Mesabolivar pseudoblechroscelis
González-Sponga 1998: 27
Mesabolivar eberhardi
Venezuela
Venezuela (Monagas)
Blechroscelis Simon, 1893
M. eberhardi
Blechroscelis araguanus
On Venezuelan pholcid spiders (Araneae, Pholcidae)
Huber, Bernhard A.
Villarreal, Osvaldo
European Journal of Taxonomy
2020
2020-10-01
718
1
317
3ZXDJ
Huber, 2000
Huber
2000
[580,1008,1689,1716]
Arachnida
Pholcidae
Mesabolivar
Animalia
Araneae
163
164
Arthropoda
species
eberhardi
Figs 597–598, 1021-1022, 1025, 1048 Synonymy, see Huber 2018: 34.
Mesabolivar pseudoblechroscelis(misidentification) – González-Sponga 1998: 27, figs 40–41 ( ♀only). Notes Mesabolivar eberhardiis a common inhabitant of native and disturbed habitats in much of northern South America and one the most ubiquitous pholcids in Venezuela. Variation in the details of genitalic shape has been noted long ago ( Huber 2000), but this referred to specimens from Peru, Brazil, and Colombia, far from the typelocality in Venezuela (Monagas). All Venezuelan specimens seen in this and previous studies ( Huber 2000, 2018) are unproblematic representatives of this species. This is also true of the six species described by González-Sponga (2011a)under BlechroscelisSimon, 1893. They have been synonymized with M. eberhardibefore ( Huber et al.2014a), and the reexamination of all typesin Nov. 2018confirmed the synonymies. However, a few errors in González-Sponga’s (2011a)work remain to be corrected: For “ Blechroscelis araguanus”, only 1 ♂ holotypeis mentioned in the original description, but the female is also described and illustrated. We assume that this is based on MAGS 1393 (MIZA 105765), 2 ♀♀, 3 juvs from the same locality (La Montañita; corrected coordinates: 10.213° N, 67.176° W).
The coordinates of the typelocality of “ Blechroscelis blechroscelis”, Cerro Guaiquinima, are approximately 5.758° N, 63.567° W(instead of 10.758°N). The coordinates of the typelocality of “ Blechroscelis copeyensis”, Cerro CopeyNational Park, are probably 11.047° N, 63.891° W(rather than 11.033°N). Most of the measurements are probably correct, but some are obviously wrong, casting doubt on the precision in general. For example, metatarsus 4 is given as shorter than metatarsus 3 inthe holotypeof “ B. blechroscelis”; femur 3 is given as longer than femora 2 and 4 inthe female of “ B. andinensis”; and carapace shape is given as highly variable among ‘species’ (width =1.0–1.7 × length), while in fact is it highly consistent but heavily dependent on the angle of view.
