Olmi 2004d: 88 Azevedo et al . 2010: 882 Guglielmino & Olmi 2015: 251 Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea) Olmi, Massimo Copeland, Robert S. Noort, Simon Van Zootaxa 2019 2019-07-10 4630 1 1 619  Guglielmino & Olmi 2015: 251  Olmi, 2004 Olmi 2004 [195,608,1903,1930] Insecta Dryinidae Deinodryinus GBIF Animalia Hymenoptera 247 248 Arthropoda species rusticus  ( Figs 88A–D, 95B)     Deinodryinus rusticus  Olmi 2004d: 88;  Azevedo et al. 2010: 882;  Guglielmino & Olmi 2015: 251. Description. ♂( Fig. 95B). Fully winged; body length 2.9–4.6 mm( holotype 2.9 mm); paratypesfrom Madagascar, 18°28.24’S 47°57.36’E, longer than holotype( 3.8–4.6 mm). Colour of holotype: head black, except mandible testaceous; antenna testaceous; mesosoma and metasoma black; legs testaceous, except coxae, clubs of femora and part of metatibia brown; paratypesfrom Madagascar, 18°28.24’S 47°57.36’E, with legs completely testaceous, except for brown basal spot on metacoxa. Usually large specimens with legs totally or mostly testaceous, small specimens with legs mostly brown. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 11:4:11:7:8:7:6.5:6.5:7:9. Head shiny, strongly punctate, unsculptured among punctures (head punctures very strong; head almost rugose, mainly on frons; in many specimens from Madagascarfrons reticulate rugose); vertex without two oblique keels from lateral ocelli to occipital carina; frontal line usually almost complete, missing only in front of anterior ocellus; frontal line occasionally only visible in anterior half of frons; occipital carina complete; POL = 8; OL = 4; OOL = 6.5; OPL = 9; TL = 5; in some specimens OPL shorter than OOL, or approximately as long as OOL. Mesoscutum shiny, as sculptured as head. Notauli incomplete, usually reaching approximately 0.5–0.6 × length of mesoscutum; rarely notauli reaching 0.8–0.9 × length of mesoscutum. Mesoscutellum shiny, punctate, unsculpured among punctures. Metanotum shiny, unsculptured. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity almost completely reticulate rugose, with small central smooth and shiny area, without longitudinal or transversal keels; occasionally propodeal declivity with two tracks of longitudinal keels and median area completely dull and reticulate rugose. Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein slightly longer than proximal part (13:12), or as long as proximal part. Paramere ( Fig. 88A) with slender and long distal inner branch (branch showing mosaic sculpture ( Fig. 88A) in holotypeand in specimen from Madagascar, 12°17.17’S 49°22.00’E; mosaic sculpture not visible in few paratypesfrom Madagascar, 18°28.24’S 47°57.36’Eand 24°39.13’S 43°59.48’E( Figs 88B, C); in almost all specimens inner side of paramere with large area sculptured by numerous papillae. In specimen from Madagascar, 24°14.26’S 46°17.16’E, distal inner branch very short ( Fig. 88D). Tibial spurs 1/1/2.  ♀. Unknown.   Material examined. Types:  ♂ holotype: MADAGASCAR: Mahajanga, Ankoririka Reserve, 10.6 km13° NE of Tsaramandroso, 16°16.2’S 46°2.55’E,  210 m,  9–14.IV.2001, MT, tropical dry forest,  Fisher, Griswold et al. leg., BLF 3665( CAS).  