Gonatodes kendallii Smith 1922:268 Gonatodes affinis Smith 1925:23 1930:16 Gonatodes flavolineatus Nicholls 1949:47 Cnemaspis flavolineatus Manthey & Grossmann 1997:211 Leong & Lim, 2003:9 Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia Grismer, Lee Wood, Perry L. Anuar, Shahrul Riyanto, Awal Ahmad, Norhayati Muin, Mohd A. Sumontha, Montri Grismer, Jesse L. Onn, Chan Kin Quah, Evan S. H. Pauwels, Olivier S. A. Zootaxa 2014 2014-10-31 3880 1 1 147  Leong & Lim, 2003:9  WGL7 (Nicholls, 1949) Nicholls 1949 [151,633,583,610] Reptilia Gekkonidae Cnemaspis Animalia Squamata 79 80 Chordata species flavolineata   Fig. 43      Gonatodes kendallii Smith 1922:268     Gonatodes affinis Smith 1925:23;  1930:16     Gonatodes flavolineatus Nicholls 1949:47     Cnemaspis flavolineatus Manthey & Grossmann 1997:211;  Leong & Lim, 2003:9     Holotype. ZRC 2.6777. Type locality: “the Gap below Fraser’s Hill, on the Pahang-Selangor boundary”, Peninsular Malaysiaat approximately  800 min elevation.   Diagnosis.Maximum SVL 39.2 mm; nine supralabials; nine infralabials; ventral scales keeled; five or six contiguous, pore-bearing precloacal scales with round pores; 23 paravertebral tubercles; body tubercles linearly arranged, absent on flanks; tubercles present in lateral caudal furrows; no ventrolateral caudal tubercles; lateral caudal row present anteriorly; caudal tubercles encircle tail anteriorly; subcaudals keeled, no enlarged median scale row; two postcloacal tubercles on each side of tail base; no enlarged femoral or subtibial scales; subtibials keeled; no enlarged submetatarsal scales; and 23 subdigital fourth toe lamellae; large, black, round spots on nape and anterior portion of body; light vertebral stripe variably present (Tables 6,7).   Redescription of species.  Theredescription of this species is based on a specimen ( LSUHC 8079) collected from the Gapbelow Fraser’s Hill, Pahangand the holotype( ZRC 2.6777). Maximum SVL 38.6–39.2 mm; head oblong in dorsal profile, moderate in size, somewhat narrow, flattened, distinct from neck; snout short, flat in lateral profile; postnasal region constricted medially, flat; scales of rostrum keeled, raised, larger than similarly shaped scales on occiput; weak, supraorbital ridges and frontorostral sulcus; canthus rostralis smoothly rounded; eye large; extra-brillar fringe scales largest anteriorly; pupil round; ear opening oval, taller than wide; rostral slightly concave, divided dorsally by longitudinal groove; rostral bordered posteriorly by supranasals and two smaller scales and laterally by first supralabials; 9 R,L raised supralabials of similar size; 9 R,L infralabials, decreasing in size slightly posteriorly; nostrils elliptical, oriented posterolaterally, bordered posteriorly by small, granular, postnasal scales; mental large, triangular, bordered posteriorly by three postmentals, outer two largest; gular scales raised, keeled; throat scales larger, raised, keeled. Body slender; small, keeled, dorsal scales equal in size throughout body, intermixed with much larger, multicarinate tubercles more or less linearly arranged; tubercles extend from occiput to base of tail; no tubercles on flanks; 23 paravertebral tubercles; pectoral scales raised, keeled, not elongate; abdominal scales slightly larger than dorsals, flat, keeled; five or six precloacal pores; forelimbs moderately long, slender; dorsal scales of brachium raised, keeled; dorsal scales of forearm keeled, raised; ventral scales of brachium keeled, raised, juxtaposed; ventral scales of forearm smooth, raised, juxtaposed; palmar scales smooth, juxtaposed, raised; digits long with an inflected joint; claws recurved; subdigital lamellae unnotched; lamellae beneath first phalanges wide; lamellae beneath phalanx immediately following inflection granular, lamellae of distal phalanges wide; interdigital webbing absent; fingers increase in length from first to fourth with fourth and fifth nearly equal in length; hind limbs slightly longer and thicker than forelimbs; dorsal scales of thigh keeled, raised, juxtaposed; scales of anterior margin of thigh keeled; ventral scales of thigh keeled; subtibial scales keeled, flat, imbricate, with no enlarged anterior row; plantar scales smooth, juxtaposed, raised; no enlarged submetatarsal scales beneath first metatarsal; digits elongate with an inflected joint; claws recurved; subdigital lamellae unnotched; lamellae beneath first phalanges wide; lamellae beneath phalanx immediately following inflection granular, lamellae of distal phalanges wide; interdigital webbing absent; toes increase in length from first to fourth with fourth being slightly longer than fifth; 23 subdigital lamellae on fourth toe; caudal scales arranged in segmented whorls; dorsal caudal scales raised anteriorly, weakly keeled, juxtaposed; middorsal and lateral caudal furrows; no row of enlarged, median subcaudal scales; subcaudal scales keeled; caudal tubercles encircle tail anteriorly; caudal tubercles present in lateral furrow; two enlarged postcloacal tubercles on lateral surface of hemipenal swellings at base of tail.  Color pattern in life based on LSUHC 8079 ( Fig. 43).Dorsal ground color brown to olive-brown; small, dark and light markings on top of head; faint, dark lines radiate out from eyes; elongate, paired, dark, paravertebral markings edging a wide, cream-colored vertebral stripe extending from nape to base of tail; flanks slightly lighter in color; large, white, dorsal tubercles in vertebral region extending from nape onto the tail; limbs mottled with dark and light, irregularly shaped blotches; no black shoulder patches or postscapular band; faint, dark and light brownish bands encircle tail; ventral surfaces of head, body, and limbs dull beige, immaculate; subcaudal region darker.   Distribution.  Cnemaspis flavolineatais known only from the typeof the Gap below Fraser’s Hill, Pahang( Nicholls 1949; Fig. 3).  Natural history. This species is known from only two specimens. No collection data were provided with the holotypebut LSUHC 8079was taken on  26 August 2006at night while sleeping on the underside of a horizontal branch at the edge of a slow moving, marshy stream edged by large granite boulders in hill dipterocarp forest at  900 min elevation ( Fig. 43).   Remarks.Although  Cnemaspis flavolineataand  C. temiah sp. nov.are very similar in morphology and color pattern, the molecular analysis indicates that they are not the same species or each others closest relatives ( Fig. 2).  Cnemaspis flavolineatacan be separated from  C. temiah sp. nov.by having a distinct series of large, white, transversely arranged, vertebral tubercles extending from the nape onto the tail which do not occur in  C. temiah sp. nov.Additionally, the anteriormost caudal tubercles in  C. flavolineataencircle the tail whereas no caudal tubercles encircle the tail in  C. temiah sp. nov.   Material examined.  Malaysia: Pahang; the Gapbelow Fraser’s Hill ZRC 2.6777( holotype) and LSUHC 8079. 3407812435 ZRC Malaysia 800 Peninsular 79 80 ZRC 2.6777 1 holotype 3407812449 SVL, R The Fraser's Hill Gap 79 80 LSUHC 8079, ZRC 2.6777 1 Pahang holotype 3407812370 2006-08-26 LSUHC 900 80 81 LSUHC 8079 1 holotype 3407812420 ZRC, LSUHC Malaysia Fraser's Hill Gap 80 81 ZRC 2.6777, LSUHC 8079 1 Pahang holotype