Material examined VENEZUELA– Amazonas• 1 ♂, 1 ♀, MIZA 105616( MAGS 1100), San Juan de Manapiare[ 5.324° N, 66.055° W], Jul. 1987(A. González). – Anzoátegui• 3 ♂♂, 1 ♀, 4 juvs, MIZA 105776( MAGS 1036), near Sabana de Uchire[approximately 10.02° N, 65.52° W], 2 Mar. 1987(A.R. Delgado, M.A. González-S.). – Aragua• 1 ♀, ZFMK(Ar 21966), Puerto Colombia( 10.5025° N, 67.6057° W), 25 ma.s.l., 2 Dec. 2018(B.A. Huber, O. VillarrealM.)• 1 ♂, ZFMK(Ar 21967), between Maracay and Puerto Colombia( 10.4304° N, 67.5998° W), 380 ma.s.l., 2 Dec. 2018(B.A. Huber, O. VillarrealM.)• 2 ♀♀, 3 juvs, MIZA 105765( MAGS 1393), La Montañita, carretera Tejerias-Tiara[ 10.213° N, 67.176° W], 28 Feb. 1994(A.R. Delgado, M.A. GonzálezS.)• 1 ♀, 1 juv., MIZA 105642( MAGS 161), Colonia Tovar[approximately 10.405° N, 67.303° W], 2 May 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 2 ♀♀, 1 juv., CAS(9027227), Henri Pittier National Park, Rancho Grande[ 10.349° N, 67.684° W], cloud forest, 29 Dec. 1970(W.B. Peck). – Bolívar• 2 ♂♂, 2 ♀♀, ZFMK(Ar 21968), and 1 ♂, 1 ♀, 1 juv.in pure ethanol, ZFMK(Ven18-162), Ciudad Guayana, Parque La Llovizna( 8.3130° N, 62.6724° W), 50 ma.s.l., 11 Nov. 2018(B.A. Huber, O. VillarrealM.)• 2 ♂♂, 4 ♀♀, ZFMK(Ar 21969), and 1 ♂in pure ethanol, ZFMK(Ven18-174), same locality but 8.3112° N, 62.6742° W, 30 ma.s.l., 14 Nov. 2018• 1 ♂, MIZA 105628( MAGS 1369), Los Pijiguaos[approximately 6.57° N, 66.81° W], May 1992(J.M. AyalaL.)• 1 ♂, MIZA 105696( MAGS 1322), Guri[ 7.77° N, 63.04° W], 9 Oct. 1991(A.R. Delgado, M.A. González-S.)• 2 ♂♂, MIZA 105717( MAGS 1191), Camp 4, Cerro Guaiquinima(Las cuevas), 980 ma.s.l., 18 Jan. 1990( C. Toro). – Capital• 1 ♀, 1 juv., MIZA 105592( MAGS 1028), near Hotel Humboldt[ 10.542° N, 66.875° W], 17 Jan. 1987(C.E. Avila). 6 ♂♂, 2 ♀♀, 7 juvs, MIZA 105672( MAGS 1167), El Ávila National Park, Quebrada Quintero[ 10.518° N, 66.851° W], 19 Aug. 1989(A.R. Delgado, E. González, M.A. González-S.)• 1 ♂, 1 ♀, 1 juv., MIZA 105728( MAGS 1171), El Ávila National Park, Los Venados[ 10.536° N, 66.898° W], 9 Sep. 1989(E. González-S., M.A.González-S.)• 6 ♂♂, 5 ♀♀, 3 juvs(with one parasitized egg-sac), MIZA 105699( MAGS 1374), Las Canoas[ 10.547° N, 66.932° W], 1200 ma.s.l., 12 Oct. 1992(E. GonzálezL., E. GonzálezS., M.A.González-S.). – Carabobo• 1 ♀, MIZA 105605( MAGS 336), San Esteban[ 10.425° N, 68.013° W], 4 Aug. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 4 ♂♂, 1 juv., MIZA 105749( MAGS 333), same data. – Falcón• 2 ♂♂, 2 ♀♀, ZFMK(Ar 21970), and 1 ♂in pure ethanol, ZFMK(Ven18-177), Península de Paraguaná, Cueva del Guano ( 11.9000° N, 69.9479° W), 140 ma.s.l., 16 Nov. 2018(B.A. Huber, O. VillarrealM.)• 2 ♂♂, 2 ♀♀, ZFMK(Ar 21971), and 1 ♀in pure ethanol, ZFMK(Ven18-180), Península de Paraguaná, near Cueva del Guano( 11.9026° N, 69.9456° W), 140 ma.s.l., 16 Nov. 2018(B.A. Huber, O. VillarrealM.)• 6 ♂♂, 4 ♀♀, 1 juv., ZFMK(Ar 21972), and 1 ♀in pure ethanol, ZFMK(Ven18-185), Península de Paraguaná, Cerro Santa Ana( 11.8177° N, 69.9468° W), 480 ma.s.l., 17 Nov. 2018(B.A. Huber, O. VillarrealM.)• 1 ♂, 1 ♀, ZFMK(Ar 21973), Medanos de Coro( 11.5690° N, 69.7147° W), 3 ma.s.l., 17 Nov. 2018(B.A. Huber, O. VillarrealM.)• 2 ♂♂, 1 ♀, ZFMK(Ar 21974), and 1 ♀in pure ethanol, ZFMK(Ven18-191), Sierra de San Luis, E Curimagua( 11.1748° N, 69.6273° W), 960 ma.s.l., 18 Nov. 2018(B.A. Huber, O. VillarrealM.)