], without date, lgt. F. Psota;  40 spec.( BMNH): Nilgiri Hills[without specified locality], without date, lgt. H. L. Andrewes;  2 spec.( IRSNB): Shempagonur[= Shenbaganur,  6 kmof Kodaikanal, ca. 10°14'N 77°30'E], without date and collector.   Uttarakhand: 1 spec.( BMNH): Akeswar, S Garhwal, Gauri Dutt. [ 29°48'N 78°37'E],  23.xi.1923, without collector;  2 spec.( IRSNB): Dehradun, Dobhalwala[ 30°20'N 78°2'37''E], without date and collector;  6 spec.( BMNH): W Almora, [ca. 29°36'N 79°34'E], without date, lgt. H. G. Champion.   INDONESIA:  East Nusa Tenggara:5 spec. ( NMPC): Sumba,  20 kmS of Waingapu, Wairinding[ 9°51'5''S 120°18'32''E],  30.i.–2.ii.2001, lgt. P. Votruba.   North Sumatra: 1 spec.( NHMW): Prapat[= Parapat] env., elev.  1000 m[ 2°39'46''N 98°56'7''E],  17.ii.1990, lgt. S. Schödl.   West Sumatra: 2 spec.( BMNH): Fort de Kock[= Bukittinggi, 0°18'20''S 100°22'9''E], elev.  920 m,  x.1922, lgt. E. Jacobson;  5 spec.( IRSNB): same locality, 1925, lgt. E. Jacobson.   LAOS: Bolikhamxai: 1 spec.( NHMB):  70 kmNEE Vientiane, elev.  150 m, 18°16'N, 103°11'E,  27– 30.iv.1997, lgt. V. Kubáň;  28 spec.( NHMB, NMPC):  8 kmNE Ban Nape, elev.  600 m, 18°21'N, 105°8'E,  1– 18.v.2001, lgt. V. Kubáň; 1 spec. ( FHGC): Ban Nape, Kaew Nua P. [ 18°18'7''N 105°4'24''E],  18.iv.–1.v.1998, lgt. E. Jendek & O. Šauša.   Houaphan: 2 spec.( NMPC): Phou Pane Mt., elev.  1480–1510 m, 20°13'9''N 103°59'54''E,  1– 16.vi.2009, lgt. V. Kubáň.   Louangphabang: 1 spec.( NHMB): Thong Khan, elev.  750 m, 19°35'N 101°58'E,  11– 21.v.2002, lgt. V. Kubáň.   Xiangkhouang: 2 spec.( NMPC): Phou Sane Mt.,  30 kmNE Phonsavan, elev.  1420 m, 19°38.2'N 103°20.2'E,  10–30.v.2009, lgt. V. Kubáň.   MALAYSIA:  Kedah: 3 spec. ( ASKC): Pulau Langkawi, environs of hotel Berjaya[= Pantai Kokvillage, 6°22'N, 99°40.5'E],  7–22.xi.2009, lgt. U. Schmidt.   Melaka: 2 spec.( BMNH): Malacca[ 2°11'53''N 102°14'59''E], without date, lgt. Wallace.   Pahang: 1 spec.( BMNH): Taman Negara[= Taman Negara National Park, ca. 4°41'N 102°34'E],  1–13.iii.1984, lgt. L. Jessop.   Perak: 1 spec.( BMNH): “ Perak” [without specified locality], without date and collector.   NEPAL: Gandaki:1 spec. ( FHGC): Sikles Mts., überh. Garlang, elev.  2000 m[ 28°19'N 84°0'E],  29.vii.1995, lgt. Fabrizi, Schmidt, Jäger.   SINGAPORE: 8 spec.( BMNH, IRSNB): Singapore[ 1°21'N 103°50'E], without date and collector.   SRI LANKA: 2 spec.( BMNH): “ Ceylon” [without specified locality], without date and collector;  1 spec.( IRSNB): same locality, without date, lgt. F. Chapuis.   Sabaragamuwa: 1 spec.( BMNH): Kitulgalle[= Kitulgala, 6°59'30''N 80°26'54''E], without date, lgt. C. Lewis.   Uva: 1 spec.( FHGC):  2 milesNW Haldummulla, elev.  1097 m[ 6°47'26''N 80°51'53''E],  2.iii.1962, lgt. Brinck, Anderson & Cederholm(111).  Western Province:5 spec. ( BMNH): Colombo [ 6°55'N 79°50'30''E],  14–15.xi.1915, lgt. M. Cameron;  1 spec.( FHGC): lagoon  17 milesN of Colombo[ca. 6°45'N 79°54'E],  11.i.1962, lgt. Brinck, Anderson & Cederholm(8);  1 spec.( FHGC): Yakkala,  18 milesNE Colombo[ 7°5'25''N 80°2'2''E],  15–21.i.1962, lgt. Brinck, Anderson & Cederholm(11).   