Hemigymnochaeta laticeps Zumpt, 1953c: 490 The Calliphoridae of Namibia (Diptera: Oestroidea) Kurahashi, Hiromu Kirk-Spriggs, Ashley H. Zootaxa 2006 2006-09-28 1322 1 1 131 4V69 1240433 Corti, 1895: 142 Corti 142 1895 [352,751,411,436] Insecta Calliphoridae Hemigymnochaeta Animalia Diptera 66 67 Arthropoda genus    TYPESPECIES:  Hemigymnochaeta luteaCorti, 1895[=  Ochromyia unicolorBigot, 1888], by monotypy.  NOTES: A large genus restricted to the Afrotropical Region; species of which develop in various kinds of organic matter, such as mushrooms, dung and faeces. Some species drop their eggs in and between the openings of temporary nests of driver ants (Zumpt 1953c: 481). Engel and Cuthbertson (1937: 10, as  Tricyclea resurgens) and Ferrar (1987: 85–86) note females of  H. apiciferaCurran, 1931, collected at fungus material removed by termites ( Isoptera) from their nest in Zimbabwe. The Namibian species appear to be primarily associated with the fungus gardens of termites and the fruiting bodies of associated fungi.    Hemigymnochaeta bequaertiCurran, 1931a: 14.  Figs. 41, 63.    TYPE LOCALITY: Liberia.  DISTRIBUTION: Widespread West Africa to southern Africa: Democratic Republic of Congo, Kenya, Liberia, Namibia*, Nigeriaand South Africa(Natal, Transvaal).  MATERIAL:  4♀, Salambalacampsite,  1–3.iii.2001, Kirk­Spriggs(1) ( HT)faeces;  2♀, same except:  3–4.iii.2001, Kirk­Spriggs(1) ( HT)fish;  2♂, 17♀, same except:  26–28.ii.2001, Marais& Kirk­ Spriggs(1) ( HT)fruit;  2♂, 4♀, same except:  8–10.ii.2004, Kirk­Spriggs(1) ( HT)fruit;  1♂, 5♀, same except: ( MT);  1♂, same except:  22–24.ii.2001, Kirk­Spriggs(1) & Marais( MT);  1♂, same except:  1–4.iii.2001, Kirk­Spriggs(1) ( PT)baited millipede;  2♀, Bum Hillcampsite ( Kwando River),  10–13.ii.2004, Kirk­Spriggs(1) ( HT)fruit;  1♀, same except: ( MT);  1♀, Toggenburg591 campsite,  18–20.ii.2004, Kirk­Spriggs(1) ( HT)fruit;  4♂, 2♀, Popa Fallsrestcamp,  13–16.ii.2004, Kirk­Spriggs(1) ( MT), riverine forest;  1♀, same except: ( HT)fruit.  NOTES: Biology and immature stages unknown. Reared from mushrooms collected near Agatha, South Africaby Zumpt (1953c: 488). In Namibiacollected in hanging traps baited with fermenting fruit bait, rotting fish and faeces, in Malaise traps, and in a pitfall trap baited with a dead millipede. North­eastern Namibia; in the ‘arid’ and ‘mesic’ savanna biome ( Fig. 63). Most abundant in February ( vide Table 2).    Hemigymnochaeta laticepsZumpt, 1953c: 490.  Figs. 42, 64.    TYPE LOCALITY: Rhodesia [= Zimbabwe].  DISTRIBUTION: Botswana, Namibia*, South Africa(Transvaal), Uganda, Upper Voltaand Zimbabwe.  MATERIAL: 3♀, Salambala campsite, 8–10.ii.2004, Kirk­Spriggs(1) ( HT) fruit; 2♀, same except: ( MT).  