• 2 ♀♀, MIZA 105621( MAGS 1132), Paraguaná, Cerro Capuchino, Montecano[approximately 11.94° N, 69.97° W], 27 Jan. 1988(R. Ramirez)• 1 ♀, MIZA 105795( MAGS 738), San Juan de los Cayos[ 11.173° N, 68.408° W], 3 Oct. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 1 ♀, MIZA 105796( MAGS 711), Riecito[ 10.906° N, 68.767° W], 30 Sep. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 2 ♂♂, 2 ♀♀, ZFMK(Ar 21975), and 1 ♂, 1 ♀, 1 juv.in pure ethanol, ZFMK(Ven20-151), forest near Santa Cruz de La Alegría( 10.8795° N, 68.4949° W), 100 ma.s.l., 15 Feb. 2020(B.A. Huber, O. Villarreal M., Q. AriasC.). – Guárico• 1 ♂, 1 ♀, MIZA 105695( MAGS 988), El Morrito, road Altagracia de Orituco to San Francisco de Macaira[ 9.916° N, 66.283° W], 10 Apr. 1982(R. Torrealba)• 1 ♀, 1 juv., MIZA 1808, Agua Salada[approximately 9.163° N, 65.775° W], 540 ma.s.l., “sistema cuevas y cañones”, 22 Nov. 1997(O. Villarreal). – Lara• 4 ♂♂, 5 ♀♀, ZFMK(Ar 21976), and 1 ♂, 1 ♀in pure ethanol, ZFMK(Ven18-198), between Coro and Barquisimeto, El Rodeo( 10.7240° N, 69.3008° W), 400 ma.s.l., 19 Nov. 2018(B.A. Huber, O. VillarrealM.)• 2 ♂♂, 1 ♀, ZFMK(Ar 21977), and 1 ♀in pure ethanol, ZFMK(Ven18-201), between Barquisimeto and Boconó( 9.5906° N, 69.8343° W), 1370 ma.s.l., 20 Nov. 2018(B.A. Huber, O. VillarrealM.). – Mérida• 2 ♀♀, (paratypes of M. pseudoblechroscelis), MIZA 105737( MAGS 986part), near Mesa Bolívar[ 8.474° N, 71.598° W], 25 Dec. 1981(A.R. Delgado, J.A. GonzálezD., M.A.González-S.)• 1 ♂, MIZA 105611( MAGS 1053), Mesa Bolívar[ 8.474° N, 71.598° W], 21 Dec. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 6 ♂♂, 3 ♀♀, 2 juvs, MIZA 105744( MAGS 835), Represa General J.A. Páez[ 8.900° N, 70.617° W], 5 Sep. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 2 ♀♀, 1 juv., MIZA 105683( MAGS 838), Dtto Rivas Dávila, between La Playa and San Pablo[ 8.28° N, 71.80° W, ~ 1350 ma.s.l.], 4 Sep. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 1 ♂, 1 ♀, 1 juv., ZFMK(Ar 21978), Las Piedras, ‘site 1’ ( 8.8939° N, 70.6448° W), 1710 ma.s.l., forest remnant, 7 Feb. 2020(B.A. Huber, O. VillarrealM., Q. AriasC.)• 7 ♂♂, 3 ♀♀, ZFMK(Ar 21979–80), and 1 ♂, 2 ♀♀in pure ethanol, ZFMK(Ven20-108), Las Piedras, ‘site 2’ ( 8.9002° N, 70.6279° W), 1700 ma.s.l., 7 Feb. 2020(B.A. Huber, O. VillarrealM., Q. AriasC.)• 6 ♂♂, 1 ♀, ZFMK(Ar 21981), forest above Caño Azul( 8.8543° N, 71.3651° W), 280 ma.s.l., 13 Feb. 2020(B.A. Huber, O. VillarrealM., Q. AriasC.). – Miranda• 1 ♂, 1 ♀in pure ethanol, ZFMK(Ven18-149), El Ávila National Park, between Sabas Nieves and La Silla( 10.5245° N, 66.8566° W), 1600 ma.s.l., 7 Nov. 2018(B.A. Huber, O. VillarrealM.)• approximately 4 ♂♂, 10 ♀♀, 10 juvs, MIZA 105595( MAGS 1022), Boca de Cura[ 10.20° N, 66.30° W], 17 Jan. 1987(A.R. Delgado, M.A. González-S.)• approximately 10 ♂♂, 2 ♀♀, 2 juvs, MIZA 105598( MAGS 1021), same data• 1 ♀, MIZA 105619(separated from MAGS 1111), road Carenero-Chirimena[approximately 10.57° N, 66.15° W], 9 Feb. 1988(A.R. Delgado, M.A. González-S.)