Aegidium minor Paulian 1984: 72 Revision of the mainland species of the Neotropical genus Aegidium Westwood (Coleoptera: Scarabaeidae: Orphninae) Frolov, Andrey V. Akhmetova, Lilia A. Vaz-de-Mello, Fernando Z. Journal of Natural History 2017 2017-05-17 51 19 - 20 1035 1090 Paulian, 1984 Paul. R. Paulian 1984 [475,840,141,166] Insecta Hybosoridae Aegidium Animalia Coleoptera 46 1080 Arthropoda species minor  ( Figure 15(a–g))      Aegidium minor Paulian 1984: 72   Type locality  Ecuador.   Type material examined   Holotype, maleat MNHNlabeled ‘ Equador Balzen[III]/52559/Fry Coll. 1905.100./ Holotype/  Aegidium minor Paul. R. Pauliandet./HOLOTYPE’.  Additional material examined   ECUADOR. Bolívar: one maleat MNHN, Balzapamba, M. de Mathan,  3 April 1894leg. Cotopaxi:  one maleat MHNG, Chugchilán,  2600 m,  February 1983, G. Onoreleg. Los Ríos:  one maleat CMN, Patricia Pilar,  47 kmS Santo Domingo, Palenque River,  230 m, FIT,  May–July 1985, S. Peckleg.;  two femalesat CMN, Patricia Pilar,  47 kmS Santo Domingo, Palenque River,  250 m, malaise-FIT rainforest,  5 May–25 July 1985, S. Peckand J. Peck leg.;  one maleat CEMT, Provincia de Los Ríos,  16 February 1983, S. Sandovalleg.   Diagnosis   Aegidium minoris similar to Ae. cribratumand Ae. asperatumin the shape of the aedeagus but differs from them by the parameres having small sensory areas apicad of lateral teeth ( Figure 16(b)). From the former species it also differs in having more protruding anterior angles of frontoclypeus, and from the later by sparser punctation of elytra.   Description Male ( Figure 15(a,b,d–f)). Body length 12.0–17.0 mm. Colour uniformly blackish brown. Anterior margin of frontoclypeus slightly convex in middle, slightly sinuate each side of medial convexity, with strongly protruding anterior angles, without distinct border or slightly upturned. Frontoclypeus punctate with rounded punctures separated by about 1–2 puncture diameters. Eyes relatively large: width about 1/7 distance between eyes in dorsal view. Pronotum with widely rounded lateral margins, slightly wider than elytra, 1.6 times wider than length, 0.55 times length of elytra. Posterior angles widely rounded, indistinct. Anterior margin bordered, border interrupted medially and not reaching anterior angles. Base of pronotum without furrow, punctate with a row of rounded punctures. Pronotal disc deeply excavated, with flattened area mediobasally. Lateral pronotal processes long, horn-shaped in lateral view, not protruding past lateral margin outline of pronotum in dorsal view. Medial excavation of pronotum punctate with rounded punctures, lateral processes smooth. Scutellum narrow subtriangular, about 1/11 length of elytra. Elytra 1.2 times longer than wide, with humeral and apical umbones, widest in basal 1/3, and tapering apically in dorsal view. Elytral carinae feebly distinct. Elytra covered with relatively rounded punctures separated by 1–3 puncture diameters, becoming denser at apices. Macropterous. Mesotibiae without tuft of setae ventroapically. Inner apical spur of mesotibia shorter than basal mesotarsomere and not curved downwards. Aedeagus with relatively long (0.55 times length of phallobase), tapering parameres ( Figure 15(d)). Parameres almost as wide as apical part of phallobase in dorsal view ( Figure 16(e)). Ventrobasal transverse plate of parameres distinct. Lateral teeth of parameres feebly protruding past paramere outline in dorsal view ( Figure 15(e)). Lateral sides of parameres with small sensory areas apicad of lateral teeth ( Figure 15(b,d), arrowed). Endophallus without strongly sclerotised sclerites. Cranial part of spiculum gastrale narrow, tapering and rounded apically ( Figure 15(f)). Female ( Figure 15(c)) differs from male in having protibial spur, relatively smaller pronotum without armature and in absence of inner protibial tooth. Body length of examined specimens 13.0 mm.  Variation Pronotum shape in males varies from fully developed as described above to ill-defined lateral pronotal processes and shallow central pronotal excavation.   Distribution The species is known from a few localities in the western Ecuadorian Andes ( Figure 15(g)). 3757961367 MNHN Ecuador Paul. R. Paulian Balzen 46 1080 1 holotype 3757961365 1894-04-03 1985-07-31 1985-05 MNHN Balzapamba, M Ecuador 2600 true Bolivar Patricia Pilar 46 1080 1 1 1 Cotopaxi 3757961424 [273,1156,642,666] 1983-02 1985-07-31 1985-05 MHNG G. Onore Ecuador 2600 true Chugchilan Patricia Pilar 46 1080 1 1 1 Cotopaxi 3757961415 1985-05 1985-07-31 1985-05 CMN Peck Ecuador 230 true Palenque River Patricia Pilar 46 1080 1 1 1 Cotopaxi 3757961313 1985-05-05 1985-07-25 1985-05-05 CMN Balzapamba, M Ecuador 250 true Palenque River Patricia Pilar 46 1080 1 1 1 Cotopaxi 3757961419 [214,1082,781,805] 1983-02-16 1985-07-31 1985-05 CEMT Sandoval Ecuador 2600 true Provincia de Los Rios Patricia Pilar 46 1080 1 1 1 Provincia de Los Rios