Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean Hogans, W. E. Zootaxa 2017 4244 1 1 38 Gnanamuthu, 1957 Gnanamuthu 1957 [151,605,404,430] Maxillopoda Pennellidae Pennella Animalia Siphonostomatoida 29 30 Arthropoda species robusta   Synonyms.None.   Type host and locality.  Paraexocoetus brachypteris; Indian Ocean. Morphology.Size: 55 mm. Papillae: partial coverage, spherical, clavate and branching tubiform, random arrangement. Holdfasts: two only, short. First antenna with three segments, second with three segments. Plumes: simple, single or double.   Remarks. Species inquirendae. An intermediate sized species of  Pennelladescribed from a flyingfish,, based on a single specimen. Hogans (1988a)considered  P. robustaas species inquirendaebased on a lack of type material, although the original description and figures do portray the species. The cephalothoracic papillae (tubiform and spherical and configured in an elongate field- Figure 17b), of  P. robustaare different than those found in any other species of  Pennellaand much different from other species also found on flyingfish. Egg strings were present on the specimen originally described. Not reported since the original record.