• 1 ♀, MIZA 105623( MAGS 1121), Hacienda Santa Rosalia, alrededores de Arag̹ita[approximately 11.215° N, 66.458° W], 16 Jul. 1988(A.R. Delgado, M.A. González-S.)• 3 ♂♂, 3 juvs, MIZA 105633( MAGS 1087), Zamora, Salmerón[approximately 10.472° N, 66.369° W], 15 Aug. 1987(A.R. Delgado, M.A. González-S.)• 1 ♂, MIZA 105686( MAGS 966), 10 kmN de Guatire[approximately 10.55° N, 66.58° W], Hacienda Santa Rosa, 19 Feb. 1982(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 1 ♂, 3 ♀♀, MIZA 105697( MAGS 845), near Tacarigua de Mamporal(“ a 1 kmde Tacarigua de Mamporal en la via a Rio Chico”) [ 10.382° N, 66.147° W], 31 Oct. 1981(A.R. Delgadode G., M.A. GonzálezS.)• 3 ♂♂, MIZA 105747( MAGS 1441), San Antonio de los Altos[ 10.369° N, 66.973° W], “Urb. Las Minas, Ed. Altair”, 1998 ( M. García)• 2 ♂♂, 6 ♀♀, 4 juvs, MIZA 105704( MAGS 242), El Castaño, carretera a Arag̹ita [ 10.283° N, 66.400° W], 27 Jun. 1981(A.R. Delgadode G., J.A. GonzálezD., M.A. GonzálezS.)• 1 ♀, MIZA 105721( MAGS 1093), Guatopo National Park, Macanilla, 5 Sep. 1987(A.R. Delgado, M.A.González-S.)• 1 ♂, 1 ♀, MIZA 105690( MAGS 506), El Ávila National Park(“estribaciones del Avila”), El Paraiso[ 10.530° N, 66.819° W], ~ 1500 ma.s.l., 16 Aug. 1981(J.A. GonzálezD.)• 1 ♂, 1 juv., MIZA 105617( MAGS 1109), La Julia, via Pico Naiguatá[between 10.50° N, 66.81° Wand 10.54° N, 66.78° W], 15 Oct. 1987(H. Bird)• 1 ♀, ZFMK(Ar 21982), El Ávila National Park, near La Julia,‘site 3’ ( 10.5066° N, 66.8119° W), 1090 ma.s.l., forest near dry brook bed, 22 Feb. 2020(B.A. Huber, O. VillarrealM.)• 2 ♂♂, ZFMK(Ar 21983), El Ávila National Park, near La Julia, trail to Rancho Grande( 10.5164° N, 66.8089° W), 1460 ma.s.l., degraded forest along small stream, 22 Feb. 2020(B.A. Huber, O. VillarrealM.)• 1 ♂, 1 ♀, MIZA 105670( MAGS 1169), Rio Aricagua[ 10.583° N, 66.234° W], 27 Aug. 1989( M.A.González-S.)• 1 ♀, MIZA 105800( MAGS 985), Dtto. Brión, Hacienda La Maravilla, 6 Jun. 1981( A. Utrera)• 7 ♂♂, 3 ♀♀, 3 juvs, MIZA 105594( MAGS 1030), Estanque Ayala[ 10.505° N, 66.763° W], 25 Jan. 1987(A.R. Delgadode G., E.S.S., E.G.L., M.A. GonzálezS.)• 1 ♂, 2 ♀♀, 2 juvs, MIZA 105668( MAGS 1165), old road Caracas-Charallave, Los Amarillos[ 10.367° N, 66.945° W], Jul. 1989(A.R. Delgado, M.A. González-S.)• 2 ♂♂, 1 ♀, CAS(9027359), Caracas, Hacienda La Trinidad[approximately 10.42° N, 66.85° W, 1100 ma.s.l.], 28 Dec. 1970(W.B. Peck)• 1 ♀, 1 juv., MIZA, El Volcán, Topotepuy[ 10.417° N, 66.851° W, ~ 1450 ma.s.l.], 11–13 Nov. 2019(O. Villarreal, J. Rodriguez). – Monagas• 1 ♂, 8 juvs, MIZA 105710( MAGS 1318), Cueva del Guácharo[ 10.174° N, 63.552° W], 12 Oct. 1991(A.R. Delgado, M.A. González-S.)