Paratypes: MADAGASCAR: Antananarivo,   11.5 km47° SSE of Anjozorobe,   3 km41° NE of Andranomay, 18°28.24’S 47°57.36’E,  1300 m,  5–13.XII.2000, MT, montane rainforest,  Fisher, Griswold et al. leg., BLF 2375,  6♂♂( 4 in CAS, 2 in MOLC). Other material: MADAGASCAR: Antananarivo,  46 kmNE of Ankazobe, Ambohitantely, 18°11.88’S 47°16.89’E,  700 m,  15.X–1.XI.2004, MT, sclerophyl forst, M. Irwin & R. Harin’Hala leg., MA-27-19,  1♂( MOLC); Antsiranana,  3 kmW of Sakalava Beach, white dunes, 12°17.17’S 49°22.00’E,  40 m,  27–30.I.2001, MT, MA- 01-03-03, M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg.,  1♂( CAS); same locality label,  21–23.I.2001,MA- 01-03-01,  3♂♂( CAS); Antsiranana, Sakalava Beach, 12°15.46’S 49°23.51’E,  10 m,  27.IV–13.V.2001, dwarf littoral forest, MT, across sandy trail, MA-01-04B-08, M.E. Irwin & R. Harin’Hala leg.,  1♂( CAS); Antsiranana, Binara Forest,  7.5 km230° SW Daraina, 13°15.18’S 49°37.00’E,  375 m,  1.XII.2003, MT, tropical dry forest, B.L. Fisherleg., BLF9557,  7♂♂( CAS); Antsiranana, SAVA Region, Vohemar District, Antsahabelelarain forest,  9 kmSW Daraina, Binara, 13°15.03’S 49°37.00’E,  182 m,  17–24.XI.2010, MT, in humid forest, M. Irwin, R. Harin’Hala leg., MG-58-07,  1♂( CAS); same locality label,  8–15.XII.2010, MG-58-10,  1♂( CAS); same locality label,  11–19.I.2011, MG-58-15,  1♂( CAS); Antsiranana, SAVA Region, Vohemar District, Andranotsimatydry forest,  9 kmNE Daraina, 13°10.17’S 49°42.04’E,  90 m,  14–21.XI.2010, MT, dry dense forest, M. Irwin, R. Harin’Hala leg., MG-57-07,  1♂( CAS); same locality label,  19–26.XII.2010, MG-57-12,  2♂♂( CAS, MOLC); same locality label,  14–21.II.2011, MG-57-19,  1♂( CAS); same locality label,  11-18.IV.2011, MG-57-27,  3♂♂( 2 in CAS, 1 in MOLC); Antsiranana, Diana Region, Orangeadry forest, Ramena, Baiede dune,  900 mE of Camp Minier,  1–8.VI.2011, 12°13.97’S 49°21.99’E, M. Irwin, R.Harin’Hala leg., MT, dry forest, elev. 152 m, MG- 62-25,  1♂( CAS); same locality label,  12–19.I.2011, MG-62-05,  1♂( CAS); Antsiranana, Ambato Forest,  26.6 km33° NE Ambanja, 13°27.52’S 48°33.06’E,  150 m,  8.XII.2004, MT, rainforest, B.L. Fisherleg., BLF11518,  1♂( CAS); Antsiranana, Montagne d’Ambre National Park, 12°31.13’S 49°10.45’E,  1125 m,  12–14.V.2001, MT, MA- 01-01D-10, R. Harin’Hala leg.,  3♂♂( CAS); same locality label,  14–30.V.2001, R. Harin’Hala leg., MA-01-01D- 11,  1♂( CAS); Antsiranana, Orangea,  3 kmE of Ramenanear fort, 12°14.49’S 49°22.17’E,  65 m,  23–27.I.2001, MT, littoral forest on sand, MA- 01-05-02, M.E. Irwin, E.I. Schlinger & R. Harin’Hala leg.,  1♂( CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21°15.05’S 47°24.43’E,  1130 m,  16.X–8.XI.2001, MT, mixed tropical forest, R. Harin’Hala leg., MA-02-09B-01,  3♂♂( 2 in CAS, 1 in MOLC); same locality label,  4– 12.III.2002,  1♂( CAS); same locality label, 21°15.05’S 47°24.43’E,  1130 m,  15–25.VII.2002, MA-02-09B-35,  1♂( CAS); same locality label,  24.V–4.VI.2002, MA-02-09B-30,  5♂♂( CAS); same locality label,  7–17.V.2003, MA- 02-09B-60,  1♂( CAS); same locality label,  27.VI–7.VII.2003, MA-02-09B-65,  1♂( CAS); Fianarantsoa, Rano- mafana National Park, Belle Vueat Talatakely, 21°15.99’S 47°25.21’E,  1020 m,  19–26.II.2002, MT, secondary tropical forest, R. Harin’Hala leg., MA-02-09C-17,  3♂♂( CAS); same locality label,  15–28.V.2003, MA-02-09C- 61,  1♂( MOLC);same locality label,  28.IV–5.V.2002, MA-02-09C-27,  1♂( CAS);same locality label  22–28.XI.2001, MA-02-09C-04,  2♂♂( CAS, MOLC); same locality label,  16.X–8.XI.2001, MA-02-09C-01,  1♂( CAS); Fianarant- soa, Ranomafana National Park, Vohiparara, 21°13.57’S 47°22.19’E, at broken bridge,  1110 m,  15–25.VII.2002, MT, high altitude rainforest, R. Harin’Hala leg., MA-02-09A-35,  5♂♂( 3 in CAS, 2 in MOLC); same locality label,  14–21.I.2002,  2♂♂( CAS, MOLC); same locality label,  15–21.XII.2001, MA-02-09A-07,  1♂( CAS), 1 gynander specimen ( MOLC); same locality label,  26–31.III.2002, MA-02-09A-22,  3♂♂( CAS); Fianarantsoa, Ranomafana National Park, Vohiparara, 21°13.67’S 47°22.19’E, at broken bridge,  1110 m,  25.VII–3.VIII.2002, MT, high altitude rainforest, R. Harin’Hala leg., MA-02-08A-36,  1♂( CAS); Fianarantsoa, Befotaka-Midongy National Park, Papango  27.7 kmS Midongy-South Mt. Papango, 23°50.07’S 46°57.49’E,  940 m,  13–19.XI.2006, MT, B.L.  