Vedalia cardinalis Mulsant 1850: 906 Novius cardinalis : Crotch, 1874: 283 Eurodolia cardinalis : Weise, 1895: 150 Rodolia cardinalis : Weise, 1905: 220 Weise, 1916: 50 Macronovius Korschefsky, 1931: 99 Sasaji, 1971: 238 Gordon, 1972: 25 Gordon, 1985: 666 Ren et al ., 2009: 160 Ślipiński, 2007: 143 Rodolia aegyptiaca Sicard, 1907: 67 Korschefsky, 1931: 99 Macronovius cardinalis : Weise, 1922: 104 Macronovius cardinalis Weise, 1922: 104 Korschefsky, 1931: 98 Revision Of The Australian Coccinellidae (Coleoptera). Genus Novius Mulsant Of Tribe Noviini Pang, Hong Tang, Xue-Fei Booth, Roger G. Vandenberg, Natalia Forrester, Juanita Mchugh, Joseph Ślipiński, Adam Annales Zoologici 2020 2020-03-30 70 1 1 24 (Mulsant) Mulsant 1850 [290,625,919,941] Insecta Coccinellidae Novius GBIF Animalia Coleoptera 5 6 Arthropoda species cardinalis  ( Figs 1–5, 12–18, 46, 47, 72–75, 89, 94, 95, 108–111)      Vedalia cardinalis Mulsant 1850: 906. TL: Australia.     Novius cardinalis: Crotch, 1874: 283.     Eurodolia cardinalis: Weise, 1895: 150.     Rodolia cardinalis: Weise, 1905: 220;  Weise, 1916: 50(  Macronoviusgroup);  Korschefsky, 1931: 99;  Sasaji, 1971: 238;  Gordon, 1972: 25;  Gordon, 1985: 666; Gordon, 1990: 291;  Ren et al., 2009: 160;  Ślipiński, 2007: 143.     Rodolia aegyptiaca Sicard, 1907: 67;  Korschefsky, 1931: 99.     Macronovius cardinalis: Weise, 1922: 104.    Macronovius cardinalisab. obnubilatus  Weise, 1922: 104;  Korschefsky, 1931: 98.    Diagnosis.Typical color forms of  Novius cardinalisare easily distinguished, especially from superficially similar  N. bellusdue to the color pattern with red elytra, each having two transverse to oblique blotches in addition to the black sutural stripe. Rare atypical forms can be distinguished by the stout penis guide of the male genitalia.    Description.Length 2.7–4.2 mm. Body oval, widest near middle, 1.1–1.3 times longer than wide. Color pattern variable. Head uniformly dark, pronotum dark with anterior part reddish. Elytra of typical form red with black medially expanded, sutural stripe and two transverse to oblique irregular spots on each elytron. Melanic forms rare, with black spots on elytra expanding and coalescing to a large degree ( Figs 1, 2, 46, 47, 72–75, 89). Interocular distance in frontal view 2.0–2.3 times eye width. Male genitalia ( Figs 12–17, 108–111): parameres slender, narrow at base, expanded apically; penis guide stout, broad at base, strongly narrowed apically (in lateral view), about as long as or slightly longer than parameres, apically pointed. Penis weakly pointed apically. Female genitalia as in Figs 94, 95.     Typematerial.“Nouvelle Hollande” (collect. Hope) ( OUM).   Other specimens examined.   