THAILAND: Uthai Thani: 1 spec. ( NHMW):  25 kmNW Lan Sak,  240 kmN of Bangkok[ca. 15°34'N 99°31'E],  iii.1989, lgt. Thielen.   VIETNAM:  Bac Thai: 1 spec.( NMPC): Tam Dao[ National Park, ca. 21°38'N, 105°30'E], without date, lgt. P. Marhoul.  1 spec.( IRSNB): “ Tonkin, Foret de Kep”, without date and collector.   Gia Lai: 20 spec.( NHMW):  40 kmNW An Khe, Buon Luoi, elev.  620–750 m, 14°10'N 108°30'E,  28.iii.–12.iv.1995, lgt. Pacholátko & Dembický.   Hoa Binh: 1 spec.( IRSNB): “ Chobo” [= Hoa Binhprovince, without precise locality], without date, lgt. H. Perrot;  1 spec.( BMNH): Hoa Binh[ca. 20°50'N 105°20'E],  viii.1918, lgt. R. V. de Salvaza; 36 spec.,  1 male( IRSNB): same locality, without date, lgt. A. de Cooman.   Lam Dong: 1 spec.( NHMW): Da Latcity [ 11°56'N 108°27'E],  27–21.iv.1994, lgt. Pacholátko & Dembický.   Nghe An: 3 spec.( BMNH): Lactho[= Lac Tho, 21°4'N 106°6'E], without date, lgt. de Cooman;  66 spec.( IRSNB): same locality, without date and collector.  Published records (not examined). INDIA: Andhra Pradesh:Kurnool district, Betamcherla; Prakasan district, Matur [= Martur] (both Mukhopadhyay 2007). Manipur:Andro, Imphal ( Mukhopadhyay & Sengupta 2004). Sikkim:Mongan ( Mukhopadhyay & Sengupta 2003). West Bengal:without detailed localities ( Mukhopadhyay 2007).  LAOS: Loung Namtha:Louang Namtha env. ( Hebauer & Ryndevich 2005).  THAILAND: Chiang Mai: Chiang Mai ( Hebauer & Ryndevich 2005).   Diagnosis.The species may be very easily distinguished from all remaining Asian  Pachysternumby the combination of pale pronotum with large M-shape spot, elytra pale with at least some of elytral intervals 2–10 completely dark ( Fig. 62), and deeply bisinuate outer margin of anterior tibia ( Fig. 46). By the coloration of pronotum it resembles  P. cardoniwhich is much larger and wider and never bears the combination of entirely pale elytral interval 2 and largely dark interval 4, and the apical portion of its median lobe is gradually narrowing apicad (narrow and nearly parallel-sided in  P. nigrovittatum, Fig. 32b). The elytral coloration of  P. nigrovittatummay resemble pale specimens of  P. sulawesicum, but may be easily distinguished from it by deeply bisinuate outer margin of anterior tibiae (continually arcuate in  P. sulawesicum) and distinctly narrowed apical portion of the median lobe (nearly parallel-sided throughout in  P. sulawesicum).   Redescription(based on specimens from Sri Lanka). Body elongate oval, shape of elytra sexually dimorphic, female with very distinctly pronounced humeral portion of elytra. Body length 2.2–3.0 mm ( lectotype: 2.3 mm); body width 1.4–2.0 mm ( lectotype 1.6 mm).  Coloration( Fig. 62). Head brown widely pale reddish median portion of frons and transverse ridge. General coloration of pronotum pale reddish, bearing M-shaped black spot of variable extent, leaving wide lateral portions, anteromedian part and a pair of large posterior submedian parts reddish; posterior margin black. Elytra pale reddish brown, with alternating pale and black intervals as follows: interval 1 entirely black, interval 2 entirely pale, interval 3 pale except of short black portion in anterior third, interval 4 black in anterior four fifths, pale apically, interval 5 entirely pale, interval 6 similar to 4, but with short pale area subbasally, intervals 7–9 pale subbasally (forming pale humeral spot), otherwise black except for apical-most portion, Interval 10 (lateralmost one) pale. Ventral surface of head and prothorax reddish brown, meso- and metathorax dark brown, abdominal ventrites 1–4 largely pale reddish posteriorly, ventrite 5 entirely pale reddish. Legs pale reddish.  