• 1 ♂, 2 ♀♀, 3 juvs, MIZA 105752( MAGS 1313), same locality, 14 Oct. 1991( M. de García, A.R. Delgado, M.A. GonzálezS.). – Nueva Esparta• 1 ♀, MIZA 105780( MAGS 1007), Cerro Copei[ 11.047° N, 63.891° W], Dec. 1986(C.E. ContrerasA.). – Portuguesa• 1 ♂, 1 ♀, MIZA 105574( MAGS 1400), Paez, Acarigua(Pozo Blanco) [approximately 9.52° N, 69.15° W], 7 Jul. 1995(A.R. Delgado, M.A. González-S.). – Táchira• 5 ♂♂, 2 ♀♀, 1 juv., ZFMK(Ar 21984), and 1 ♀in pure ethanol, ZFMK(Ven20-123), SE Pregonero, forest near La Trampa( 7.9236° N, 71.7152° W), 1300 ma.s.l., 10 Feb. 2020(B.A. Huber, O. Villarreal M., Q. Arias C.). – Trujillo• 7 ♂♂, 3 ♀♀, ZFMK(Ar 21985–86), and 2 ♀♀in pure ethanol, ZFMK(Ven18-216), between Boconó and Burbusay( 9.3945° N, 70.2674° W), 1820 ma.s.l., 22 Nov. 2018(B.A. Huber, O. VillarrealM.)• 1 ♀, 1 juv., MIZA 105762( MAGS 1386), Boconó, Miticún[ 9.235° N, 70.264° W], 26 Dec. 1993(E. GonzálezS., M.A. GonzálezS.)• 1 ♂, 1 ♀, MIZA 105778( MAGS 1011), near Boconó, Hacienda La Encomienda[ 9.284° N, 70.317° W], 27 Dec. 1986(R. Manzanilla)• 1 ♂(palps missing), MIZA 1809, route to Buena Vista via La Gira, 650 ma.s.l., 14 May 2005( O. VillarrealM., H. Escalona)• 1 ♂, ZFMK(Ar 21987), and 1 juv.in pure ethanol, ZFMK(Ven20-140), near El Encanto( 9.7562° N, 70.7418° W), 150 ma.s.l., among rocks, 13 Feb. 2020(B.A. Huber, O. Villarreal M., Q. AriasC.). – La Guaira• 3 ♂♂, 1 ♀, ZFMK(Ar 21988), and 1 ♂in pure ethanol, ZFMK(Ven18-159), El Limón, above road Colonia Tovar-Puerto Cruz( 10.4566° N, 67.2548° W), 1535 ma.s.l., 9 Nov. 2018(B.A. Huber, O. VillarrealM.)• 4 ♂♂, 2 ♀♀, 3 juvs, MIZA 105742( MAGS 993), Hacienda El Limón[approximately 10.475° N, 67.283° W], 26 Nov. 1986(A.R. Delgado, M.A. González-S.)• 3 ♂♂, 2 ♀♀, 1 juv., MIZA 105777( MAGS 983), same locality, 12 Feb. 1982(M.A. GonzálezS.)• 2 ♀♀, ZFMK(Ar 21989), El Limón, ‘site 1’ ( 10.4788° N, 67.3010° W), 600 ma.s.l., forest remnant along small stream, 21 Feb. 2020(B.A. Huber, O. VillarrealM.)• 2 ♂♂, 2 ♀♀, 1 juv., ZFMK(Ar 21990), and 1 ♀in pure ethanol, ZFMK(Ven20-171), El Limón, ‘site 2’ ( 10.4774° N, 67.2819° W), 1235 ma.s.l., forest along stream, 21 Feb. 2020(B.A. Huber, O. VillarrealM.)• 1 ♀, 1 juv., MIZA 105583( MAGS 1412), Galipán[approximately 10.55° N, 66.89° W], 26 Jun. 1996(J.A. GonzálezD.). – Yaracuy• 1 ♂, MIZA 105641( MAGS 1085), Yurubi National Park[approximately 10.48° N, 68.66° W], 11 Sep. 1986(E. Yustiz)• 1 ♀, MIZA 1810, Minas de Aroa[approximately 10.422° N, 68.878° W], “tunel de la casa de montaña”, 22 Dec. 1998(O. Villarreal)• 6 ♂♂, 3 ♀♀, ZFMK(Ar 21991), and 2 ♀♀, 1 juv.in pure ethanol, ZFMK(Ven20-157), Yurubi National Park( 10.4913° N, 68.6564° W), 140 ma.s.l., forest along stream, 16 Feb. 2020(B.A. Huber, O. VillarrealM., Q. AriasC.)• 2 ♂♂, ZFMK(Ar 21992), Guaquira, ‘site 1’ ( 10.2951° N, 68.6535° W), 120 ma.s.l., forest along stream, 16 Feb. 2020(B.A. Huber, O. VillarrealM., Q. AriasC.)• 3 ♂♂, 4 ♀♀, ZFMK(Ar 21993), Guaquira, ‘site 2’ ( 10.2807° N, 68.6530° W), 150 ma.s.l., forest along stream, 17 Feb. 2020(B.A. Huber, O. VillarrealM., Q. AriasC.).
Distribution This species is widespread in northern South America (cf. Huber 2018: fig. 724). The map in Fig. 1048 shows only the known Venezuelan records and those in neighboring regions.