Australian Capital Territory: CSIRO, Canberra,  15-18.xii.1980, R. D. Pope(10, BMNH);  Uriarra,  6.xii.1980, R. D. Pope(1, BMNH);  Deakin,  22-23.xi.1980, R. D. Pope(1, BMNH);  CSIRO, Canberra, 23.xii.1080, R. D. Pope(2, BMNH);  Honeysuckle Ck. T.S., old quarry (  1100m),  2.iii.1986, C. Reid, on  Euc.& Acacia(1, ANIC);  Kangaroo Creek,  8.iii.1972, S. Misko, R. Kohout(1, ANIC);  35.16S, 149.06E, Black Mt,  600m,  xii.1987, M.E. Irwin, ex Malaise trap, ANIC(1, ANIC);  35.34S 148.47E, Mt. Gingera,  15.xii.1987, A.A. Calder, foliage of  Euc.pauciflora(1, ANIC);  Canberra,  14.vii.1964, G.S. Snoball, Ex Canberraaccession, ex Bermada(14, ANIC);  Black Mt.A.C. T., 1.3.30, A. Tonnoir(2, ANIC).   Tasmania: Hobart, J.J. Waiker(1, BMNH);  Tasmania(2, BMNH);  Launceston(2, ANIC);  Tasmania(1, ANIC).   New South Wales: Sydney, C. Darwin, 87-42 (2, BMNH);  Jervis Bay,  xi.1924, F.A. Rodway(1, BMNH); N. Sydney, G.E. Bryant,  xii.1908(1, BMNH);  Blue Mts., G.E. Bryant.  xii.1908(1, BMNH);  Tuglo Res., 151.’6’E -32.’4’S, 30.xi- 1.xii.80, R. D. Pope(2, BMNH);  J.C. Goudie, Sydney, Lea(1, VM);  Exeter,  i.1935, H.J. Carter(1, ANIC);  Blue Mts, E.W. Ferguson Collection(5, ANIC);  Sydney, E.W. Ferguson Collection(1, ANIC);  2 mi.S. of Myrtle Ck., Braemor Forest, Casino-Grafton Rd., 34.325S 149.52E,  8.xi.1969, S. Misko(1, ANIC);  Kosciuszko, 17-11-38, A.L. Tonnolr(1, ANIC);  Badgery’s Lookout, nr. Tallong,  27.ii.1969, Britton, Upton, Misko(1, ANIC);  Sydney,  22.iii.1927, Nicholson(4, ANIC);  2kmS. of Tallong,  18.ii.1974, S. Misko(1, ANIC); c. 1kmN. of Binalong, 12.xi.73, S. Misko(1, ANIC);  Cattle Creek Rd. SW of Captains FlatNSW,  26.xii.1976, W. Allen(1, ANIC);  105kmSW. of Nowta on Nerriga-Nowta Rd.,  19.i.1971, woodlands, S. Misko& K. Pullen(1, ANIC);  Sydney, W.W. Froggatt(2, ANIC);  N.E. University, Armidale,  1.ii.1967, C.W. Frazier. ANIC Uni. of New England Coll., Donated1983 (3, ANIC);  Wauchope, ( 31.27S 152.44E), 72kmW. on Oxley Highway, 4.1.70, wet sclerophyll, at light, Britton, Holloway, Misko(1, ANIC);  36.125S 148.43E, Dainer Gap, NSW,  6.ii.1974, P. Morrow, ANIC(1, ANIC);  28.29S 152.24E, Tooloom Plateau,  14kmW Urbenville,  14.ii.1984, NSW, I.D. Naumann(1, ANIC);  35.30S 150.18E, KioloaSF  15kmNE Batemans Bay,  ix.1986, M.G. Robinson, flight interc. trap (1, ANIC);  35.30S 150.18E, KioloaSF  15kmNE Batemans Bay,  viii.1986, M.G. Robinson, flight interc. trap (1, ANIC);  36.12S 148.43E, Dainers Gap, NSW.,  6.iii.1974, P. Moorow(2, ANIC);  R. Bogan, J. Amstrong(1, ANIC);  35.30S 150.18E, KioloaSF  15kmNE Batemans Bay, Jan.87, M.G. Robinson, flight interc. trap (1, ANIC);  28.29S 152.24E, Tooloom Plateau,  14kmW Urbenville,  14.ii.1984, I.D. Naumann(1, ANIC);  Sydney,  14.viii. 1964, R. G. Lukins, Ex Canberraaccession, ex Bermada(16, ANIC);  Biocon, Sydney,  ii.1965, V. S. Wright, Exmaterials from L.H.I. on  30.i.1965(9, ANIC);  Mt. Tomah, Blue Mts., 2/32. J. Amstrong(1, ANIC);  The Kurrajong Blue Mts, J. Amstrong(1, ANIC);  The Flags  30mS. of Walcha,  27.i.1963, C.W. Frazier(1, ANIC);  A. Dyce, 20.5.49, Leeton(1, ANIC);  The Kurrajong Blue Mts, J. Amstrong(1, ANIC).   