Fisher et al. leg., BLF14774,  1♂( CAS); Mahajanga, Namoroka National Park,  16.9 km317° NW Vilanandro, 16°24.24’S 45°18.36’E,  100 m,  12–16.XI.2002, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 6581,  6♂♂( CAS); Mahajanga, Namoroka National Park, 16.40494S 45.31294E,  21–31.X.2016, MT, C. Villemant leg.,  1♂( MNHN); Mahajanga, Sofia District,  45 kmS Antsohihy,  5 kmW of Anjiamangirana, elev  97 m,  5–8 May 2011, 15° 09.42’S 47° 47.05’E, M.Irwin, R.Harin’Hala, MT, Analagnambe dry degraded forest, MG-61-11,  1♂( CAS); same locality label,  2–6.I.2011, MG-61-04,  1♂( CAS); Mahajanga, Beanka Forest Reserve,  50.2 kmE Maintirano, 18°01’35’’S 44°03’02’’E,  250 m,  19–26.X.2009, tropical dry forest on tsingy, MT, B.L.  Fisher et al. leg., BLF22787,  2♂♂( CAS); Mahajanga, Analamanitra Forest,  14 kmNE Misinjo, 15°55’S 45°57’E,  65 ft.,  23–30.X.2007, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG-38-06,  1♂( CAS); same locality label,  12–19.II.2008, MG-38-18,  1♂( CAS); Mahajanga, Bemarivo Special Reserve,  23.8 km223° SW Besalampy, 16°55.30’S 44°22.06’E,  19– 23.XI.2002, 30 m, sifted litter (leaf mold, rotten wood), in tropical dry forest, Fisher, Griswold et al. leg., BLF6693,  4♂♂( CAS); Mahajanga, Ankarafantsika National Park, Tsimaloto Forest,  18.3 km49° NE Tsaramandroso, 16°13’41’’S 46°08’37’’E, 135 M,  2–8.IV.2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF3600,  5♂♂( 4 in CAS, 1 in MOLC); Mahajanga, Besalampy District, Marofototrapalm forest,  17 kmW of Besalampy, 16°43.30’S 44°25.42’E,  35 ft.,  24.IX–1.X.2007, MT, palm trees on sand, M. Irwin, R. Harin’Hala leg., MG-42B-01,  1♂( CAS); same locality label, Marofototradry forest,  170 ft.,  19–26.XI.2007, dry wash in forest, MG-42A-09,  1♂( CAS); same locality label, Marofototradry forest,  170 ft.,  1–8.X.2007, dry wash in forest, MG-42A-02,  1♂( MOLC); Mahajanga, Tsingy de Bemaraha National Park,  3.4 km93° E Bekopaka, Tombeau Vazimba, 19°08.31’S 44°49.41’E,  50 m,  6–10.XI.2001, MT, tropical dry forest, Fisher, Griswold et al. leg., BLF 4233,  1♂( CAS); Toama- sina, Andasibe National Park, 18°55.58’S 48°24.47’E, botanic garden near entrance to park,  1025 m,  16–23.XI.2001, MT, tropical forest, R. Harin’Hala leg., MA-01-08B-20,  1♂( CAS); Toliara, Mahafaly Plateau,  6.2 km74° ENE of Itampolo, 24°39.13’S 43°59.48’E,  80 m, MT, spiny forest thicket,  21–25.II.2002,   Fisher, Griswold et al. leg., BLF5759,  2♂♂( CAS, MOLC); Toliara, Andohahela Nat. Park, Manantalinjo Forest,  33.6 km63° ENE Amboasary,  7.6 km99° E Hazofotsy, 24°49.1’S 46°36.36’E,  150 m,  12–16.I.2002, MT, spiny forest thicket, Fisher, Griswold et al. leg., BLF 4811,  15♂♂( 13 in CAS, 2 in MOLC); Toliara, Beza Mahafaly Reserve, parcelle I near research sta- tion, 23°41.19’S 44°35.46’E,  165 m,  1–8.II.2002, MT, dry deciduous forest, R. Harin’Hala leg., MA-02-14A-14,  2♂♂( CAS, MOLC); same locality label,  25.XII.2001–  2.I.2002,  1♂( CAS); same locality label,  9–20.IX.2002,  3♂♂( 2 in CAS, 1 in MOLC); same locality label,  23.III–2.IV.2003,  1♂( CAS); same locality label,  29.III–10.IV.2002,  1♂( CAS); Toliara, Beza Mahafaly Reserve, parcelle II near Bellevue, 23°41.39’S 44°34.53’E,  180 m,  15.X– 10.XI.2001, MT, spiny forest, M.E. Irwin, F.D. Parker & R. Harin’Hala leg., MA-02-14B-01,  3♂♂( 1 in CAS, 2 in MOLC); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24°56.21’S 46°37.60’E,  180 m,  16–17.XII.2002, MT, transitional forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02-20-09,  1♂( CAS); same locality label,  10–20.VII.2003,  2♂♂( CAS); Toliara, Mikea Forest, NW of Manombo, 22°54.22’S 43°28.53’E,  30 m,  20– 27.XI.2001, MT, deciduous dry forest, R. Harin’Hala