1987-07
MIZA, MAGS
Gonzalez
Venezuela
5.324
San Juan de Manapiare
78
-66.055
165
166
MIZA 105616, MAGS 1100
2
1
1
Amazonas
1987-03-02
R
Delgado
10.02
Aragua
779
-65.52
165
166
MIZA 105776, MAGS 1036
8
1
3
Aragua
2018-12-02
ZFMK
Huber & Villarreal
25
10.5025
Puerto Colombia
7
-67.6057
Ar
165
166
1
1
Aragua
2018-12-02
ZFMK
Huber & Villarreal
380
10.4304
Ar
7
-67.5998
165
166
1
1
Aragua
1994-02-28
R
Delgado & Gonzalez
10.213
La Montanita
77
-67.176
165
166
MIZA 105765, MAGS 1393
5
2
Aragua
1981-05-02
R
Delgado & Gonzalez & M. A. Gonzalez
10.405
Colonia Tovar
77
-67.303
165
166
MIZA 105642, MAGS 161
2
1
Aragua
cloud forest
1970-12-29
CAS
Peck
10.349
Rancho Grande
77
-67.684
Henri Pittier National Park
165
166
3
2
Bolivar
2018-11-11
ZFMK
Huber & Villarreal
Ar
50
8.313
Ciudad Guayana
7
-62.6724
Ven
165
166
7
3
3
Bolivar
2018-11-14
ZFMK
Huber & Villarreal
Ar
30
8.3112
Ven
7
-62.6742
Ar
165
166
7
4
3
Bolivar
1992-05
MIZA, MAGS
Ayala
6.57
Los Pijiguaos
782
-66.81
165
166
MIZA 105628, MAGS 1369
1
1
Bolivar
1991-10-09
R
Delgado
7.77
Guri
781
-63.04
165
166
MIZA 105696, MAGS 1322
1
1
Bolivar
1990-01-18
MIZA, MAGS
C. Toro & Capital
Camp
980
Cerro Guaiquinima
Cerro Guaiquinima
165
166
MIZA 105717, MAGS 1191
2
2
Bolivar
1987-01-17
1989-08-19
1987-01-17
R
Avila & Delgado & Gonzalez
Hotel Humboldt
10.518
Quebrada Quintero
77
-66.851
El Avila National Park
165
166
MIZA 105592, MAGS 1028, MIZA 105672, MAGS 1167
17
3
6
Bolivar
1989-09-09
MIZA, MAGS
M. A.
10.536
Los Venados
77
-66.898
El Avila National Park
165
166
MIZA 105728, MAGS 1171
3
1
1
Bolivar
1992-10-12
MIZA, MAGS
Gonzalez & M. A.
1200
10.547
Las Canoas
77
-66.932
165
166
MIZA 105699, MAGS 1374
14
5
6
Carabobo
1981-08-04
R
Delgado & Gonzalez & M. A. Gonzalez
10.425
San Esteban
77
-68.013
165
166
MIZA 105605, MAGS 336
1
1
Carabobo
1981-08-04
MIZA, MAGS
Delgado & Gonzalez & M. A. Gonzalez
10.425
Falcon
77
-68.013
165
166
MIZA 105749, MAGS 333
5
4
Falcon
2018-11-16
ZFMK
Huber & O. Villarreal
Ar
140
11.9
Peninsula de Paraguana
7
-69.9479
Ven
165
166
5
2
3
Falcon
2018-11-16
ZFMK
Huber & Villarreal
Ar
140
11.9026
Peninsula de Paraguana
7
-69.9456
Ven
165
166
5
3
2
Falcon
2018-11-17
ZFMK
Huber & Villarreal
Ar
480
11.8177
Peninsula de Paraguana
7
-69.9468
Ven
165
166
12
5
6
Falcon
2018-11-17
ZFMK
Huber & Villarreal
3
11.569
Medanos de Coro
7
-69.7147
Ar
165
166
2
1
1
Falcon
2018-11-18
ZFMK
Huber & Villarreal
Ar
960
11.1748
Sierra de San Luis
7
-69.6273
Ven
165
166
4
2
2
Falcon
1988-01-27
R
Ramirez
Paraguana
11.94
Montecano
776
-69.97
Cerro Capuchino
165
166
MIZA 105621, MAGS 1132
2
2
Falcon
1981-10-03
R
Delgado & Gonzalez & M. A. Gonzalez
166
167
11.173
San Juan de los Cayos
77
-68.408
San Juan
165
166
MIZA 105795, MAGS 738
1
1
Falcon
1981-09-30
R
Delgado & Gonzalez & M. A. Gonzalez
10.906
Riecito
77
-68.767
166
167
MIZA 105796, MAGS 711
1
1
Falcon
2020-02-15
ZFMK
Huber & Villarreal M. & Q. Arias
Ar
100
10.8795
Guarico
7
-68.4949
Ven
166
167
7
3
3
Guarico
1982-04-10
R
Torrealba
9.916
El Morrito
77
-66.283
Altagracia de Orituco
166
167
MIZA 105695, MAGS 988
2
1
1
Guarico
1997-11-22
MIZA
Villarreal
540
9.163
Agua Salada
77
-65.775
166
167
MIZA 1808
2
1
Lara
2018-11-19
ZFMK
Huber & Villarreal
400
10.724
Ven
7
-69.3008
Ar
166
167
11
6
5
Lara
2018-11-20
ZFMK
Huber & Villarreal
1370
9.5906
Merida
7
-69.8343
Ar
166
167
4
2
2
Merida
1981-12-25
R
Delgado & Gonzalez & M. A.