NOTES: Biology, life history and immature stages unknown. In Namibiacollected in a Malaise trap and a hanging trap baited with fermenting fruit. North­eastern Namibia; in the ‘mesic’ savanna biome ( Fig. 64). Recorded in February ( vide Table 2).    Hemigymnochaeta maraisi  sp. nov.  Figs. 15–20, 39, 65.    TYPE LOCALITY: Namibia.  DISTRIBUTION: Katima Mulilo and Tsumkwe Districts of north­eastern Namibia, in the ‘mesic’ savanna biome ( Fig. 65).  MATERIAL: videdescription, p. 28.   Hemigymnochaeta trichaeta  sp. nov.  Figs. 21–26, 40, 66.   TYPE LOCALITY: Namibia.  DISTRIBUTION: Bwabwata National Park of north­eastern Namibia, in the ‘mesic’ savanna biome ( Fig. 66).  MATERIAL: videdescription, p. 32.   Hemigymnochaeta unicolor(Bigot, 1888: 608) ( Ochromyia).  Figs. 27–32, 67.   TYPE LOCALITY: Sierra Leone.  DISTRIBUTION: Widespread in Afrotropical Region: Benin, Botswana, Democratic Republic of Congo, Ethiopia, Gambia, Ghana, Kenya, Liberia, Madagascar, Malawi, Mozambique, Namibia, Nigeria, Sierra Leone, Somalia, South Africa( Cape, Natal, Transvaal), Togo, Uganda, Zambia* and Zimbabwe.  PREVIOUS RECORDS: Grootfontein [ 19°57'S, 18°12'E]; Okakarara [ 20°58'S, 17°43'E]; Omatjette [ 21°05'S, 15°05'E]; Epukiro [ 21°07'S, 19°12'E] (Zumpt 1953c: 488).    KNOWN RECORDS: Grootfontein [ 19°33'S, 18°05'E],  30.iv.1950; Omatjette [ 21°05'S, 15°05'E], 11.iv.1950(NMSA)  MATERIAL:  1♂, Ondundozonananandana Mts.,  16.v–22.vi.1986, Griffin(1) ( PT);  1♂, 1♀, Okaukuejo­Duikersdrinkrds. junction on,  15.v–16.vi.1986, Griffin(1) ( PT), H65447;  2♂, 1♀, Etosha Game Parkat:  20.v–18.vi.1986, Griffin(1) ( PT), H65568/H65569;  3♀, Wolfnes­Okondekards., W of junction of,  16.v–12.vi.1986, Griffin(1) ( PT) H65403;  2♂, 4♀, Ghaub47,  16–26.vii.1986, Irish(1) ( MT);  1♀, Sturmfeld252,  iii.1986, Tölken, H65186;  1♂, Tobiroen,  15.v.1986, Curtis;  1♂, 1♀, Dorsland,  14.v–16.vi.1986, Griffin(1) ( PT);  1♀, Hereroland­Oosat:  20.i.1984, Irish(1), H58092;  1♂, Okahandja6.xii.[19]72, Gaerdes, H25130;  1♀, Sprokieswoud,  7.viii–15.ix.1987, Griffin(1) ( PT);  1♀, Waterberg Plateau Park(2),  2.ii–1.iii.1993, Pusch( PT);  2♂, 11♀, Ruacana area,  14.iii.1997, Koch( MT);  3♂, 2♀, Helio(2),  27.iii–4.v.1988, Griffin(1) ( PT);  1♂, 2♀, Okaukuejo, SE1915 Bb,  vi.1991, Versfeld;  1♀, B de la Bat Restcamp,  9–13.iv.1993, Green( MT);  1♂, 1♀, Persianer705,  25.ii.1994, Marais;  1♂, 1♀, Popa Falls(1),  26.ii–1.iii.1992, Marais& Pusch(LT);  1♀, Nakatwa,  8–13.iii.1992, Pusch& Marais( MT);  1♂, Mushare, Sof,  27.iii–28.iv.1988, Griffin(1) ( PT);  1♂, Wolfsnes, Wof,  24.iii–10.v.1988, Griffin(1) ( PT);  2♂, 4♀, Waterberg Plateau Park, Onjoka,  iv.1992, Green( MT);  1♀, Grootberg191: Khoadi// HôasM24,  24–27.v.2001, Marais& Kirk­Spriggs(1) ( PT);  1♀, Shaka: Salamabala[ sic– Salambala] M8,  27.ii–2.iii.2001, Kirk­Spriggs(1) ( MT);  1♂, 1♀, mopane in SalambalaM4,  