leg., MA-02-18A-03,  1♂( CAS); same locality label,  23.IX– 3.X.2002, MA-02-18A-34,  2♂♂( MOLC); same locality label,  19.II–1.III.2002,  2♂♂( CAS, MOLC); same local- ity label,  23.VII–6.VIII.2002,  1♂( CAS); same locality label,  6–16.I.2002, MA-02-18A-09,  1♂( CAS); same locality label,  17–28.I.2002, MA-02-18A-11,  3♂♂( 2 in CAS, 1 in MOLC); Toliara, Mikea Forest, NWof Manom- bo, 22°54.80’S 43°28.93’E,  37 m,  23.VII–6.VIII.2002, MT, spiny forest, R. Harin’Hala leg., MA-02-18B-29,  1♂( CAS); same locality label,  8–18.III.2002, MA-02-18B-16,  1♂( CAS); Toliara, Andohahela National Park, Ihazo- fotsy ParcelleIII, 24°49.85’S 46°32.17’E,  80 m,  20.V–3.VI.2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02-21-20,  4♂♂( 3 in CAS, 1 in MOLC); same locality label,  13–24.II.2003, MA- 02-21-10,  1♂( CAS); Toliara, Andohahela National Park, Ihazofotsy ParcelleIII, 24°50.09’S 46°29.21’E,  80 m,  7–17.XII.2003, MT, dry spiny forest, M. Irwin, F. Parker & R. Harin’Hala leg., MA- 02-21-41,  1♂( CAS); Toliara, Berenty Special Reserve,  8 kmNW Amboasary, 25°00.40’S 46°18.20’E,  85 m,  24.III–3.IV.2003, MT, gallery forest, M. Irwin, F. Parker, R. Harin’Hala leg., MA- 02-22-20,  2♂♂( CAS); same locality label,  27.XII.2002–  7.I.2003,  1♂( CAS); same locality label,  25.II–7.III.2003,  2♂♂( CAS); same locality label,  26.I–5.II.2003, MA- 02-22-14,  1♂( CAS); Toliara, Zombitse National Park, near ANGAP office, 22°53.19’S 44°41.53’E,  840 m,  9–19.XI.2001, MT, deciduous spiny forest, M.E. Irwin, F.D. Parker & R. Harin’Hala leg., MA-02-13A-02,  1♂( CAS); Toliara, Ivohibe Forest,  55 kmN Tolagnaro, 24°34.08’S 47°12.14’E,  200 m,  2–4.XII.2006, MT, B.L.  Fisher et al. leg., BLF15448,  4♂♂( 3 in CAS, 1 in MOLC); Toliara, Vohidava Forest,  88.9 kmN Amboasary, 24°14.26’S 46°17.16’E,  500 m,  6–8.XII.2006, B.L.  Fisher et al. leg., BLF15694,  1♂( MOLC); Toliara, Androimpano Forest,  3 kmE of Itampolo, 24°39.47’S 43°57.37’E,  45 m,  25.XII.2008–  1.I.2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG-54B-09,  2♂♂( CAS); same locality label,  6–13.XI.2008, MG-54B-03,  8♂♂( 6 in CAS, 2 in MOLC); Toliara, Androimpano Forest,  5 kmE of Itampolo, 24°39.02’S 43°57.79’E,  130 m,  14–21.V.2009, MT, path in dry forest, M. Irwin, R. Harin’Hala leg., MG- 54C-27,  7♂♂( CAS); Toliara, Ankazomena,  8 kmN of Ambohimahavelona village, 23°25.82’S 43°50.05’E,  122 m,  21–31.X.2008, MT, dry forest, M. Irwin, R. Harin’Hala leg., MG-51-01,  2♂♂( CAS); Toliara, Ranobe, 23°02.783’S 43°36.617’E,  20 m,  27.X–3.XI.2003, MT, spiny forest thicket, MGF082, Frontier Wilderness Project,  3♂♂( CAS); Toliara, Ifaty, near Hotel Paradisia, 23°10.78’S 43°37.01’E,  9 m,  26.V–5.VI.2002, in coastal dunes, MT, vegetation in sandy area, R. Harin’Hala leg., MA- 02-16-28,  1♂( CAS); same locality label,  13.X-12.XI.2001, MA- 02-16-01,  1♂( CAS); Toliara, Makay Mts., 21°13.11’S, 45°19.26’E,  500 m,  24.XI–12.XII.2010, MT, gallery forest, sandy soil, B.L.  Fisheret al. leg., BLF25357,  1♂( CAS); Toliara, Tsifota, 20 kmN Manombo,  1 kmE of Tsifota village, 22°49.08’S 43°22.36’E,  15 m,  30.X–9.XI.2008, MT, spiny forest, M. Itwin, R. Harin’Hala leg., MG-48-03,  2♂♂( CAS); Toliara,  5 kmSE Manombo, Andranovato, 22°49.08’S 43°30.13’E,  18 m,  20–30.X.2008, MT, in  Euphorbiaforest, M. Irwin, R. Harin’Hala leg., MG-49-01,  3♂♂( CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Forest, 24°02.91’S 43°45.14’E,  120 m,  24–29.X.2008, MT, dense dry forest, M. Irwin, R. Harin’Hala leg., MG- 53A-01,  6♂♂( CAS); same locality label,  4–12.XII.2008, transitional forest, MG-53C-07,  2♂♂( CAS); same local- ity label,  24–29.X.2008, across trail at base of escarpment, dense dry forest, MG-53B-01,  1♂( CAS).   