South Australia: C E Cole, Hobart, 4317/3, 4317/4 (2, VM).   Western Australia: Kalamunda, SW Australia,  14.iii-14.iv.1914, R.E. Turn- er, 1914-349 (1, BMNH);  Yanchep, 32mls. N. of Perth,  20-31.xii.1935, R. E. Turner(1, BMNH); nr Busselton,  14.iii.1981, R. D. Pope(1, BMNH); H.J. Carter Coll.P.20 4.22. (1, VM);  Perth, E. T. Smith (2, VM);  Katanning,  11.x.1951, I.F.B. Common(1, ANIC);  Stirling Ra.NP. by S. R. Caravan Pk.,  30.xii.1985, C. Reid, on  Euc.wandoo(1, ANIC);   45kmE Geraldton,  20-23.viii.1987, C. Reid, ex  Acacia(1, ANIC).   Victoria: Diamond Creek, 18& 21.i.81, R. D. Pope(1, BMNH); J.E. Dixon(12, VM);  Melbourne,  16.i.1928, G.F. Hill(1, VM);  Baxter, J.E. Dixon(5, VM); 12 Mi. W.of Neerim,  30.xii.1964, Neboiss(1, VM);  Carrum, V29.12.30, C. Oke(1, VM);  Western District, J.E. Dixon(1, VM);  Bogong Plains,  5699-6000 ft.,  i.1928, F.E. Wilson, (1, VM);  Stoney Creak, 14.ii,1973, LocalityVB (1, VM);  Mooroopna.  19.xii.1937, A.D. Butcher(1, VM);  Melbourne, P.E. Wilson.F.E. (1, VM);  Grampians,  10.vi.1975, S. Neser(1, ANIC).   Queensland: Lamington NP,  ii.1983, M. Lowman(1, ANIC);  Indooroopilly,  xii.1994, Labbred: fed on  Icerya aegyptiaca(3, ANIC).   Northern Territory: 23.37S 133.54E, 10kmN by E of Alice Spring,  6. xi.1979, T. Weir(2, ANIC);  23.38S 133.52E, 7kmN by W of Alice Spring,  8. xi.1979, T. Weir, (1, ANIC).   Norfolk Islands: Norfolk Is. 29.05S 167.59E,  15. i.1992, D.P.A, Sands(1, BMNH).   Distribution.Originally Australian, but due to its introduction as a biocontrol agent, this species is now cosmopolitan ( Fig. 18). 2609449620 1980-12-15 1980-12-18 1980-12-15 CSIRO, R, BMNH D. Pope Canberra 5 6 1 Australian Capital Territory 2609449616 [966,1440,427,449] 1980-12-06 R, BMNH D. Pope Uriarra 5 6 1 Australian Capital Territory 2609449633 [816,1344,458,480] 1980-11-22 1980-11-23 1980-11-22 R, BMNH D. Pope Deakin 5 6 1 Australian Capital Territory 2609449666 CSIRO, R, BMNH Canberra & D. Pope 5 6 1 Australian Capital Territory 2609449628 1986-03-02 T, ANIC Honeysuckle Ck. & C. Reid & Euc. 1100 5 6 1 Australian Capital Territory 2609449630 1972-03-08 R, ANIC S. Misko & Kohout Kangaroo Creek 5 6 1 Australian Capital Territory 2609449679 1987-12 ANIC Irwin 600 -35.16 Black Mt 717 149.06 5 6 1 Australian Capital Territory 2609449607 1987-12-15 ANIC Calder & Euc. -35.34 Mt. Gingera 716 148.47 5 6 1 Australian Capital Territory 2609449694 1964-07-14 ANIC G. S. Snoball & Ex Canberra & Bermada Canberra 5 6 1 Australian Capital Territory 2609449705 T, ANIC, BMNH A. Tonnoir & Hobart, J. J & Waiker Black Mt. 5 6 1 Tasmania 2609449629 T, ANIC, BMNH A. Tonnoir & Hobart, J. J & Waiker Black Mt. 5 6 1 