Acrotaphus franklini Gauld, 1991: 329 Gauld, 1991: 329 Review of the New World genus Acrotaphus Townes, 1960 (Hymenoptera: Ichneumonidae: Pimplinae), with descriptions of fifteen new species Pádua, Diego G. Sääksjärvi, Ilari E. Monteiro, Ricardo F. Oliveira, Marcio L. Zootaxa 2020 2020-01-09 4719 1 1 62 9QRS Gauld, 1991 Gauld 1991 [151,566,1490,1517] Insecta Ichneumonidae Acrotaphus Animalia Hymenoptera 25 26 Arthropoda species franklini  ( Figs 82–90)     Acrotaphus franklini   Gauld, 1991: 329. Holotype ♀, Costa Rica(MNCR) [examined by digital image]   Diagnosis.This species can be distinguished from all other species of  Acrotaphusby the combination of the following characters: 1) margin of gena behind eyes flat in dorsal view; 2) margin of gena 0.6–0.7× length of eye in dorsal view; 3) posterior ocelli separated from eyes by 0.4–0.7× its diameter in dorsal view; 4) mesosoma black, except scutellum orange or mesosoma orange with propleuron, pronotum, mesoscutum and anterior and ventral region of mesopleuron black; 5) metasoma orange, with tergites V+ black; 6) ovipositor robust, 1.3–1.6× as long as hind tibia.   Comments.Additional characters to the original description ( ♀) are as follows: body [12.5] 12.0–17.0 mm; mar- gin of gena behind eyes flat in dorsal view; margin of gena [0.4] 0.6–1.0× length of eye in dorsal view; tarsal claw with basal lobe quadrangular, with claw apex slightly overtaking the posterior margin of lobe. Male.( Figs 85–86). Similar to female in structure and coloration, except margin of gena 0.6–0.7× length of eye in dorsal view; posterior ocelli separated from eyes by 0.6× its diameter in dorsal view; sternite IX ( Fig 88) longer than high, with lateral and anterior margins strongly sclerotised, lateral margin slightly concave and posterior margin concave, with few bristles widely spaced posteriorly. Genital capsule ( Figs 89–90): Paramere truncated apically, slightly narrower than the parameral lamina, with bristles except in posterior margin; volsellar lamina with bristles anteroapically spaced in anterior margin and with a set of four to five bristles in anterolateral region, just below the base of the digitus; cuspis robust with about four to five teeth aligned in the mid part until apex and with three to four few bristles spaced in apical region; digitus with teeth on apex, rounded distally and angular ventrally, with the margins of the apical region narrower than base; aedeagus with few teeth on apex and its apex slightly curved downward in lateral view.   Distribution. Brazil*, Colombia**, Costa Rica, Ecuador** and Peru* ( Fig 91).  Biological notes.One male was collected from the nest of the  Trypoxylon( Trypargilum) lactitarseSaussure, 1867 (Crabronidae). This record is similar to the strange case documented by Palacio (1999), who found a species of the  Acrotaphusfrom the nest of the  Trigonopsis violascens(Dalla Torre, 1897) (Sphecidae). Palacio (1991) suggested this was an accidental case, where the solitary wasp captures and imprisons an already parasitized spider (with egg/larva of the  Acrotaphus). This is the first record of the genus  Acrotaphusin nest of Crabronidae.   