Atheta ( Lamiota ) keeni Casey, 1910a: 17
Atheta ( s. str. ) innocens Casey, 1910a: 39
Atheta ( Lamiota ) achromata Casey, 1911a: 82
Atheta ( Lamiota ) profecta Casey, 1911a: 83
Atheta ( Liogluta ) keeni : Fenyes, 1920: 209
Atheta ( s. str. ) innocens : Fenyes, 1920: 214
Atheta ( Liogluta ) achromata : Fenyes, 1920: 209
Atheta ( Liogluta ) profecta : Fenyes, 1920: 209
Atheta ( Liogluta ) keeni : Bernhauer & Scheerpeltz, 1926: 657
Atheta ( s. str. ) innocens : Bernhauer & Scheerpeltz, 1926: 644
Atheta ( Liogluta ) achromata : Bernhauer & Scheerpeltz, 1926: 656
Atheta ( Liogluta ) profecta : Bernhauer & Scheerpeltz, 1926: 658
Atheta ( Liogluta ) keeni : Moore & Legner, 1975: 364
Atheta ( s. str. ) innocens : Moore & Legner, 1975: 363
Atheta ( Liogluta ) achromata : Moore & Legner, 1975: 353
Atheta ( Liogluta ) profecta : Moore & Legner, 1975: 370
Lamiota keeni : Seevers, 1978: 263
Xenota innocens : Seevers, 1978: 270
Lamiota achromata : Seevers, 1978: 263
Lamiota profecta : Seevers, 1978: 263
Atheta ( Liogluta ) vasta : Klimaszewski & Winchester, 2002: 39
Lohse & Smetana, 1985: 297
Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes
Gusarov, Vladimir I.
Zootaxa
2003
2003-11-17
353
1
1
134
5VWRS
Casey, 1910 a
Casey
1910
[264,584,804,830]
Insecta
Staphylinidae
Atheta
Animalia
Coleoptera
36
37
Arthropoda
species
keeni
( Fig. 97in Klimaszewski & Winchester 2002; Figs. 41–46in this paper)
Atheta( Lamiota) keeni Casey, 1910a: 17. Atheta( s. str.) innocens Casey, 1910a: 39, syn. nov. Atheta( Lamiota) achromata Casey, 1911a: 82, syn. nov. Atheta( Lamiota) profecta Casey, 1911a: 83, syn. nov. Atheta( Liogluta) keeni: Fenyes, 1920: 209(as valid species). Atheta( s. str.) innocens: Fenyes, 1920: 214(as valid species). Atheta( Liogluta) achromata: Fenyes, 1920: 209(as valid species). Atheta( Liogluta) profecta: Fenyes, 1920: 209(as valid species). Atheta( Liogluta) keeni: Bernhauer & Scheerpeltz, 1926: 657(as valid species). Atheta( s. str.) innocens: Bernhauer & Scheerpeltz, 1926: 644(as valid species). Atheta( Liogluta) achromata: Bernhauer & Scheerpeltz, 1926: 656(as valid species). Atheta( Liogluta) profecta: Bernhauer & Scheerpeltz, 1926: 658(as valid species). Atheta( Liogluta) keeni: Moore & Legner, 1975: 364(as valid species). Atheta( s. str.) innocens: Moore & Legner, 1975: 363(as valid species). Atheta( Liogluta) achromata: Moore & Legner, 1975: 353(as valid species). Atheta( Liogluta) profecta: Moore & Legner, 1975: 370(as valid species). Lamiota keeni: Seevers, 1978: 263(as valid species). Xenota innocens: Seevers, 1978: 270(as valid species). Lamiota achromata: Seevers, 1978: 263(as valid species). Lamiota profecta: Seevers, 1978: 263(as valid species). Atheta( Liogluta) vasta: Klimaszewski & Winchester, 2002: 39( nec Lohse & Smetana, 1985: 297) (as valid species).