Tasmania 2609449624 [1085,1329,798,820] BMNH Tasmania 5 6 1 Tasmania 2609449636 ANIC Launceston 5 6 1 Tasmania 2609449701 [979,1214,829,851] ANIC, BMNH Tasmania & Sydney, C & Darwin New South Wales 5 6 1 New South Wales 2609449650 ANIC, BMNH Tasmania & Sydney, C & Darwin New South Wales 5 6 1 New South Wales 2609449646 1924-11 BMNH Rodway Jervis Bay 5 6 1 New South Wales 2609449692 1908-12 BMNH Bryant Sydney 5 6 1 New South Wales 2609449623 1908-12 BMNH Blue Mts., G. E. Bryant. 5 6 1 New South Wales 2609449658 R, BMNH Tuglo Res. & D. Pope 5 6 1 New South Wales 2609449621 VM J. C. Goudie & Sydney & Lea 5 6 1 New South Wales 2609449673 [913,1326,1014,1036] 1935-01 ANIC Carter Exeter 5 6 1 New South Wales 2609449654 ANIC Blue Mts, E. W. & Ferguson Collection 5 6 1 New South Wales 2609449711 ANIC Sydney, E. W. & Ferguson Collection 5 6 1 New South Wales 2609449641 1969-11-08 ANIC S. Misko Myrtle Ck. -34.325 Casino-Grafton Rd. 461 149.52 Braemor Forest 5 6 1 New South Wales 2609449645 ANIC Kosciuszko & Tonnolr 5 6 1 New South Wales 2609449667 1969-02-27 ANIC Britton & Upton & Misko Tallong Badgery's Lookout 5 6 1 New South Wales 2609449640 1927-03-22 ANIC Nicholson Sydney 5 6 1 New South Wales 2609449693 1974-02-18 ANIC Misko Tallong 5 6 1 New South Wales 2609449631 ANIC Binalong & Misko 5 6 1 New South Wales 2609449610 1976-12-26 ANIC Allen Cattle Creek Rd. 5 6 1 New South Wales 2609449685 1971-01-19 ANIC Misko & Pullen Nowta on Nerriga-Nowta Rd. 5 6 1 New South Wales 2609449606 ANIC Sydney, W. W & Froggatt 5 6 1 New South Wales 2609449627 1967-02-01 ANIC N. E. University & Frazier. & Coll. & Donated United Kingdom Uni. of New Armidale 5 6 1 2609449617 ANIC Wauchope & Oxley Highway & Britton & Holloway & Misko United Kingdom -31.27 730 152.44 5 6 1 2609449708 1974-02-06 ANIC Morrow United Kingdom -36.125 Dainer Gap 451 148.43 5 6 1 2609449642 1984-02-14 ANIC Naumann United Kingdom -28.29 Tooloom Plateau 740 152.24 5 6 1 2609449695 1986-09 ANIC Robinson United Kingdom -35.3 Kioloa 716 150.18 5 6 1 2609449643 1986-08 ANIC Robinson United Kingdom -35.3 Kioloa 716 150.18 5 6 1 2609449688 1974-03-06 ANIC Moorow United Kingdom -36.12 Dainers Gap 713 148.43 5 6 1 2609449649 [1047,1440,1818,1840] R, ANIC Bogan, J & Amstrong United Kingdom 5 6 1 2609449639 ANIC Kioloa & Robinson United Kingdom -35.3 Jan. 716 150.18 5 6 1 2609449670 1984-02-14 ANIC Naumann United Kingdom -28.29 Tooloom Plateau 740 152.24 6 7 1 2609449614 1964-08-14 R, ANIC G. Lukins & Ex Canberra & Bermada United Kingdom Sydney 6 7 1 2609449687 1965-01-30 1965-02 1965-01-30 V, ANIC S. Wright & J. L. Hemptinne United Kingdom Biocon Ex Sydney 6 7 1 2609449653 ANIC Amstrong United Kingdom Blue Mts. Mt. Tomah 6 7 1 2609449682 ANIC The Kurrajong Blue Mts, J & Amstrong United Kingdom 6 7 1 2609449651 1963-01-27 ANIC Frazier United Kingdom 