Material examined.   Holotype: ♀, COSTA RICA, Gste[= Guanacaste] Pv.[= Provincia], Guanacaste NP[=Na- tional Park],   560 m., Est.[= Estación] Maritza Vn[= Volcán] Orosi,  iii.1989( Gauld& Mitchell), INBIO# CRI001  102074, MNCR[digital image sent by Ronald Zuñiga].   Paratype: ♀, COSTA RICA, idem holotype, but   800 m ., iv–v.1986 ( Gauld), #010823022, BNHM [examined].  Brazil: Acre, Senador Guiomard, Reserva Catuaba, 10°04’S/ 67°36’W,  07.xi.2001, Nest trapn° 427 ( E.F. Moratoleg.), ♂[with genitalia extracted], UFAC;  Amazonas, Manaus, WWF, Reserve1113, Rede Lateral Oeste,  07.xi.1985, Malaise( Bert Klein), ♂[with genitalia extracted], INPA; idem, but Reserve1208, Fazendo Esteio, PDBFF, 02°22’34’’S/ 59°52’39’’W, 03.ix.1985,  ♀, ZMUT; idem, but Reserve1301, 02°23’03’’S/ 59°51’15’’W,  22.v.1985,  ♀, ZMUT; idem, but Reserva Ducke,  14.x.1995, Foggingem  Corythophora alta(without collector), ♂[with genitalia extracted], INPA; idem, but Reserve1501, Km41,  15–25.xi.1995, Suspense trap4,  ♀, INPA.   Colombia: Amazonas, PNN [=Parque Nacional Naturale] Amacayacu, Matamata, 03°41’S/ 70°15’W,   150 m ., Red1 [sic],  11.xi.2000( A. Parenteleg.), M.3553 [?],  ♀, #52277, IAVH; idem, but Caño Matá-Matá, Malaise, ii–iii.1989 ( M. Kelsey),  ♀, #53000, IAVH; idem, but Mun.[=Municipal- ity] Leticia, Km11, Via Tarapaca, 04°07’S/ 69°57’W,   200 m., 06–21.ii.2003(W. Morales leg.), M.3782 [?],  ♀, #52999, IAVH.   Ecuador: Dept.[=Department] Orellana, Onkonegare, 00°39’25,7’’S/ 76°27’10,8’’W, a.s.l.:   216 m .,  02.vii.1995, Fogging, Lot#1067 ( T. L. Erwinleg.),  ♀, ZMUT; idem, but  08.ii.1996, Lot#1464,  ♀, ZMUT; idem, but  01.x.1996, Lot#1692,  ♀, ZMUT; idem, but  08.vii.2006, Lot#3245,  ♀, ZMUT; idem, but; idem, but  26.vii.1996, Lot#1592,  ♂, ZMUT; idem, but  08.ii.1996 Lot#1478,  2♂, ZMUT; idem, but  20.x.2005, Lot#3043,  ♂, ZMUT; idem, but  23.i.2006, Lot#3180,  ♂, ZMUT; idem, but  07.vii.2006, Lot#3201,  ♂, ZMUT; idem, but  02.vii.1995, Lot#1090,  ♂, ZMUT; idem, but  10.ii.1995, Lot#1009,  ♂, ZMUT; idem, but  07.ii.1996, Lot#1449,  ♀, ZMUT; idem, but  04.ii.1996, Lot#1417,  ♂, ZMUT; idem, but  06.x.1994, Lot#889,  ♂, ZMUT; idem, but  03.x.1996, Lot#1725,  ♀, ZMUT; idem, but  04.x.1996, Lot#1758,  ♂, ZMUT; idem, but  01.vii.1995, Lot#1148,  ♀, ZMUT; idem, but  02.x.1996, Lot#1710,  ♀, ZMUT; idem, but  08.x.1995, Lot#1266,  ♂, ZMUT; idem, but  21.vi.1994, Lot#694,  ♂, ZMUT; idem, but Yasuni, 00°37’55’’S/ 76°08’39’’W; a.s.l.:  220– 250 m ., Lot#1900,  ♂, ZMUT; idem, but  09.ii.1999, Lot#2013,  ♂, ZMUT;  Morona-Santiago, Cord.[=Cordillera] de Cutucu, 6 Km E.of Macas,  1100 m.,  17.vii.1981( M. Cooper),  ♀, BNHM.   Peru: Dept.[=Department] Huanuco, Tingo Maria, Cueva de Las Pavas,  23–27.vii.1982( C. Porter& T. O’Neill),  2 ♀, FSCA;  Dept.