Olmi 1994c: 7 Azevedo et al . 2010: 882 Guglielmino & Olmi 2015: 254 Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea) Olmi, Massimo Copeland, Robert S. Noort, Simon Van Zootaxa 2019 2019-07-10 4630 1 1 619  Guglielmino & Olmi 2015: 254  Olmi, 1994 Olmi 1994 [195,601,513,540] Insecta Dryinidae Deinodryinus GBIF Animalia Hymenoptera 252 253 Arthropoda species steineri  ( Figs 89A–C)     Deinodryinus steineri  Olmi 1994c: 7;  Azevedo et al. 2010: 882;  Guglielmino & Olmi 2015: 254.   Description. ♂. Fully winged; body length 2.9–3.2 mm. Head black, except mandible testaceous; antenna testa- ceous; mesosoma black; metasoma brown; legs testaceous, except pro- and metacoxa partly black; occasionally legs testaceous, except metacoxa partly black. Antenna filiform, with setae at most as long as breadth of antennomeres; antennomeres in following proportions: 11:6:11.5:8:8:7:7:7.5:7:12. Head shiny, strongly punctate (punctures large and deep, similar to areolae; some regions of head seem reticulate rugose; in specimen from Madagascar, 21°15.99’S 47°25.21’E, frons completely reticulate rugose); frontal line complete; occipital carina complete; POL = 8.5; OL = 4; OOL = 7; OPL = 6; TL = 3. Mesoscutum shiny, punctate, unsculptured among punctures. Notauli incomplete, reaching approximately 0.7–0.8 × length of mesoscutum. Mesoscutellum and metanotum shiny, punctate, unsculptured among punctures. Mesopleuron punctate, unsculptured among punctures. Metapleuron shiny, unsculptured. Metapectal-propodeal disc reticulate rugose, without strong transverse posterior keel; propodeal declivity without longitudinal keels, with central shiny unsculptured area. Forewing with dark distal transverse band; distal part of 2r-rs&Rs vein longer than proximal part (17:13). Paramere ( Figs 89A-C) with one large apical band wrapping penis. Tibial spurs 1/1/2.  ♀. Unknown.   Material examined. Type:  ♂ holotype: MADAGASCAR: Fianarantsoa,  7 kmW Ranomafana,  900 m,  1– 7.III.1990, MT, small clearing, montane rain forest, W.E. Steinerleg. ( USNM). Other material: MADAGASCAR: Antsiranana, Montagne d’Ambre National Park, 12°30.52’S 49°10.53’E,  960 m,  21–26.I.2001, MT, M.E. Irwin, E.I. Schlinger& R. Harin’Halaleg., MA-01-01A-01,  1♂( CAS); Antsiranana, Marojejy National Park,  5 kmW Manantenina village, Camp Mantella, 14°26.29’S 49°46.44’E,  490 m,  28.IV–7.V.2005, MT, low altitude rainforest, M. Irwin& R. Harin’Halaleg., MA-31-20,  1♂( MOLC); same locality label,  28.VIII–11.IX.2005, MA-31-29,  1♂( CAS); same locality label,  18–25.III.2005, MA-31-16,  1♂( CAS); Fianarantsoa, Ranomafana National Park, Belle Vueat Talatakely, 21°15.99’S 47°25.21’E,  1020 m,  15–22.XI.2001, MT, secondary tropical forest, R. Harin’Halaleg., MA-02-09C-03,  1♂( CAS); same locality label,  16.X–8.XI.2001, MA-02-09C-01,  2♂♂( CAS, MOLC); Fian- arantsoa, Ranomafana National Park, radio tower at forest edge, 21°15.05’S 47°24.43’E,  1130 m,  16.X–8.XI.2001, MT, mixed tropical forest, R. Harin’Halaleg., MA-02-09B-01,  1♂( MOLC); Fianarantsoa, Ranomafana National Park, Vohiparara, 21°13.57’S 47°22.19’E, at broken bridge,  1110 m,  14–21.I.2002, MT, high altitude rainforest, R. Harin’Halaleg., MA-02-09A-12,  1♂( CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21°13.67’S 47°22.19’E, at broken bridge,  1110 m,  12–19.II.2002, MT, high altitude rainforest, R. Harin’Halaleg., MA-02- 08A-16,  1♂, 1 gynander ( CAS); Toamasina, Andasibe National Park, 18°55.58’S 48°24.47’E, botanic garden near entrance to park,  1025 m,  1–5.IX.2001, MT, tropical forest, R. Harin’Halaleg., MA-01-08B-11,  2♂♂( CAS); Toamasina, Sainte Marie Island, Ambohidena Forest,  22.8 km44° NW Ambodifotatra, 16°49.28’S 49°57.51’E,  20 m,  21–24.XI.2005, MT, littoral rainforest, Brian L. Fisher et al. leg., BLF12842,  1♂( CAS).   