24–26.ii.2001, Marais& Kirk­Spriggs(1) ( YP);  1♀, Simataa: Salambala B6,  24–27.ii.2001, Marais& Kirk­ Spriggs(1) ( YP);  4♀, Shaka: Salambala M8,  27.ii–2.iii.2001, Kirk­Spriggs(1) ( MT);  11♂, 10♀, Salambala, 4 kmNE of,  25.ii–1.iii.2001, Kirk­Spriggs(1) & Marais( MT);  1♀, Salambala forest,  23–29.xii.2002, Kirk­Spriggs(2) ( MT);  2♂, 3♀, Salambalacampsite,  28–29.xii.2002, Kirk­ Spriggs(2) ( HT)fish;  1♂, same except:  3–4.iii.2001, Kirk­Spriggs(1) ( HT)fish;  1♂, 3♀, same except:  23–28.xii.2002, Kirk­Spriggs(2) ( HT)mango;  18♂, 24♀, same except:  26–28.ii.2001, Marais& Kirk­Spriggs(1) ( HT)fruit;  4♂, 7♀, same except:  22–24.ii.2001, Kirk­Spriggs(1) & Marais( MT);  6♀, Salambalapan,  1–4.iii.2001, Kirk­Spriggs(1) ( MT); 1♀, same except: 23–27.xii.2002;  7♀, Simanya:  Okavango River,  23–24.i.1998, Kirk­Spriggs(1) & Marais( MT)primary woodland;  3♀, Dussi, 10 kmW of,  29–30.xii.1998, Kirk­Spriggs(1), Marais& Mann( MT);  2♂, 5♀, Divuju:  Okavango River,  1.i.1999, Kirk­Spriggs(1), Marais& Mann( MT) & (S);  1♂, Xawashapan, 2 kmW,  26–27.xii.1998, Kirk­Spriggs(1), Marais& Mann( MT);  1♂, Xawashapan,  26.xii.1998, Marais& Kirk­Spriggs(1) (S);  3♀, Namib Desert Res, Stn., Kuiseb Riv.,  14–26.ii.1997, Kapofi& Irwin( MT)riparian vegetation;  1♀, Varianto771/2,  30.iii–1.iv.2003, Kirk­Spriggs(1) ( MT); 1♀, same except: Kirk­Spriggs(1) & Mey (LT);  3♂, 2♀, same except: ( HT)fungi;  5♂, 9♀, same except:  29–30.iii.2003, Kirk­Spriggs(1) ( HT)fruit;  2♂, Mile46,  25–27.iii.2003, Kirk­Spriggs(1) ( MT);  1♂, 1♀, same except: ( HT)fruit;  1♂, 1♀, Rundu( Kavango Lodge),  28.iii.2003, Kirk­Spriggs(1) ( HT)fruit; 4♂, 2♀, same except: 27–29.iii.2003, Kirk­Spriggs(1) & Mey (LT);  4♀, Somerkoms521,  6–8.ii.2001, Kirk­ Spriggs(1), Marais& Wheeler( MT);  3♀, Luchando village,  26–27.ii.2001, Kirk­Spriggs(1) & Marais( MT);  1♂, 4♀, Xawashapan,  27.xii.1998, Kirk­Spriggs(1) ( MT);  1♂, 3♀, Mannheim Research Station,  31.i–7.ii.2000, Kirk­Spriggs(1) ( MT)mango plantation;  1♀, Salambalacampsite,  1–4.iii.2001, Kirk­Spriggs(1) ( PT)baited millipede;  3♀, Hippo Lodge( Zambezi River),  6–7.ii.2004, Kirk­Spriggs(1) ( HT)fruit;  2♂, 5♀, Toggenburg591 campsite,  18–20.ii.2004, Kirk­Spriggs(1) ( HT)fruit;  1♀, same except: ( MT);  6♂, 18♀, Popa Fallsrestcamp,  13–16.ii.2004, Kirk­Spriggs(1) ( MT), riverine forest;  3♀, Bum Hillcampsite ( Kwando River),  10–13.ii.2004, Kirk­Spriggs(1) ( MT);  14♂, 17♀, same except: ( HT)fruit;  1♂, 4♀, Salambalacampsite,  8–10.ii.2004, Kirk­Spriggs(1) ( MT);  3♂, 6♀, same except: ( HT)fruit;  6♂, 5♀, Omatjenne Agricultural Research Station,  7.ii.2004, Shihepo, fruiting  Termitomyces schimperi[all with associated puparia, 3♂, larva coll:  7.ii.2004, pupated:  15.ii.2004, emerged:  22.ii.2004; 3♂, 2♀, larva coll:  7.ii.2004, pupated:  14.ii.2004, emerged:  21.ii.2004; 3♀, larva coll:  7.ii.2004, pupated:  15.ii.2004, emerged:  21.ii.2004]; 2♂, 7♀, same except: fruiting  Termitomyces schimperi[all with associated puparia, larva coll: 5.ii.2004, pupated: 14.ii.2004, emerged: 20.ii.2004];  1♂, Veterinary ServicesHQ ( KM1)(2),   5–12.iii.2004, Mudenga( NU­L);  1♂, Hungorobravine at(1):  25.iv.2000, Meakin/ Raleigh Int.