Atheta ( Lamiota ) keeni Casey, 1910a: 17 Atheta ( s. str. ) innocens Casey, 1910a: 39 Atheta ( Lamiota ) achromata Casey, 1911a: 82 Atheta ( Lamiota ) profecta Casey, 1911a: 83 Atheta ( Liogluta ) keeni : Fenyes, 1920: 209 Atheta ( s. str. ) innocens : Fenyes, 1920: 214 Atheta ( Liogluta ) achromata : Fenyes, 1920: 209 Atheta ( Liogluta ) profecta : Fenyes, 1920: 209 Atheta ( Liogluta ) keeni : Bernhauer & Scheerpeltz, 1926: 657 Atheta ( s. str. ) innocens : Bernhauer & Scheerpeltz, 1926: 644 Atheta ( Liogluta ) achromata : Bernhauer & Scheerpeltz, 1926: 656 Atheta ( Liogluta ) profecta : Bernhauer & Scheerpeltz, 1926: 658 Atheta ( Liogluta ) keeni : Moore & Legner, 1975: 364 Atheta ( s. str. ) innocens : Moore & Legner, 1975: 363 Atheta ( Liogluta ) achromata : Moore & Legner, 1975: 353 Atheta ( Liogluta ) profecta : Moore & Legner, 1975: 370 Lamiota keeni : Seevers, 1978: 263 Xenota innocens : Seevers, 1978: 270 Lamiota achromata : Seevers, 1978: 263 Lamiota profecta : Seevers, 1978: 263 Atheta ( Liogluta ) vasta : Klimaszewski & Winchester, 2002: 39 Lohse & Smetana, 1985: 297 Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes Gusarov, Vladimir I. Zootaxa 2003 2003-11-17 353 1 1 134 5VWRS Casey, 1910 a Casey 1910 [264,584,804,830] Insecta Staphylinidae Atheta Animalia Coleoptera 36 37 Arthropoda species keeni  ( Fig. 97in Klimaszewski & Winchester 2002; Figs. 41–46in this paper)      Atheta( Lamiota) keeni Casey, 1910a: 17.     Atheta( s. str.) innocens Casey, 1910a: 39,  syn. nov.     Atheta( Lamiota) achromata Casey, 1911a: 82,  syn. nov.     Atheta( Lamiota) profecta Casey, 1911a: 83,  syn. nov.     Atheta( Liogluta) keeni: Fenyes, 1920: 209(as valid species).     Atheta( s. str.) innocens: Fenyes, 1920: 214(as valid species).     Atheta( Liogluta) achromata: Fenyes, 1920: 209(as valid species).     Atheta( Liogluta) profecta: Fenyes, 1920: 209(as valid species).     Atheta( Liogluta) keeni: Bernhauer & Scheerpeltz, 1926: 657(as valid species).     Atheta( s. str.) innocens: Bernhauer & Scheerpeltz, 1926: 644(as valid species).     Atheta( Liogluta) achromata: Bernhauer & Scheerpeltz, 1926: 656(as valid species).     Atheta( Liogluta) profecta: Bernhauer & Scheerpeltz, 1926: 658(as valid species).     Atheta( Liogluta) keeni: Moore & Legner, 1975: 364(as valid species).     Atheta( s. str.) innocens: Moore & Legner, 1975: 363(as valid species).     Atheta( Liogluta) achromata: Moore & Legner, 1975: 353(as valid species).     Atheta( Liogluta) profecta: Moore & Legner, 1975: 370(as valid species).     Lamiota keeni: Seevers, 1978: 263(as valid species).     Xenota innocens: Seevers, 1978: 270(as valid species).     Lamiota achromata: Seevers, 1978: 263(as valid species).     Lamiota profecta: Seevers, 1978: 263(as valid species).     Atheta( Liogluta) vasta: Klimaszewski & Winchester, 2002: 39( nec  Lohse & Smetana, 1985: 297) (as valid species).   Type material.  Lectotypeof  Atheta keeni(here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “  Lamiota keeniCsy.”, “TYPE USNM 39480” (red label), “CASEY bequest 1925” ( NMNH).  Paralectotypes: 5♂♂, 5♀♀, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “  keeni PARATYPEUSNM 39480” (red label), “CASEY bequest 1925” ( NMNH).   Lectotypeof  Atheta innocens(here designated): ♀, “ Lane Co., Or.[egon]”, “  Atheta innocensCsy.”, “TYPE USNM 39270” (red label), “CASEY bequest 1925” ( NMNH).   Lectotypeof  Atheta achromata(here designated): ♂(without head and prothorax), “Metlakatla, B.[ritish] Col.[umbia] Keen”, “  achromataCsy.”, “TYPE USNM 39481” (red label), “CASEY bequest 1925” ( NMNH).   