[=Department] Madre de Dios, Los Amigos, 380955,769E/8610042,474N, a.s.l.:  240.2 m.,  14–21.viii.2008, Malaise trap( I. Gómezleg.),  ♀, ZMUT; idem, but  03–10.vii.2008,  ♀, ZMUT; idem, but Explorer’sinn Amazonlogde,   161 m., 12°50’30’’S/ 69°17’31’’W; 29.viii.2009(L. Sulca leg.),  ♀, ZMUT; CU [= Cusco], Shivankorenica., 11°42’S/ 72°54’W, ca.   300 m., 27.x.1997, Malaise trap (S. Cordova leg.),  ♀, ZMUT;  Dept.[=Department] of Loreto, Iquitos area, Allpahuayo, clay,  20.x–05.xi.1998, Malaise trap( I.E. Sääksjärvi& R. Jussila et al.leg.), APHI, C2/5,  ♀, ZMUT; idem, but  04–20.x.1998(I.E. Sääksjärvi& R. Jussila et al.leg.), APHI, C2/4,  ♀, ZMUT; idem, but  17.xi–03.xii.1998, APHI, C2/7,  ♀, ZMUT; idem, but  17.xii–20.i.1998, APHI, C2/9,  ♀, ZMUT; idem, but white sand,  29.i–20.ii.2000( Sääksjärvi et al.leg.), APHI, I3/1,  ♀, ZMUT; idem, but  22.v–11.vi.2000, APHI, E3/7,  ♀, ZMUT; idem, but  08–24.iii.2000, APHI, G2/3,  3♀, ZMUT;   24.iii–16.iv.2000, APHI, E3/4,  ♀, ZMUT; idem, but  24.i–20.ii.2000, APHI, G1/1,  ♀, ZMUT; idem, but  29.vi–16.vii.2000, APHI, E3/9,  ♀, ZMUT; idem, but clay,  22.v–11.vi.2000, APHI, H1/7,  ♀, ZMUT; idem, but  29.vi–16.vii.2000, APHI, J1/9,  ♀, ZMUT; idem, but 30°58’29’’S/ 73°25’46’’W,  07–13.xi.2011( Gómez& Sääksjärvileg.),  ♀, ZMUT; idem, but  14–20.xi.2011,  ♀, ZMUT; idem, but  01–16.viii.2000,  ♀, ZMUT; idem, but Mishana, varillal,  01–16.ix.1998( I.E. Sääksjärvi& R. Jussila et al.leg.), APHI, B3/2,  ♀, MUSM; idem, but clayish soil,  16.ix–01.x.1998, APHI, A2/3,  ♀, MUSM; idem, but white sand,  29.vi.2000, G1(16)  ♀, MUSM; idem, but  08.iii.2000, J2(2)  ♀, MUSM; idem, but  01–15.xii.2000, I1(16),  ♀, MUSM; idem, but  08.xi–01.xii.2000, J1,  ♀, MUSM; idem, but  22.v–11.vii.2000, J1(7),  2♀, MUSM. 2545463078 [367,547,1588,1613] MNCR Costa Rica Costa Rica 25 26 1 1 holotype 2545462898 1989-03 74-10-20 1989-03 COSTA, RICA, INBIO, MNCR Gste & Guanacaste & Pv. & Provincia & Guanacaste NP & Park & Est. & Estacion & Maritza Vn & Volcan & Orosi & Gauld & Mitchell & Ronald Zuniga 560 26 27 CRI001 561 1 holotype 2545462868 [812,1149,689,716] COSTA, RICA 26 27 1 1 paratype 2545462874 COSTA, RICA 800 26 27 1 holotype 2545462879 [178,673,725,752] BNHM Gauld 26 27 1 paratype 2545462895 2001-11-07 E. F. Morato Brazil -10.066667 Reserva Catuaba 1298 -67.6 Senador Guiomard 26 27 1 Acre paratype 2545462793 1985-11-07 Malaise & Bert Klein Brazil Amazonas Reserve Manaus 26 27 1 Acre paratype 2545462893 [230,790,832,859] 1985-11-07 PDBFF Fazendo Esteio Brazil Amazonas Reserve Manaus 26 27 1 Acre paratype 2545462802 [1243,1361,833,859] ZMUT Brazil 26 27 1 1 Acre paratype 2545462798 1985-05-22 ZMUT Brazil -2.3841665 Reserve 21 -59.854164 26 27 1 Acre paratype 2545462924 [790,908,868,894] ZMUT Brazil 26 27 1 1 Acre paratype 2545462838 1995-10-14 ZMUT Fogging Brazil Reserva Ducke 26 27 1 Acre paratype 2545462804 1995-11-15 1995-11-25 1995-11-15 ZMUT Fogging Brazil Km Reserve 26 27 1 Acre paratype 2545462878 [520,623,940,966] INPA Brazil 26 27 1 1 Acre paratype 2545462897 Colombia Amazonas 150 -3.6833334 