(LT) Bberg UV 31;  1♀, Upper Hungorobravine at:  7.iv.2000, Meakin/ Raleigh Int( MT)  Bberg Mal36;  1♀, Fallsrock ravine at:  17.iv.2000, Meakin/ Raleigh Int.( MT)  Bberg Mal37.  NOTES: The commonest species of the genus in Africa (Zumpt 1953c: 486) bred from mushrooms in South Africa. Cuthbertson (1935: 17, as  H. clara) notes that flies are attracted to the honeydew of scale insects ( Hemiptera: Coccidae) on fruit trees and were occasionally observed on fresh cattle dung. He briefly discusses the breeding habits and notes a large number of flies as being attracted to a mushroom growing out of a large termite hill in Zimbabwe. Later (Cuthbertson 1939: 145, as  Tricyclea clara) also notes the occurrence of larvae in wild mushrooms in December and January in Zimbabwe. In Namibiacollected in pitfall, yellow pan and Malaise traps; by sweeping, and at UV­light. Further collected in hanging traps baited with fermenting mango, fruit bait, rotting fish and rotting fungi, and in a pitfall trap baited with a dead millipede. Most significantly, the species has been reared in numbers from the fruiting bodies of the fungus  Termitomyces schimperi(Pat.), which grows from termite hills in the rainy season. Occurring at low and high elevations on the Brandberg ( 700 m, 1170 m, 1920 m). Widely distributed in central, north and north­eastern Namibia; in the ‘arid’ and ‘mesic’ savanna, nama­karoo and desert biomes ( Fig. 67). Recorded throughout the year, except the colder months of September, October and November; most abundantly in February ( vide Table 2). Cuthbertson (1935: 17, as  H. clara) briefly describes the egg, larvae and puparium and illustrates: the larval anal compartment, lateral aspect (Plate IV, 15a); posterior spiracles, from behind ( ibid, 15b); cephaloskeleton, lateral aspect and detail of the mouthooks ( ibid, 15d and 15f); anterior spiracles ( ibid, 15e), and the pupal posterior spiracles, from behind ( ibid, 15c).    Hemigymnochaeta varia(Hough, 1898: 178) ( Parochromyia).  Fig. 68. TYPE LOCALITY: Somalia. DISTRIBUTION: Widespread in East Africato southern Africa: Angola,  Democratic Republicof Congo, Kenya, Namibia, Rwanda, South Africa( Cape, Natal, Transvaal), Somalia, Tanzania,   Ugandaand Zimbabwe. PREVIOUS RECORD: Omatjette [ 21°05'S, 15°05'E] (Zumpt 1953c: 490). KNOWN RECORDS: Gobabis [ 22°45'S, 18°97'E], 11.iii.1971; Okakuejo [=Okaukuejo] [ 19°18'S, 15°93'E], 13.iii.1971; Omatjette [ 21°05'S, 15°05'E], 11.iv.1950(NMSA).  