External morphology. Clypeus with fine and rather sparse punctation consisting of slightly semicircular punctures of two slightly different sizes separated by ca. 1–2× puncture width. Punctation of frons similar to that on clypeus, only with slightly larger punctures. Interstices on head without microsculpture. Larger punctures on pronotum semicircular, deeply impressed, much larger than small punctures; small punctures transverse, divided from each other by 2–3× puncture width. Pronotal interstices without microsculpture. Prosternum with distinct median carina. Elytral series consisting of large, shallow punctures; interval punctation with very fine semicircular punctures much smaller than serial ones, with sparsely intermixed slightly larger setiferous punctures of the same shape; interstices with weakly developed mesh-like microsculpture. Elytral series weakly impressed. Preepisternal elevation of mesothorax without posterolateral pits. Femoral lines on metaventrite indistinctly angulate, median portion of metaventrite bearing moderately coarse and dense setiferous punctation, interstices without microsculpture; lateral portions with very coarse and dense, densely pubescent punctation. Anterior tibia rather narrow, outer margin deeply bisinuate, outer series of spines interrupted at the place of the sinuation.   Malegenitalia( Fig. 32). Tegmen 1.3–1.4 mmlong, median lobe 1.3–1.4 mmlong. Phallobase ca. 2× as long as parameres, basally slightly asymmetrical but without detached manubrium. Median lobe widest in basal third, weakly narrowing to apical third, narrow and nearly parallel-sided in apical third, apex rounded; gonopore subapical, rather indistinct; lateral pubescent lobes distinct only subapically. Sternite 9 narrow, without V-shaped apical emargination.  Variation.Rather variable species, with the variation concerning mainly the coloration, dorsal microsculpture, body size and slightly also the extent of sexual dimorphism in body shape. Pale specimens ( Fig. 62c) have largely reddish elytra with dark T-shape spot (consisting of dark elytral interval 1 and bark base of elytra) and dark spots at midlength of intervals 4 and 6–9 and subbasally on intervals 6–7. The extent of dark area is increasing in other specimens through the most typical coloration mentioned in the redescription ( Figs. 62a–b), to dark specimens with largely black elytra and very apparent reddish spots at bases and in apical portions of intervals 2, 3, 5 and 8– 10 and narrowly pale lateral margin of elytron ( Fig. 62g). The coloration of pronotum varies in the same extent, bearing a median and a pair of submedian dark spots in palest specimens ( Fig. 62c), the dark M-shaped spot in