Matamata 1307 -70.25 Amacayacu 26 27 1 paratype 2545462906 [576,1207,974,1002] 2000-11-11 A. Parente Colombia Red 26 27 1 paratype 2545462849 [1223,1432,974,1002] IAVH Colombia 26 27 1 1 paratype 2545462939 [151,857,1011,1038] IAVH M. Kelsey Colombia Malaise Cano Mata-Mata 26 27 1 paratype 2545463071 [872,1082,1010,1037] IAVH Colombia 26 27 1 1 paratype 2545462884 IAVH Colombia Mun. 200 -4.116667 Km 1307 -69.95 Leticia 26 27 1 paratype 2545462782 IAVH Colombia 26 27 1 1 paratype 2545463011 [327,1427,1081,1109] Ecuador 216 -0.6571389 Onkonegare 1 -76.452995 Dept. 26 27 1 Orellana paratype 2545462796 1995-07-02 T Fogging & Lot & L. Erwin Ecuador 26 27 1 Orellana paratype 2545462869 [717,833,1119,1145] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545463094 [842,1197,1118,1145] 1996-02-08 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545463065 [1206,1322,1119,1145] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462864 1996-10-01 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545463025 [410,528,1154,1180] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462876 [539,916,1153,1180] 2006-07-08 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462819 [928,1045,1154,1180] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462840 1996-07-26 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545463016 [285,404,1190,1216] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462894 [417,776,1189,1216] 1996-02-08 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462886 [790,923,1190,1216] ZMUT Ecuador 26 27 2 2 Orellana paratype 2545462827 [935,1301,1189,1216] 2005-10-20 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462792 [1314,1433,1190,1216] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462956 [151,500,1225,1252] 2006-01-23 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462885 [511,627,1226,1252] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462931 [637,1007,1225,1252] 2006-07-07 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462829 [1018,1135,1226,1252] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462815 1995-07-02 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462896 [236,354,1261,1287] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545463073 [365,727,1260,1287] 1995-02-10 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462850 [740,858,1261,1287] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462912 [869,1231,1261,1287] 1996-02-07 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462806 [1244,1362,1261,1287] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462828 1996-02-04 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462911 [455,572,1297,1323] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462926 [584,930,1297,1323] 1994-10-06 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545463044 [942,1060,1297,1323] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462812 [1071,1432,1296,1323] 1996-10-03 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462927 [151,271,1333,1359] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462909 [284,652,1332,1359] 1996-10-04 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545463036 [666,785,1333,1359] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545463093 [799,1182,1332,1359] 1995-07-01 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462799 [1195,1315,1333,1359] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462794 1996-10-02 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462930 [410,528,1369,1395] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545463075 [539,899,1367,1395] 1995-10-08 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462834 [912,1030,1369,1395] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462811 [1041,1396,1368,1395] 1994-06-21 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462836 ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462918 [242,940,1403,1430] ZMUT Ecuador 235 -0.6319445 Yasuni 21 -76.144165 26 27 1 Orellana paratype 2545462843 [940,1072,1404,1430] Ecuador Lot 26 27 1 Orellana paratype 2545462814 [1081,1197,1404,1430] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545462805 1999-02-09 ZMUT Lot Ecuador 26 27 1 Orellana paratype 2545462941 [283,401,1440,1466] ZMUT Ecuador 26 27 1 1 Orellana paratype 2545463072 1981-07-17 de Cutucu & Km E. & M. Cooper Ecuador 1100 Macas Cord. 26 27 1 Morona-Santiago paratype 2545463057 [306,430,1476,1502] BNHM Ecuador 26 27 1 1 Morona-Santiago paratype 2545462848 1982-07-23 1982-07-27 1982-07-23 T C. Porter & O'Neill Peru Dept. Cueva de Las Pavas Tingo Maria 26 27 1 Huanuco paratype 2545463074 [445,580,1511,1538] FSCA Peru 26 27 2 2 Huanuco paratype 2545462851 2008-08-14 2008-08-21 2008-08-14 I. Gomez Peru 240 Los Amigos Dept. 26 27 1 Madre de Dios paratype 2545463062 [968,1089,1547,1573] ZMUT Peru 26 27 1 1 Madre de Dios paratype 2545462820 [1102,1394,1547,1573] 2008-07-03 2008-07-10 2008-07-03 ZMUT Peru 26 27 1 Madre de Dios paratype 2545463046 ZMUT Peru 26 27 1 1 Madre de Dios paratype 2545463070 [246,794,1582,1609] ZMUT Peru 161 Amazon Explorer's 26 27 1 Madre de Dios paratype 2545463045 [151,271,1619,1645] ZMUT Peru 26 27 1 1 Madre de Dios paratype 2545462873 [284,999,1617,1645] Peru 300 -11.7 Shivankoreni 1295 -72.9 26 27 1 Cusco paratype 2545463029 [215,332,1655,1681] ZMUT Peru 26 27 1 1 Cusco paratype 2545462856 1998-10-20 1998-11-05 1998-10-20 R, APHI I. E. Saaksjarvi & Jussila Peru Dept. Allpahuayo Iquitos area 26 27 C2 1 Loreto paratype 2545463056 [691,808,1690,1716] ZMUT Peru 26 27 1 1 Loreto paratype 2545462832 1998-10-04 1998-10-20 1998-10-04 R, APHI Saaksjarvi & Jussila Peru 26 27 C2 1 Loreto paratype 2545462970 [400,517,1726,1752] ZMUT Peru 26 27 1 1 Loreto paratype 2545463013 [528,967,1725,1752] 1998-11-17 1998-12-03 1998-11-17 APHI Peru 26 27 C2 1 Loreto paratype 2545462852 [1002,1120,1726,1752] ZMUT Peru 26 27 1 1 Loreto paratype 2545463060 1998-01-20 1998-12-17 1998-01-20 APHI Peru 26 27 C2 1 Loreto paratype 