Lectotypeof  Atheta profecta(here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “  profectaCsy.”, “TYPE USNM 39482” (red label), “CASEY bequest 1925” ( NMNH).   Paralectotype(see discussion) of  Homalota vastaMäklin in Mannerheim, 1853: ♀, “Sitka”, “Holmberg”, “  Homalota vastam.Sitkha”, “Typus” (green label), “Mus. Zool. H:fors. Spec. Typ. No. 2250.  Homalota vastaMäklin”, “  Homalota vastaMäkl. Holotypus/  Liogluta vasta. HolotypusLohse fix. 1985” ( MZHF).  Additional material.   UNITED STATES: Alaska: ♀, Valdez, 61º09.05'N 146º21.80'W,  20 m, in forest litter,  Alnus, ferns,  Verathrum,  12.vii.1998( V.I.Gusarov) ( SPSU).   Diagnosis.See Klimaszewski and Winchester (2002; as  Atheta vasta).   Discussion.The lectotypeof  At. innocensagrees with the types of  At. keeniin external characters and the shape of the spermatheca. The lectotypeof  At. profectahas no head and prothorax but agrees with the types of  At. keeniin the shape of the aedeagus and abdominal tergum 8. The lectotypeof  At. profectais poorly pigmented, but otherwise it agrees completely with  At. keeniin external characters, including male secondary features, and in the shape of the aedeagus. The type series of  Homalota vastaMäklin in Mannerheim, 1853(examined) includes two specimens( Lohse & Smetana 1985). One of them is a female conspecific with  Atheta keeniand the other is a female belonging to the genus  Lioglutaand illustrated by Lohse and Smetana (1985). This second female bears the following labels: “Sitka”, “Holmberg”, “Mus. Zool. H:fors. Spec. Typ. No. 2251.  Homalota vastaMäklin”. From the description and illustrations in Lohse and Smetana (1985)it is clear that they meant to designate the second specimen (a female of  Liogluta) as the lectotypeof  Homalota vasta. Unfortunately, when they cited the labels of the lectotype, they listed the labels of the first specimen (a female of  Atheta keeni). Moreover, it is the first specimen that had been labeled as the lectotype. The latter circumstance is especially misleading and originally I was going to accept what the lectotypelabel said, and consider  Atheta keeniand  Homalota vastato be synonymous. While identifying some athetines from British Columbiafor Jan Klimaszewski ( Klimaszewski & Winchester 2002) I labeled the specimens of  Atheta keenias  At. vasta. However, Lohse and Smetana are very explicit in that they meant the specimen of  Liogluta, illustrated and redescribed by them, to be the lectotypeof  Homalota vasta.Therefore I accept their intent to designate the  Lioglutaspecimen as the lectotype, disregard their misplaced lectotypelabel and consider  At. keenito be a valid species, different from  Liogluta vasta. In the type series of  Homalota vastaI labeled the specimen of  Lioglutaas lectotypeand the specimen of  At. keenias paralectotype.   Atheta keeniis the typespecies of the subgenus  Lamiota Casey, 1910a(by original designation). Subgeneric assignment of  At. keeniand the status of the name  Lamiotarequires further study.  Atheta keenihas the pronotal pubescence of the typeI and cannot be placed in the genus  Liogluta Thomson, 1858.   Distribution.  Atheta keeniis known from Alaska, British Columbiaand Oregon. 3321919520 NMNH 36 37 1 1 lectotype 3321919606 NMNH 36 37 10 5 5 paralectotype 3321919363 NMNH Lane Co. Lane Co. 36 37 1 1 lectotype 3321919591 NMNH 37 38 1 1 lectotype 3321919572 NMNH 37 38 1 1 lectotype 3321919372 MZHF 37 38 1 1 paralectotype 3321919569 1998-07-12 SPSU Verathrum & V. I. Gusarov United States of America 20 61.150833 Valdez 10 -146.36333 37 38 1 1 Alaska