most specimens ( Figs. 62a–b, d–e) and a being largely dark at pronotal disc with only a weakly developed pale spots submedially in dark specimens ( Figs. 62f–g). Dorsal microsculpture usually consists of rather weak meshlike microsculpture which is often rather inconspicuous, but in extremely dark and/or small specimens the microsculpture is often very distinct. The body shape of males is constant in all specimens examined, but the shape of females may vary from rather short and wide specimens with weakly pronounced shoulders to very distinctly elongate specimens with highly pronounced shoulders and elytra strongly narrowing apicad (the most elongate females are known from the island of Sumba). The shape of the anterior tibia as well as of the male genitalia (including the median lobe and sternite 9) is very constant in all examined specimens irrespectively to their variation in the above characters, which we therefore consider as intraspecific.   Biology.Coprophagous species inhabiting excrements of various herbivorous mammals.   Distribution.Widespread throughout the Oriental Region from Nepaland Sri Lankathrough India, whole continental SE Asia and some of the islands of Sundaland (Sumatra, Sumba). Records are surprisingly missing for whole lowland northern part of India(and  P. nigrovittatumseems therefore be absent from semidesert areas) and from many islands of Sundaland (in this case most probably because of the collecting bias). 3499776350 ZMUM 33 34 1 1 lectotype 3499776404 2010-07-13 SYSU China Nanjing County 24.866667 Hexi 1249 117.23333 33 34 1 3499776478 1980-09-27 BMNH P. M. Hammond 23.15 Guangzhou 1257 113.28333 Guangdong 33 34 1 3499776325 2004-05-28 SYSU F. L. Jia 23.353611 Lubao 21 112.89611 Sanshui 33 34 17 3499776384 2006-04-23 2006-04-26 2006-04-23 SYSU F. L. Jia Yangmei Gaoming 33 34 14 3499776373 [422,1177,1647,1671] 1985-02-29 SYSU Lianjiang County Wu Wu Wuling 33 34 1 3499776424 1986-05-09 SYSU Xinyi County Wu Wu Wu Wu 33 34 2 3499776450 [508,1241,1685,1709] 1986-06-07 1986-06-13 1986-06-07 SYSU Wu Wu 33 34 13 3499776368 1985-08-25 SYSU Guangzhou 33 34 9 3499776441 1985-10-10 SYSU Guangzhou 33 34 4 3499776385 1986-08 1988-01 1986-08 BMNH, SYSU 24.170555 Wu Wu 21 107.248886 33 34 10 3499776443 1996-01-20 NHMW M. A. Jach 250 19.1444 Hainan 1 108.9469 33 34 1 1 3499776364 BMNH, NHMW J. J. Walker Hong Kong 22.4 Hong Kong 1260 114.13333 33 34 4 3499776313 1996-01-14 NHMW M. A. Jach Cook Islands 22.524445 Shue Ha 21 114.219444 33 34 1 3499776369 NHMW G. de Rougemont Hong Kong 22.283333 Hong Kong 1260 114.13333 33 34 3 3499776431 BMNH J. J. Walker 30.033333 Chusan Is. 1224 122.11667 Zhejiang 33 34 1 3499776301 2008-05-07 NMPC India 280 34 35 27.0325 Bhalukpong 20 92.60667 Arunachal Pradesh 33 34 1 1 3499776370 [414,1291,189,214] BMNH 27.833334 Sudiya 1235 95.75 Assam 34 35 2 3499776352 2002-08-04 2002-08-12 2002-08-04 NMPC India 15 15.013056 Madgaon 21 74.03278 Margao 34 35 1 3499776432 BMNH 15.85 Belgaum 925 74.55 Karnataka 34 35 1 3499776470 1999-01-02 NHMW D. Boukal 600 8.766666 Trivandrum 1300 77.11667 Kerala 34 35 1 3499776315 [247,1191,338,362] BMNH Malabar 34 35 1 3499776359 1947-02 