Hosts. Unknown.   Distribution. Madagascar.   Remarks. A gynander specimen has been collected in the following locality: MADAGASCAR: Ranomafana National Park, Vohiparara, 21°13.57’S 47°22.19’E, at broken bridge, 1110 m, 15–21.XII.2001, MT, high altitude rainforest, R. Harin’Hala leg., MA-02-09A-07, one gynander specimen (MOLC). Description: general aspect of ♂; fully winged; body length 3.9 mm; head black, except mandible testaceous, clypeus testaceous, exept posterior margin black, one testaceous spot on malar space; antenna testaceous-ferruginous; mesosoma black; metasoma brown; legs testaceous-ferruginous. Antenna filiform, with setae shorter than breadth of antennomeres; antennomeres in following proportions: 9:6:8:11:10:10:8:8:7:11. Head dull, strongly rugose and sculptured by many irregular longitudinal keels; vertex without two oblique keels from lateral ocelli to the occipital carina; frontal line absent; occipital carina complete; POL =11; OL = 5; OOL = 9; OPL = 8.5; TL = 6; greatest breadth of lateral ocelli about as long as OL. Mesoscutum dull, as sculptured as head. Notauli incomplete, reaching approximately 0.25 × length of mesoscutum; notauli hardly visible among longitudinal keels. Mesoscutellum shiny, partly rugose and partly sculptured by deep punctures, similar to areolae. Metanotum rugose. Metapectal-propodeal disc dull, reticulate rugose, without transverse posterior keel; propodeal declivity completely reticulate rugose, without longitudinal or transverse keels. Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein much longer than proximal part (30:13). ♂genitalia absent. Ovipositor present. Tibial spurs 1/1/2. 2275277060 tropical dry forest 2001-04-09 2001-04-14 2001-04-09 CAS Fisher & Griswold Madagascar 210 -16.27 Ankoririka Reserve, 10.6 km 13 ° NE of Tsaramandroso 92 46.0425 Fisher 248 249 BLF 3665 1 1 Mahajanga holotype 2275276930 2001-04-09 2001-04-14 2001-04-09 CAS Madagascar 210 -16.27 Ankoririka Reserve 92 46.0425 Fisher 248 249 1 1 Mahajanga paratype 2275277732 Madagascar Fisher 248 249 1 Antananarivo paratype 2275277581 2004-10-15 2004-11-01 2004-10-15 CAS, MOLC Madagascar 700 -18.198 Ambohitantely 13 47.2815 248 249 6 6 Antananarivo 2275276952 2001-01-27 2001-01-30 2001-01-27 MOLC Madagascar 40 -12.286167 3 km W of Sakalava Beach 13 49.366665 248 249 1 1 Antsiranana 2275276970 2001-01-21 2001-01-23 2001-01-21 CAS Madagascar 40 -12.286167 3 km W of Sakalava Beach 13 49.366665 248 249 1 1 Antsiranana 2275277462 2001-04-27 2001-05-13 2001-04-27 CAS Madagascar 10 -12.257667 Sakalava Beach 13 49.391834 248 249 3 3 Antsiranana 2275277155 [206,1432,1412,1439] 2003-12-01 CAS Madagascar 375 -13.253 Binara Forest 13 49.616665 248 249 1 1 Antsiranana 2275276695 2010-11-17 2010-11-24 2010-11-17 CAS Madagascar Region 182 -13.2505 Antsahabelela 13 49.616665 Vohemar District 248 249 7 7 Antsiranana 2275276541 [716,1250,1520,1547] 2010-12-08 2010-12-15 2010-12-08 CAS Madagascar Region 182 -13.2505 Antsahabelela 13 49.616665 Vohemar District 248 249 1 1 Antsiranana 2275277287 2011-01-11 2011-01-19 2011-01-11 CAS Madagascar Region 182 -13.2505 Antsahabelela 13 49.616665 Vohemar District 248 249 1 1 Antsiranana 2275277222 2010-11-14 2010-11-21 2010-11-14 CAS Madagascar Region 90 -13.1695 Andranotsimaty 13 49.70067 Vohemar District 248 249 1 1 Antsiranana 2275276826 [620,1157,1629,1655] 2010-12-19 2010-12-26 2010-12-19 CAS Madagascar Region 90 -13.1695 Andranotsimaty 13 49.70067 Vohemar District 248 249 1 1 Antsiranana 2275276620 2011-02-14 2011-02-21 2011-02-14 CAS, MOLC Madagascar Region 90 -13.1695 Andranotsimaty 13 49.70067 Vohemar District 248 249 2 2 Antsiranana 2275277138 [784,1298,1665,1691] 2011-04-11 2011-04-18 2011-04-11 CAS Madagascar