Hosts. Unknown.   Distribution. Madagascar.   Remarks. One gynander specimen of  D. steineriis known from Madagascar, Fianarantsoa, Ranomafana National Park, Vohiparara, 21°13.67’S 47°22.19’E(CAS). Description: general aspect of a ♂, without chelae. Fully winged; body length 3.7 mm. Head black, except mandible testaceous (with brown teeth) and anterior half of clyp- eus testaceous; antenna testaceous, except antennomeres 8–10 brown; mesosoma black; metasoma brown-black; legs testaceous-ferruginous. Antenna filiform, setose; antennomeres in following proportions: 10:5:6:11:10:10:9:8:8:11. Head dull, reticulate rugose and with many irregular longitudinal keels; frontal line absent; occipital carina complete; POL = 10; OL = 5; OOL = 7; OPL = 8; TL = 7; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, reticulate rugose and with many irregular longitudinal keels. Notauli incomplete, reaching about 0.5 × length of mesoscutum. Mesoscutellum and metanotum dull, reticulate rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing completely slightly darkened. Distal part of 2r-rs&Rs vein much longer than proximal part (29:11). Specimen with ovipositor, without ♂genitalia. This specimen is attributed tentatively to  D. steineri, because collected together with one normal ♂specimen in the same locality. The reticulate rugose sculpture of the mesoscutum is not present in any species collected in the same locality (  D. harinhalai,  madecassus,  pulcher,  rusticus,  steineri,  suavis). The above sculpture is possibly present in the still unknown ♀of  D. steineri. 2275276802 1990-03-01 2001-01-26 1990-03-01 USNM W. E. Steiner & M. E. Irwin & E. I. Schlinger & R. Harin'Hala Madagascar 900 -12.508667 Montagne d'Ambre National Park 13 49.1755 252 253 2 1 Fianarantsoa holotype 2275277079 2005-04-28 2005-05-07 2005-04-28 CAS M. Irwin & R. Harin'Hala Madagascar 490 -14.438167 Camp Mantella 13 49.774 Marojejy National Park 252 253 1 1 Antsiranana holotype 2275276974 [645,1389,1371,1398] 2005-08-28 2005-09-11 2005-08-28 MOLC M. Irwin & R. Harin'Hala Madagascar 490 -14.438167 Camp Mantella 13 49.774 Marojejy National Park 252 253 1 1 Antsiranana holotype 2275277053 2005-03-18 2005-03-25 2005-03-18 CAS M. Irwin & R. Harin'Hala Madagascar 490 -14.438167 Camp Mantella 13 49.774 Marojejy National Park 252 253 1 1 Antsiranana holotype 2275278127 2001-11-15 2001-11-22 2001-11-15 CAS R. Harin'Hala Madagascar Ranomafana National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 252 253 1 1 Fianarantsoa holotype 2275277565 [395,1122,1479,1506] 2001-10-16 2001-11-08 2001-10-16 CAS R. Harin'Hala Madagascar Ranomafana National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 252 253 1 1 Fianarantsoa holotype 2275277621 2001-10-16 2001-11-08 2001-10-16 CAS, MOLC R. Harin'Hala Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 252 253 2 2 Fianarantsoa holotype 2275276988 2002-01-14 2002-01-21 2002-01-14 MOLC R. Harin'Hala Madagascar 1110 -21.226166 Vohiparara 12 47.369835 Ranomafana National Park 252 253 1 1 Fianarantsoa holotype 2275277680 2002-02-12 2002-02-19 2002-02-12 CAS R. Harin'Hala Madagascar 1110 -21.227833 Vohiparara 12 47.369835 Ranomafana National Park 252 253 1 1 Fianarantsoa holotype 2275277595 2001-09-01 2001-09-05 2001-09-01 CAS R. Harin'Hala Madagascar 1025 -18.926332 Andasibe National Park 13 48.407833 252 253 1 1 Toamasina holotype 2275277225 2005-11-21 2005-11-24 2005-11-21 CAS Brian L. Fisher Madagascar 20 -16.821333 Ambohidena Forest 13 49.9585 Sainte Marie Island 252 253 2 2 Toamasina holotype 2275276799 [1058,1176,1804,1830] CAS Madagascar 252 253 1 1 Toamasina holotype