8.474
Mesa Bolivar
77
-71.598
166
167
MIZA 105737, MAGS 986
2
2
Merida
paratype
1981-12-21
R
Delgado & Gonzalez & M. A. Gonzalez
8.474
Mesa Bolivar
77
-71.598
166
167
MIZA 105611, MAGS 1053
1
1
Merida
paratype
1981-09-05
R
Delgado & Gonzalez & M. A. Gonzalez
8.9
Represa General J. A. Paez
77
-70.617
Represa General
166
167
MIZA 105744, MAGS 835
11
3
6
Merida
paratype
1981-09-04
R
Delgado & Gonzalez
1350
8.28
Dtto Rivas Davila
781
-71.8
166
167
MIZA 105683, MAGS 838
3
2
Merida
paratype
forest remnant
2020-02-07
ZFMK
Huber & Villarreal & Q. Arias
1710
8.8939
Las Piedras
7
-70.6448
Ar
166
167
3
1
1
Merida
paratype
2020-02-07
ZFMK
Huber & O. Villarreal & Q. Arias
Ar
1700
8.9002
Las Piedras
7
-70.6279
Ven
166
167
13
5
8
Merida
paratype
2020-02-13
ZFMK
Huber & Villarreal & Q. Arias
280
8.8543
Miranda
7
-71.3651
Ar
166
167
7
1
6
Miranda
paratype
2018-11-07
ZFMK
Huber & Villarreal
1600
10.5245
El Avila National Park
7
-66.8566
Ven
166
167
2
1
1
Miranda
paratype
1987-01-17
R
Delgado
10.2
Boca de Cura
779
-66.3
166
167
MIZA 105595, MAGS 1022
24
10
4
Miranda
paratype
1987-01-17
MIZA, MAGS
Delgado
10.2
Boca de Cura
779
-66.3
166
167
MIZA 105598, MAGS 1021
14
2
10
Miranda
paratype
1988-02-09
R
Delgado
10.57
Carenero-Chirimena
779
-66.15
166
167
MIZA 105619, MAGS 1111
1
1
Miranda
paratype
1988-07-16
R
Delgado
11.215
Hacienda Santa Rosalia
77
-66.458
Hacienda Santa Rosalia
166
167
MIZA 105623, MAGS 1121
1
1
Miranda
paratype
1987-08-15
R
Delgado
10.472
Salmeron
77
-66.369
Zamora
166
167
MIZA 105633, MAGS 1087
6
3
Miranda
paratype
1982-02-19
R
Hacienda Santa Rosa & Delgado & G., J. A. Gonzalez & M. A. Gonzalez
10.55
10 km N de Guatire
779
-66.58
166
167
MIZA 105686, MAGS 966
1
1
Miranda
paratype
1981-10-31
R
Delgado & Gonzalez
Tacarigua de Mamporal
10.382
Rio Chico
77
-66.147
Tacarigua de Mamporal
166
167
MIZA 105697, MAGS 845
4
3
1
Miranda
paratype
MIZA, MAGS
Ed. Altair & M. Garcia
10.369
San Antonio de los Altos
77
-66.973
San Antonio
166
167
MIZA 105747, MAGS 1441
3
3
Miranda
paratype
1981-06-27
R
Delgado & Gonzalez & M. A. Gonzalez
10.283
El Castano
77
-66.4
166
167
MIZA 105704, MAGS 242
12
6
2
Miranda
paratype
1987-09-05
R
Delgado, M. A.