Region 90 -13.1695 Andranotsimaty 13 49.70067 Vohemar District 248 249 1 1 Antsiranana 2275276763 2011-06-01 2011-06-08 2011-06-01 CAS, MOLC Baie Madagascar Diana Region 900 Ramena Orangea 248 249 3 3 Antsiranana 2275277486 [231,738,1773,1799] 2011-01-12 2011-01-19 2011-01-12 CAS Baie Madagascar Diana Region 900 Ramena Orangea 248 249 1 1 Antsiranana 2275277597 2004-12-08 CAS Madagascar 150 -13.458667 Ambato Forest 13 48.551 248 249 1 1 Antsiranana 2275277022 [882,1387,1808,1835] BLF B. L. Fisher Madagascar 248 249 BLF11518 1 Antsiranana 2275276673 2001-05-12 2001-05-14 2001-05-12 CAS Madagascar 1125 -12.518833 Montagne d'Ambre National Park 13 49.174168 248 249 1 1 Antsiranana 2275276879 [512,1051,1881,1907] 2001-05-14 2001-05-30 2001-05-14 CAS Madagascar 1125 -12.518833 Montagne d'Ambre National Park 13 49.174168 248 249 3 3 Antsiranana 2275277336 [194,1432,1916,1943] 2001-01-23 2001-01-27 2001-01-23 CAS Madagascar 65 -12.2415 Orangea 13 49.3695 248 249 1 1 Antsiranana 2275277244 2001-10-16 2001-11-08 2001-10-16 CAS Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 248 249 1 1 Fianarantsoa 2275277066 2002-03-04 2002-03-12 2002-03-04 CAS, MOLC Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 249 250 3 3 Fianarantsoa 2275276906 [291,1201,188,215] 2002-07-15 2002-07-25 2002-07-15 CAS Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 249 250 1 1 Fianarantsoa 2275277434 2002-05-24 2002-06-04 2002-05-24 CAS Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 249 250 1 1 Fianarantsoa 2275277703 [850,1370,224,251] 2003-05-07 2003-05-17 2003-05-07 CAS Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 249 250 5 5 Fianarantsoa 2275277206 [290,863,261,287] 2003-06-27 2003-07-07 2003-06-27 CAS Madagascar 1130 -21.250834 Ranomafana National Park 12 47.407166 249 250 1 1 Fianarantsoa 2275277051 2002-02-19 2002-02-26 2002-02-19 CAS Madagascar National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 249 250 1 1 Fianarantsoa 2275276318 [738,1276,332,359] 2003-05-15 2003-05-28 2003-05-15 CAS Madagascar National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 249 250 3 3 Fianarantsoa 2275277131 [190,735,368,395] 2002-04-28 2002-05-05 2002-04-28 MOLC Madagascar National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 249 250 1 1 Fianarantsoa 2275276442 [930,1432,369,395] 2001-11-22 2001-11-28 2001-11-22 CAS Madagascar National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 249 250 1 1 Fianarantsoa 2275277090 [340,992,405,431] 2001-10-16 2001-11-08 2001-10-16 CAS, MOLC Madagascar National Park 1020 -21.2665 Talatakely 12 47.420166 Belle Vue 249 250 2 2 Fianarantsoa 2275277123 2002-07-15 2002-07-25 2002-07-15 CAS Madagascar 1110 -21.226166 Vohiparara 12 47.369835 Ranomafana National Park 249 250 1 1 Fianarantsoa 2275277194 2002-01-14 2002-01-21 2002-01-14 CAS, MOLC Madagascar 1110 -21.226166 Vohiparara 12 47.369835 Ranomafana National Park 249 250 5 5 Fianarantsoa 2275277010 [316,973,512,539] 2001-12-15 2001-12-21 2001-12-15 CAS, MOLC Madagascar 1110 -21.226166 Vohiparara 12 47.369835 Ranomafana National Park 249 250 2 2 Fianarantsoa 2275276949 2002-03-26 2002-03-31 2002-03-26 CAS, MOLC Madagascar 1110 -21.226166 Vohiparara 12 47.369835 Ranomafana National Park 249 250 1 1 Fianarantsoa 2275277275 2002-07-25 2002-08-03 2002-07-25 CAS Madagascar 1110 -21.227833 Vohiparara 12 47.369835 Ranomafana National Park 249 250 3 3 Fianarantsoa 2275277629 2006-11-13 2006-11-19 2006-11-13 CAS Madagascar 940 -23.8345 Papango 12 46.958168 Befotaka-Midongy National Park 249 250 1 1 Fianarantsoa 2275277293 Fisher Madagascar 249 250 1 Fianarantsoa 2275277402 2002-11-12 2002-11-16 2002-11-12 CAS Madagascar 100 -16.404 Namoroka National Park 13 45.306 249 250 1 1 Mahajanga 