30 Walcha The Flags 6 7 1 2609449608 ANIC A. Dyce & Leeton United Kingdom 6 7 1 2609449710 [452,1073,180,447] 1914-03-14 1914-04-14 1914-03-14 ANIC, VM, R, BMNH J. Amstrong & C E Cole & Hobart Australia Kalamunda The Kurrajong Blue Mts 6 7 1 South Australia 2609449712 1914-03-14 1914-04-14 1914-03-14 ANIC, VM, R, BMNH J. Amstrong & C E Cole & Hobart Australia Kalamunda The Kurrajong Blue Mts 6 7 1 South Australia 2609449622 1914-03-14 1914-04-14 1914-03-14 ANIC, VM, R, BMNH J. Amstrong & C E Cole & Hobart Australia Kalamunda The Kurrajong Blue Mts 6 7 1 South Australia 2609449662 1935-12-20 1935-12-31 1935-12-20 R, BMNH E. Turner Australia Perth Yanchep 6 7 1 South Australia 2609449715 1981-03-14 R, BMNH D. Pope Australia Busselton 6 7 1 South Australia 2609449664 VM Carter Coll. Australia 6 7 1 South Australia 2609449700 [989,1301,364,386] T, VM Perth Australia 6 7 1 South Australia 2609449618 1951-10-11 ANIC Common Australia Katanning 6 7 1 South Australia 2609449611 1985-12-30 R, ANIC Stirling Ra. & Caravan Pk. & C. Reid & Euc. Australia 7 8 6 7 1 South Australia 2609449615 1987-08-20 1987-08-23 1987-08-20 ANIC, R, BMNH C. Reid & D. Pope Australia Diamond Creek 7 8 1 Victoria 2609449652 1987-08-20 1987-08-23 1987-08-20 ANIC, R, BMNH C. Reid & D. Pope Australia Diamond Creek 7 8 1 Victoria 2609449655 [540,767,241,263] VM Dixon Australia 7 8 1 Victoria 2609449647 [144,612,272,294] 1928-01-16 VM Hill Australia Melbourne 7 8 1 Victoria 2609449672 VM Baxter, J. E & Dixon Australia 7 8 1 Victoria 2609449637 1964-12-30 VM Mi. W. & Neboiss Australia Neerim 7 8 1 Victoria 2609449674 [248,660,333,355] V, VM Carrum & C. Oke Australia 7 8 1 Victoria 2609449605 VM Western District, J. E & Dixon Australia 7 8 1 Victoria 2609449613 1928-01 VM Wilson Australia 1783 Bogong Plains 7 8 1 Victoria 2609449680 VM Stoney Creak & Locality Australia 7 8 1 Victoria 2609449626 1937-12-19 VM Butcher Australia Mooroopna. 7 8 1 Victoria 2609449602 VM Melbourne, P. E. & Wilson. Australia 7 8 1 Victoria 2609449697 [255,772,486,509] 1975-06-10 1983-02 1975-06-10 ANIC Neser & Lamington NP & Lowman Australia Grampians 7 8 1 Queensland 2609449698 1975-06-10 1983-02 1975-06-10 ANIC Neser & Lamington NP & Lowman Australia Grampians 7 8 1 Queensland 2609449690 1979-11-06 1994-12 1979-11-06 ANIC, T Lab & Weir Australia -23.37 Indooroopilly 753 133.54 7 8 1 Northern Territory 2609449706 1979-11-06 1994-12 1979-11-06 ANIC, T Lab & Weir Australia -23.37 Indooroopilly 753 133.54 7 8 1 Northern Territory 2609449604 1979-11-08 1992-01-15 1979-11-08 T, ANIC, BMNH Weir & Norfolk Is. & D. Pope & Sands Australia -29.05 W of Alice Spring 737 167.59 7 8 1 Norfolk Islands 2609449713 1979-11-08 1992-01-15 1979-11-08 T, ANIC, BMNH Weir & Norfolk Is. & D. Pope & Sands Australia -29.05 W of Alice Spring 737 167.59 7 8 1 Norfolk Islands