Type material. Lectotypeof Atheta keeni(here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ Lamiota keeniCsy.”, “TYPE USNM 39480” (red label), “CASEY bequest 1925” ( NMNH). Paralectotypes: 5♂♂, 5♀♀, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ keeni PARATYPEUSNM 39480” (red label), “CASEY bequest 1925” ( NMNH). Lectotypeof Atheta innocens(here designated): ♀, “ Lane Co., Or.[egon]”, “ Atheta innocensCsy.”, “TYPE USNM 39270” (red label), “CASEY bequest 1925” ( NMNH). Lectotypeof Atheta achromata(here designated): ♂(without head and prothorax), “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ achromataCsy.”, “TYPE USNM 39481” (red label), “CASEY bequest 1925” ( NMNH). Lectotypeof Atheta profecta(here designated): ♂, “Metlakatla, B.[ritish] Col.[umbia] Keen”, “ profectaCsy.”, “TYPE USNM 39482” (red label), “CASEY bequest 1925” ( NMNH). Paralectotype(see discussion) of Homalota vastaMäklin in Mannerheim, 1853: ♀, “Sitka”, “Holmberg”, “ Homalota vastam.Sitkha”, “Typus” (green label), “Mus. Zool. H:fors. Spec. Typ. No. 2250. Homalota vastaMäklin”, “ Homalota vastaMäkl. Holotypus/ Liogluta vasta. HolotypusLohse fix. 1985” ( MZHF). Additional material. UNITED STATES: Alaska: ♀, Valdez, 61º09.05'N 146º21.80'W, 20 m, in forest litter, Alnus, ferns, Verathrum, 12.vii.1998( V.I.Gusarov) ( SPSU).
Diagnosis.See Klimaszewski and Winchester (2002; as Atheta vasta).
Discussion.The lectotypeof At. innocensagrees with the types of At. keeniin external characters and the shape of the spermatheca. The lectotypeof At. profectahas no head and prothorax but agrees with the types of At. keeniin the shape of the aedeagus and abdominal tergum 8. The lectotypeof At. profectais poorly pigmented, but otherwise it agrees completely with At. keeniin external characters, including male secondary features, and in the shape of the aedeagus. The type series of Homalota vastaMäklin in Mannerheim, 1853(examined) includes two specimens( Lohse & Smetana 1985). One of them is a female conspecific with Atheta keeniand the other is a female belonging to the genus Lioglutaand illustrated by Lohse and Smetana (1985). This second female bears the following labels: “Sitka”, “Holmberg”, “Mus. Zool. H:fors. Spec. Typ. No. 2251. Homalota vastaMäklin”. From the description and illustrations in Lohse and Smetana (1985)it is clear that they meant to designate the second specimen (a female of Liogluta) as the lectotypeof Homalota vasta. Unfortunately, when they cited the labels of the lectotype, they listed the labels of the first specimen (a female of Atheta keeni). Moreover, it is the first specimen that had been labeled as the lectotype. The latter circumstance is especially misleading and originally I was going to accept what the lectotypelabel said, and consider Atheta keeniand Homalota vastato be synonymous. While identifying some athetines from British Columbiafor Jan Klimaszewski ( Klimaszewski & Winchester 2002) I labeled the specimens of Atheta keenias At. vasta. However, Lohse and Smetana are very explicit in that they meant the specimen of Liogluta, illustrated and redescribed by them, to be the lectotypeof Homalota vasta.Therefore I accept their intent to designate the Lioglutaspecimen as the lectotype, disregard their misplaced lectotypelabel and consider At. keenito be a valid species, different from Liogluta vasta. In the type series of Homalota vastaI labeled the specimen of Lioglutaas lectotypeand the specimen of At. keenias paralectotype. Atheta keeniis the typespecies of the subgenus Lamiota Casey, 1910a(by original designation). Subgeneric assignment of At. keeniand the status of the name Lamiotarequires further study. Atheta keenihas the pronotal pubescence of the typeI and cannot be placed in the genus Liogluta Thomson, 1858.
Distribution. Atheta keeniis known from Alaska, British Columbiaand Oregon.
3321919520
NMNH
36
37
1
1
lectotype
3321919606
NMNH
36
37
10
5
5
paralectotype
3321919363
NMNH
Lane Co.
Lane Co.
36
37
1
1
lectotype
3321919591
NMNH
37
38
1
1
lectotype
3321919572
NMNH
37
38
1
1
lectotype
3321919372
MZHF
37
38
1
1
paralectotype
3321919569
1998-07-12
SPSU
Verathrum & V. I. Gusarov
United States of America
20
61.150833
Valdez
10
-146.36333
37
38
1
1
Alaska