MATERIAL:  1♂, 1♀, Simanya:  Okavango River,  23–24.i.1998, Kirk­Spriggs(1) & Marais( MT)primary woodland;  1♀, Ruacana area,  14.iii.1997, Koch( MT);  1♀, Nama,  20–22.xii.1998, Kirk­ Spriggs(1), Marais& Mann( MT);  3♀, Persianer705,  25.ii.1994, Marais;  3♀, Toggenburg591 campsite,  18–20.ii.2004, Kirk­Spriggs(1) ( HT)fruit.  NOTES: Cuthbertson (1935: 17, as  H. pallens) reared larvae from the broken down fungus beds of termites and adults from Aardvark burrows in termite nests. He also observed adults of both sexes on fresh cattle dung in shade. Engel and Cuthbertson (1937: 10, as  Tricyclea angola) note larvae from mushrooms in Ugandaand provide details of the immature stages. Later (Cuthbertson 1939: 17, as  Tricyclea tridentata) noted both sexes encountered on faeces, pig dung and fresh cattle dung, plus males in blossoms. In Namibiaonly collected in Malaise traps and in a hanging trap baited with fermenting fruit. Widely distributed over central and northern Namibia; apparently restricted to the ‘arid’ savanna biome ( Fig. 68). Recorded from December to March ( vide Table 2). Cuthbertson (1935: 17–18) provides a brief description of the larva and puparium and illustrates the following larval parts: anal compartment, lateral aspect (Plate IV, 16a); posterior spiracles, from behind ( ibid, 16b); anterior spiracle ( ibid, 16c); and cephaloskeleton, lateral aspect ( ibid, 16d). [264,526,828,851] LOCALITY Liberia Liberia 66 67 1 holotype [390,1044,923,947] 2001-03-01 2001-03-03 2001-03-01 HT Kirk-Spriggs Salambala 66 67 4 4 2001-03-03 2001-03-04 2001-03-03 HT Kirk-Spriggs 66 67 2 2 2001-02-26 2001-02-28 2001-02-26 HT Marais & Spriggs 66 67 19 17 2 [554,1171,987,1011] 2004-02-08 2004-02-10 2004-02-08 HT Kirk-Spriggs 66 67 6 4 2 MT 66 67 6 5 1 [542,1269,1019,1043] 2001-02-22 2001-02-24 2001-02-22 MT Kirk-Spriggs & Marais 66 67 1 1 2001-03-01 2001-03-04 2001-03-01 PT Kirk-Spriggs 66 67 1 1 2004-02-10 2004-02-13 2004-02-10 HT Kirk-Spriggs Kwando Bum Hill 66 67 2 2 Rivers [997,1267,1083,1107] MT 66 67 1 1 Rivers 2004-02-18 2004-02-20 2004-02-18 HT Kirk-Spriggs Toggenburg 66 67 1 1 Rivers 2004-02-13 2004-02-16 2004-02-13 MT Kirk-Spriggs Popa Falls 66 67 6 2 4 Rivers [956,1200,1147,1171] HT 66 67 1 1 Rivers [264,707,1436,1459] LOCALITY Zimbabwe Zimbabwe 66 67 1 holotype [264,540,1788,1811] LOCALITY Namibia Namibia 66 67 1 holotype [264,540,412,435] LOCALITY Namibia Namibia 67 68 1 holotype [264,585,636,659] LOCALITY Sierra Leone Sierra Leone 67 68 1 holotype RECORDS -21.116667 1265 19.2 67 68 1 [264,972,860,883] 1950-04-30 KNOWN, RECORDS -19.55 1271 18.083334 67 68 1 [390,1124,923,947] 1986-05-16 1986-06-22 1986-05-16 PT Griffin Ondundozonananandana Mts. 67 68 1 1 H65447 1986-05-15 1986-06-16 