2275277179 2016-10-21 2016-10-31 2016-10-21 CAS Madagascar -16.40494 Namoroka National Park 1 45.31294 249 250 6 6 Mahajanga 2275277116 2011-05-05 2011-05-08 2011-05-05 MNHN Madagascar 97 Sofia District 249 250 1 1 Mahajanga 2275276934 2011-01-02 2011-01-06 2011-01-02 CAS Madagascar 97 Sofia District 249 250 1 1 Mahajanga 2275278024 2009-10-19 2009-10-26 2009-10-19 CAS Madagascar 250 -18.026388 Beanka Forest Reserve 21 44.050556 249 250 1 1 Mahajanga 2275276835 [576,1366,909,935] Fisher Madagascar 249 250 1 Mahajanga 2275277617 2007-10-23 2007-10-30 2007-10-23 CAS Madagascar 20 -15.916667 Analamanitra Forest 1284 45.95 249 250 2 2 Mahajanga 2275276732 [741,1256,981,1007] 2008-02-12 2008-02-19 2008-02-12 CAS Madagascar 20 -15.916667 Analamanitra Forest 1284 45.95 249 250 1 1 Mahajanga 2275276785 2002-11-19 2002-11-23 2002-11-19 CAS Madagascar -16.921667 Bemarivo Special Reserve 13 44.36767 249 250 1 1 Mahajanga 2275277701 2001-04-02 2001-04-08 2001-04-02 CAS Madagascar -16.228056 Tsimaloto Forest 21 46.143612 Ankarafantsika National Park 249 250 4 4 Mahajanga 2275277289 2007-09-24 2007-10-01 2007-09-24 CAS, MOLC Madagascar 11 -16.721666 Marofototra 13 44.423668 Besalampy District 249 250 5 5 Mahajanga 2275277062 [151,1024,1233,1259] 2007-11-19 2007-11-26 2007-11-19 CAS Madagascar 52 -16.721666 Marofototra 13 44.423668 Besalampy District 249 250 1 1 Mahajanga 2275277432 2007-10-01 2007-10-08 2007-10-01 CAS Madagascar 52 -16.721666 Marofototra 13 44.423668 Besalampy District 249 250 1 1 Mahajanga 2275276948 2001-11-06 2001-11-10 2001-11-06 MOLC Madagascar 50 -19.1385 Tombeau Vazimba 13 44.8235 Tsingy de Bemaraha National Park 249 250 1 1 Mahajanga 2275276810 2001-11-16 2001-11-23 2001-11-16 CAS Madagascar 1025 -18.926332 Andasibe National Park 13 48.407833 249 250 1 1 Mahajanga 2275277825 2002-02-21 2002-02-25 2002-02-21 CAS Madagascar 80 -24.652166 Mahafaly Plateau 12 43.991333 249 250 1 1 Toliara 2275276583 Madagascar Griswold Fisher 249 250 1 Toliara 2275276557 2002-01-12 2002-01-16 2002-01-12 CAS, MOLC Madagascar Andohahela Nat. Park 150 -24.818333 Hazofotsy 92 46.606 Manantalinjo Forest 249 250 2 2 Toliara 2275276766 2002-02-01 2002-02-08 2002-02-01 CAS, MOLC Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 249 250 15 15 Toliara 2275277490 [151,888,1628,1655] 2001-12-25 2002-01-02 2001-12-25 CAS, MOLC Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 249 250 2 2 Toliara 2275277502 [903,1432,1629,1655] 2002-09-09 2002-09-20 2002-09-09 CAS Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 249 250 1 1 Toliara 2275277193 [151,881,1665,1691] 2003-03-23 2003-04-02 2003-03-23 CAS, MOLC Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 249 250 3 3 Toliara 2275277065 [889,1432,1665,1691] 2002-03-29 2002-04-10 2002-03-29 CAS Madagascar 165 -23.6865 Beza Mahafaly Reserve 12 44.591 249 250 1 1 Toliara 2275277498 2001-10-15 2001-11-10 2001-10-15 CAS Madagascar 180 -23.689833 Bellevue 12 44.5755 Beza Mahafaly Reserve 249 250 1 1 Toliara 2275277635 2002-12-16 2002-12-17 2002-12-16 CAS, MOLC Madagascar 180 -24.936832 Tsimelahy 12 46.626667 Andohahela National Park 249 250 3 3 Toliara 2275277978 2003-07-10 2003-07-20 2003-07-10 CAS Madagascar 180 -24.936832 Tsimelahy 12 46.626667 Andohahela National Park 249 250 1 1 Toliara 2275277343 2001-11-20 2001-11-27 2001-11-20 CAS Madagascar 30 -22.903667 Mikea Forest 12 43.4755 249 250 2 2 Toliara 2275277412 2002-09-23 2002-10-03 2002-09-23 CAS Madagascar 30 -22.903667 Mikea Forest 12 43.4755 249 250 1 1 Toliara 2275276455 [462,1047,1917,1943] 2002-02-19 2002-03-01 2002-02-19 MOLC Madagascar 30 -22.903667 Mikea Forest 12 43.4755 249 250 2 2 Toliara 2275276891 2002-07-23 2002-08-06 2002-07-23 CAS, MOLC Madagascar 249 