FMNH India 10.916667 Kurumbagatam 1296 79.816666 34 35 1 3499776330 FMNH F. Psota 10.766666 Negapatam 1297 79.833336 Tamil Nadu 34 35 3 3499776401 BMNH H. L. Andrewes Nilgiri Hills 34 35 40 3499776475 IRSNB 10.233334 Shenbaganur 1298 77.5 Shempagonur 34 35 2 3499776366 1923-11-23 BMNH 29.8 Akeswar 1225 78.61667 Uttarakhand 34 35 1 3499776416 IRSNB 30.333334 Dobhalwala 925 78.04361 Dehradun 34 35 2 3499776390 BMNH H. G. Champion 29.6 Almora 1226 79.566666 34 35 6 3499776338 2001-01-30 2001-02-02 2001-01-30 NMPC P. Votruba Indonesia -9.851389 Sumba 21 120.30889 East Nusa Tenggara 34 35 1 3499776434 1990-02-17 NHMW S. Schodl 1000 2.662778 Prapat 21 98.93528 North Sumatra 34 35 1 3499776354 1922-10 BMNH E. Jacobson 920 -0.30555558 Fort de Kock 21 100.36917 West Sumatra 34 35 2 3499776411 IRSNB E. Jacobson 34 35 5 3499776462 1997-04-27 1997-04-30 1997-04-27 NHMB V. Kuban Laos 150 18.266666 Bolikhamxai 1275 103.183334 Bolikhamxai 34 35 1 3499776314 2001-05-01 2001-05-18 2001-05-01 NHMB, NMPC V. Kuban 600 18.35 1275 105.13333 34 35 28 3499776429 1998-04-18 1998-05-01 1998-04-18 FHGC Iceland 18.301943 Sausa 21 105.07333 34 35 1 3499776311 2009-06-01 2009-06-16 2009-06-01 NMPC V. Kuban 1495 20.219168 Phou Pane Mt. 21 103.99833 Houaphan 34 35 2 3499776460 2002-05-11 2002-05-21 2002-05-11 NHMB V. Kuban 750 19.583334 Thong Khan 1271 101.96667 Louangphabang 34 35 1 3499776406 2009-05-10 2009-05-30 2009-05-10 NMPC V. Kuban 1420 19.636667 Phou Sane Mt. 127 103.33667 Xiangkhouang 34 35 2 3499776383 2009-11-07 2009-11-22 2009-11-07 ASKC U. Schmidt Malaysia 6.366667 Langkawi 929 99.675 Kedah 34 35 1 3499776445 BMNH Malaysia 2.1980555 Wallace 21 102.24972 34 35 2 3499776348 1984-03-01 1984-03-13 1984-03-01 BMNH L. Jessop 4.6833334 Taman Negara 1306 102.566666 Pahang 34 35 1 3499776312 BMNH Perak Perak 34 35 1 3499776316 1995-07-29 FHGC Nepal 2000 28.316668 Sikles Mts. 1232 84.0 Gandaki 34 35 1 3499776480 BMNH, IRSNB Singapore 1.35 SINGAPORE 1308 103.833336 34 35 8 3499776469 BMNH Sri Lanka SRI LANKA 34 35 2 3499776339 [698,1363,1234,1258] IRSNB F. Chapuis 34 35 1 3499776335 BMNH C. Lewis 6.991667 Kitulgalle 21 80.44833 Sabaragamuwa 34 35 1 3499776457 1962-03-02 FHGC 1097 6.7905555 Haldummulla 21 80.86472 Uva 34 35 1 3499776464 1915-11-14 1915-11-15 1915-11-14 BMNH Cameroon 6.9166665 Cameron 925 79.84167 34 35 1 3499776345 1962-01-11 FHGC 6.75 Brinck 1304 79.9 Colombo 34 35 1 3499776451 1962-01-15 1962-01-21 1962-01-15 FHGC 7.090278 Brinck 21 80.03389 Colombo 34 35 1 3499776317 1989-03 NHMW Thailand 15.566667 Lan Sak 1284 99.51667 Uthai Thani 34 35 1 3499776357 NMPC P. Marhoul Vietnam 21.633333 Tam 1263 105.5 Bac 34 35 1 3499776403 IRSNB Vietnam Foret de Kep 34 35 1 3499776437 1995-03-28 1995-04-12 1995-03-28 NHMW 685 14.166667 An Khe 1289 108.5 Gia Lai 34 35 20 3499776402 IRSNB H. Perrot Chobo Hoa Binh 34 35 1 3499776340 1918-08 BMNH R. V. de Salvaza 20.833334 Hoa Binh 1266 105.333336 34 35 37 3499776310 [385,1093,1682,1707] IRSNB A. de Cooman 34 35 1 1 3499776341 1994-04-21 1994-04-27 1994-04-21 NHMW 11.933333 Da Lat 1294 108.45 Lam Dong 34 35 1 3499776474 BMNH 21.066668 Lactho 1265 106.1 Nghe An 34 35 3 3499776323 IRSNB 34 35 66