Guatopo National Park
Guatopo National Park
166
167
MIZA 105721, MAGS 1093
1
1
Miranda
paratype
1981-08-16
MIZA, MAGS
Gonzalez
El Avila National Park
1500
10.53
El Paraiso
77
-66.819
Avila
166
167
MIZA 105690, MAGS 506
2
1
1
Miranda
paratype
1987-10-15
MIZA, MAGS
Bird
10.54
La Julia
779
-66.78
La Julia
166
167
MIZA 105617, MAGS 1109
2
1
Miranda
paratype
forest near dry brook bed
2020-02-22
ZFMK
Huber & Villarreal
Ar
1090
10.5066
El Avila National Park
7
-66.8119
El Avila National Park
166
167
1
1
Miranda
paratype
degraded forest along small stream
2020-02-22
ZFMK
Huber & O. Villarreal
El Avila National Park
Ar
1460
10.5164
El Avila National Park
7
-66.8089
El Avila National Park
166
167
2
2
Miranda
paratype
1989-08-27
MIZA, MAGS
M. A.
10.583
Rio Aricagua
77
-66.234
166
167
MIZA 105670, MAGS 1169
2
1
1
Miranda
paratype
1981-06-06
MIZA, MAGS
A. Utrera
Hacienda La Maravilla
Dtto. Brion
166
167
MIZA 105800, MAGS 985
1
1
Miranda
paratype
1987-01-25
R
Delgado & G., E. S. S., E. G. L., M. A. Gonzalez
167
168
10.505
Estanque Ayala
77
-66.763
166
167
MIZA 105594, MAGS 1030
13
3
7
Miranda
paratype
1989-07
R
Delgado
10.367
Los Amarillos
77
-66.945
Caracas-Charallave
167
168
MIZA 105668, MAGS 1165
5
2
1
Miranda
paratype
1970-12-28
CAS
Peck
1100
10.42
Hacienda La Trinidad
779
-66.85
Caracas
167
168
3
1
2
Miranda
paratype
2019-11-11
2019-11-13
2019-11-11
MIZA
Villarreal & Rodriguez
1450
10.417
Topotepuy
77
-66.851
El Volcan
167
168
2
1
Monagas
paratype
1991-10-12
R
Delgado
10.174
Cueva del Guacharo
77
-63.552
167
168
MIZA 105710, MAGS 1318
9
1
Monagas
paratype
1991-10-14
R
M. de Garcia & Delgado & Gonzalez
10.174
Nueva Esparta
77
-63.552
167
168
MIZA 105752, MAGS 1313
6
2
1
Nueva Esparta
paratype
1986-12
MIZA, MAGS
Contreras
11.047
Cerro Copei
77
-63.891
167
168
MIZA 105780, MAGS 1007
1
1
Portuguesa
paratype
1995-07-07
R
Delgado
Paez
9.52
Acarigua
779
-69.15
Acarigua
167
168
MIZA 105574, MAGS 1400
2
1
1
Tachira
paratype
2020-02-10
ZFMK
Arias C.
Ar
1300
7.9236
Trujillo
7
-71.7152
Ven
167
168
9
3
5
Trujillo
paratype
2018-11-22
ZFMK
Huber & Villarreal
1820
9.3945
Ven
7
-70.2674
Ar
167
168
12
5
7
Trujillo
paratype
1993-12-26
MIZA, MAGS
Gonzalez & M. A. Gonzalez
9.235
Miticun
77
-70.264
Bocono
167
168
MIZA 105762, MAGS 1386
2
1
Trujillo
paratype
1986-12-27
R
Manzanilla
9.284
Hacienda La Encomienda
77
-70.317
Bocono
167
168
MIZA 105778, MAGS 1011
2
1
1
Trujillo
paratype
2005-05-14
MIZA
O. Villarreal & H. Escalona
650
La Gira
Buena Vista
167
168
MIZA 1809
1
1
Trujillo
paratype
among rocks
2020-02-13
ZFMK
Huber & Villarreal M. & Q. Arias & La Guaira
Ar
150
9.7562
El Encanto
7
-70.7418
Ven
167
168
2
1
Trujillo
paratype
2018-11-09
ZFMK
Huber & O. Villarreal
Ar
1535
10.4566
El Limon
7
-67.2548
Ven
167
168
5
1
4
Trujillo
paratype
1986-11-26
R
Delgado
10.475
Hacienda El Limon
77
-67.283
167
168
MIZA 105742, MAGS 993
9
2
4
Trujillo
paratype
1982-02-12
MIZA, MAGS
Gonzalez
10.475
Hacienda El Limon
77
-67.283
167
168
MIZA 105777, MAGS 983
6
2
3
Trujillo
paratype
forest remnant along small stream
2020-02-21
ZFMK
Huber & O. Villarreal
600
10.4788
El Limon
7
-67.301
Ar
167
168
2
2
Trujillo
paratype
forest along stream
2020-02-21
ZFMK
Huber & O. Villarreal
Ar
1235
10.4774
El Limon
7
-67.2819
Ven
167
168
6
3
2
Trujillo
paratype
1996-06-26
MIZA, MAGS
Gonzalez
10.55
Galipan
779
-66.89
167
168
MIZA 105583, MAGS 1412
2
1
Yaracuy
paratype
1986-09-11
MIZA, MAGS
Yustiz
10.48
Yurubi National Park
779
-68.66
167
168
MIZA 105641, MAGS 1085
1
1
Yaracuy
paratype
" tunel de la casa de montana "
1998-12-22
MIZA
Villarreal
10.422
Minas de Aroa
77
-68.878
167
168
MIZA 1810
1
1
Yaracuy
paratype
forest along stream
2020-02-16
ZFMK
Huber & Villarreal & Q. Arias
Ar
140
10.4913
Yurubi National Park
7
-68.6564
Ven
167
168
12
5
6
Yaracuy
paratype
forest along stream
2020-02-16
ZFMK
Huber & O. Villarreal & Q. Arias
120
10.2951
Guaquira
7
-68.6535
Ar
167
168
2
2
Yaracuy
paratype
forest along stream
2020-02-17
ZFMK
Huber & O. Villarreal & Q. Arias
150
10.2807
Guaquira
7
-68.653
Ar
167
168
7
4
3
Yaracuy
paratype