250 2 2 Toliara 2275277096 [506,1025,1953,1979] 2002-01-06 2002-01-16 2002-01-06 CAS Madagascar 249 250 1 1 Toliara 2275277464 2002-01-17 2002-01-28 2002-01-17 CAS Madagascar 249 250 1 1 Toliara 2275277540 2002-07-23 2002-08-06 2002-07-23 CAS, MOLC Mikea Forest, NW Madagascar 37 250 251 -22.913334 Toliara 12 43.482166 249 250 3 3 Toliara 2275276717 2002-03-08 2002-03-18 2002-03-08 CAS Mikea Forest, NW Madagascar 37 -22.913334 Toliara 12 43.482166 250 251 1 1 Toliara 2275276991 2003-05-20 2003-06-03 2003-05-20 CAS Madagascar 80 -24.830833 Parcelle 12 46.536167 Andohahela National Park 250 251 1 1 Toliara 2275276649 [515,1259,261,287] 2003-02-13 2003-02-24 2003-02-13 CAS, MOLC Madagascar 80 -24.830833 Parcelle 12 46.536167 Andohahela National Park 250 251 4 4 Toliara 2275277588 [151,1432,296,323] 2003-12-07 2003-12-17 2003-12-07 CAS Madagascar 80 -24.834833 Ihazofotsy Parcelle 12 46.48683 Andohahela National Park 250 251 1 1 Toliara 2275276889 2003-03-24 2003-04-03 2003-03-24 CAS Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 250 251 1 1 Toliara 2275277081 [624,1240,405,431] 2002-12-27 2003-01-07 2002-12-27 CAS Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 250 251 2 2 Toliara 2275277242 2003-02-25 2003-03-07 2003-02-25 CAS Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 250 251 1 1 Toliara 2275277631 [503,1040,440,467] 2003-01-26 2003-02-05 2003-01-26 CAS Madagascar 85 -25.006666 Berenty Special Reserve 12 46.303333 250 251 2 2 Toliara 2275277140 2001-11-09 2001-11-19 2001-11-09 CAS Madagascar 840 -22.8865 Zombitse National Park 12 44.692165 250 251 1 1 Toliara 2275277463 2006-12-02 2006-12-04 2006-12-02 CAS Madagascar 200 -24.568 Ivohibe Forest 12 47.20233 250 251 1 1 Toliara 2275277447 [807,1240,548,575] Fisher Madagascar 250 251 1 Toliara 2275277385 2006-12-06 2006-12-08 2006-12-06 CAS, MOLC Madagascar 500 -24.237667 Vohidava Forest 12 46.286 250 251 4 4 Toliara 2275276644 Madagascar Fisher 250 251 1 Toliara 2275277273 2008-12-25 2009-01-01 2008-12-25 MOLC Madagascar 45 -24.657833 Androimpano Forest 12 43.956165 250 251 1 1 Toliara 2275277278 2008-11-06 2008-11-13 2008-11-06 CAS Madagascar 45 -24.657833 Androimpano Forest 12 43.956165 250 251 2 2 Toliara 2275276839 2009-05-14 2009-05-21 2009-05-14 CAS, MOLC Madagascar 130 -24.650333 Androimpano Forest 12 43.963165 250 251 8 8 Toliara 2275276971 2008-10-21 2008-10-31 2008-10-21 CAS Madagascar 122 -23.430334 Ankazomena 12 43.834167 250 251 7 7 Toliara 2275277378 2003-10-27 2003-11-03 2003-10-27 CAS Madagascar 20 -23.046383 Ranobe 1 43.610283 250 251 2 2 Toliara 2275277610 2002-05-26 2002-06-05 2002-05-26 CAS Madagascar 9 -23.179667 Hotel Paradisia 12 43.616833 Ifaty 250 251 3 3 Toliara 2275276919 [709,1260,909,935] 2001-10-13 2001-11-12 2001-10-13 CAS Madagascar 9 -23.179667 Hotel Paradisia 12 43.616833 Ifaty 250 251 1 1 Toliara 2275277092 [151,1132,944,971] 2010-11-24 2010-12-12 2010-11-24 CAS Madagascar 500 -21.2185 Makay Mts. 12 45.321 250 251 1 1 Toliara 2275277451 Fisher Madagascar 250 251 1 Toliara 2275277169 2008-10-30 2008-11-09 2008-10-30 CAS Madagascar 15 -22.818 Manombo 12 43.372665 Tsifota 250 251 1 1 Toliara 2275277500 2008-10-20 2008-10-30 2008-10-20 CAS Madagascar 18 -22.818 Andranovato 12 43.502167 250 251 2 2 Toliara 2275277950 2008-10-24 2008-10-29 2008-10-24 CAS Madagascar 120 -24.0485 Mitoho Forest 12 43.752335 Tsimanampetsotsa National Park 250 251 3 3 Toliara 2275277192 [250,788,1161,1187] 2008-12-04 2008-12-12 2008-12-04 CAS Madagascar 120 -24.0485 Mitoho Forest 12 43.752335 Tsimanampetsotsa National Park 250 251 6 6 Toliara 2275277240 2008-10-24 2008-10-29 2008-10-24 CAS Madagascar 250 251 2 2 Toliara 2275276983 [1161,1279,1197,1223] CAS Madagascar 250 251 1 1 Toliara