2545463063 [299,415,1762,1788] ZMUT Peru 26 27 1 1 Loreto paratype 2545462871 [426,1176,1761,1788] 2000-01-29 2000-02-20 2000-01-29 APHI Saaksjarvi Peru 26 27 1 Loreto paratype 2545462818 [1246,1363,1762,1788] ZMUT Peru 26 27 1 1 Loreto paratype 2545462797 2000-05-22 2000-06-11 2000-05-22 APHI Peru 26 27 E3 1 Loreto paratype 2545462908 [534,650,1797,1823] ZMUT Peru 26 27 1 1 Loreto paratype 2545463014 [661,1064,1797,1823] 2000-03-08 2000-03-24 2000-03-08 APHI Peru 26 27 G2 1 Loreto paratype 2545462823 [1098,1229,1797,1823] ZMUT Peru 26 27 3 3 Loreto paratype 2545462801 2000-03-24 2000-04-16 2000-03-24 APHI Peru 26 27 E3 1 Loreto paratype 2545463015 [304,424,1833,1859] ZMUT Peru 26 27 1 1 Loreto paratype 2545462938 [437,861,1833,1859] 2000-01-24 2000-02-20 2000-01-24 APHI Peru 26 27 G1 1 Loreto paratype 2545462881 [897,1017,1833,1859] ZMUT Peru 26 27 1 1 Loreto paratype 2545462817 2000-06-29 2000-07-16 2000-06-29 APHI Peru 26 27 E3 1 Loreto paratype 2545462875 [217,334,1869,1895] ZMUT Peru 26 27 1 1 Loreto paratype 2545462968 [344,820,1869,1895] 2000-05-22 2000-06-11 2000-05-22 APHI Peru 26 27 H1 1 Loreto paratype 2545462915 [856,972,1869,1895] ZMUT Peru 26 27 1 1 Loreto paratype 2545462842 [982,1405,1869,1895] 2000-06-29 2000-07-16 2000-06-29 APHI Peru 26 27 J1 1 Loreto paratype 2545462907 [151,270,1905,1931] ZMUT Peru 26 27 1 1 Loreto paratype 2545462936 [283,1183,1904,1932] 2011-11-07 2011-11-13 2011-11-07 ZMUT Gomez & Saaksjarvi Peru -30.974722 20 -73.42944 26 27 1 Loreto paratype 2545462807 [1197,1316,1905,1931] ZMUT Peru 26 27 1 1 Loreto paratype 2545462784 2011-11-14 2011-11-20 2011-11-14 ZMUT Peru 26 27 1 Loreto paratype 2545462903 [327,445,1941,1967] ZMUT Peru 26 27 1 1 Loreto paratype 2545462889 [457,753,1941,1967] 2000-08-01 2000-08-16 2000-08-01 ZMUT Peru 26 27 1 Loreto paratype 2545462932 [765,883,1941,1967] ZMUT Peru 26 27 1 1 Loreto paratype 2545462925 1998-09-01 1998-09-16 1998-09-01 R, APHI I. E. Saaksjarvi & Jussila Peru Mishana 26 27 B3 1 Loreto paratype 2545462855 [702,827,1977,2003] MUSM Peru 26 27 1 1 Loreto paratype 2545462890 [840,1409,1977,2003] 1998-09-16 1998-10-01 1998-09-16 APHI Peru 26 27 A2 1 Loreto paratype 2545462854 [151,275,2013,2039] MUSM Peru 26 27 1 1 Loreto paratype 2545462847 [286,745,2013,2040] 2000-06-29 MUSM Peru 26 27 1 Loreto paratype 2545463012 [752,875,2013,2039] MUSM Peru 26 27 1 1 Loreto paratype 2545462914 [886,1190,2013,2039] 2000-03-08 MUSM Peru 26 27 1 Loreto paratype 2545463042 [1197,1320,2013,2039] MUSM Peru 26 27 1 1 Loreto paratype 2545462787 2000-12-01 2000-12-15 2000-12-01 MUSM Peru 27 28 26 27 1 Loreto paratype 2545463030 [419,543,153,179] MUSM Peru 27 28 1 1 Loreto paratype 2545463003 [555,907,153,179] 2000-11-08 2000-12-01 2000-11-08 MUSM Peru 27 28 1 Loreto paratype 2545462887 [919,1043,153,179] MUSM Peru 27 28 1 1 Loreto paratype 2545463061 [1055,1431,153,179] 2000-05-22 2000-07-11 2000-05-22 MUSM Peru 27 28 1 Loreto paratype 2545463001 [151,289,189,215] MUSM Peru 27 28 2 2 Loreto paratype