1986-05-15 PT Griffin Okaukuejo-Duikersdrink 67 68 2 1 1 H65568, H65569 1986-05-20 1986-06-18 1986-05-20 PT Griffin Etosha Game Park 67 68 3 1 2 H65403 1986-05-16 1986-06-12 1986-05-16 PT Griffin Wolfnes-Okondeka 67 68 3 3 1986-07-16 1986-07-26 1986-07-16 MT Irish Ghaub 67 68 6 4 2 H65186 [473,963,1051,1075] 1986-03 Tolken Sturmfeld 67 68 1 1 [975,1319,1051,1075] 1986-05-15 Curtis Tobiroen 67 68 1 1 [312,858,1083,1107] 1986-05-14 1986-06-16 1986-05-14 PT Griffin Dorsland 67 68 2 1 1 H58092 1984-01-20 Irish Hereroland-Oos 67 68 1 1 H25130 [413,911,1115,1139] Gaerdes Okahandja 67 68 1 1 1987-08-07 1987-09-15 1987-08-07 PT Griffin Sprokieswoud 67 68 1 1 [486,1116,1147,1171] 1993-02-02 1993-03-01 1993-02-02 PT Pusch Park Waterberg 67 68 1 1 Plateau 1997-03-14 MT Koch Ruacana area 67 68 13 11 2 Plateau [659,1217,1179,1203] 1988-03-27 1988-05-04 1988-03-27 PT Griffin Helio 67 68 5 2 3 Plateau 1991-06 Okaukuejo, SE & Versfeld Bb 67 68 3 2 1 Plateau 1993-04-09 1993-04-13 1993-04-09 MT B de la Bat Restcamp & Green 67 68 1 1 Plateau [390,867,1243,1267] 1994-02-25 Marais Persianer 67 68 2 1 1 Plateau 1992-02-26 1992-03-01 1992-02-26 Marais & Pusch Popa Falls 67 68 2 1 1 Plateau [555,1107,1275,1299] 1992-03-08 1992-03-13 1992-03-08 MT Pusch & Marais Nakatwa 67 68 1 1 Plateau 1988-03-27 1988-04-28 1988-03-27 PT Mushare, S & Griffin 67 68 1 1 Plateau [685,1271,1307,1331] 1988-03-24 1988-05-10 1988-03-24 PT Wolfsnes, W & Griffin 67 68 1 1 Plateau 1992-04 MT Green Waterberg Onjoka Park 67 68 6 4 2 Plateau 2001-05-24 2001-05-27 2001-05-24 PT Marais & Kirk-Spriggs Grootberg Hoas Khoadi 67 68 1 1 Plateau 2001-02-27 2001-03-02 2001-02-27 MT Kirk-Spriggs Salamabala Shaka 67 68 1 1 Plateau 2001-02-24 2001-02-26 2001-02-24 YP Marais & Kirk-Spriggs Salambala 67 68 2 1 1 Plateau 2001-02-24 2001-02-27 2001-02-24 YP Salambala B & Marais & Spriggs Simataa 67 68 1 1 Plateau [497,1204,1467,1491] 2001-02-27 2001-03-02 2001-02-27 MT Salambala M & Kirk-Spriggs Shaka 67 68 4 4 Plateau 2001-02-25 2001-03-01 2001-02-25 MT Kirk-Spriggs & Marais Salambala 67 68 21 10 11 Plateau 2002-12-23 2002-12-29 2002-12-23 MT Kirk-Spriggs Salambala forest 67 68 1 1 Plateau 2002-12-28 2002-12-29 2002-12-28 HT Spriggs Salambala 67 68 5 3 2 Plateau [548,1115,1563,1587] 2001-03-03 2001-03-04 2001-03-03 HT Kirk-Spriggs 67 68 1 1 Plateau 2002-12-23 2002-12-28 2002-12-23 HT Kirk-Spriggs 67 68 4 3 1 Plateau 2001-02-26 2001-02-28 2001-02-26 HT Marais & Kirk-Spriggs 67 68 42 24 18 Plateau 2001-02-22 2001-02-24 2001-02-22 MT Kirk-Spriggs & Marais 67 68 11 7 4 Plateau [478,1108,1659,1683] 2001-03-01 2001-03-04 2001-03-01 MT Kirk-Spriggs Salambala 67 68 6 6 Plateau [487,1323,1691,1715] 1998-01-23 1998-01-24 1998-01-23 MT Kirk-Spriggs & Marais Simanya 67 68 7 7 Okavango River 1998-12-29 1998-12-30 1998-12-29 MT Kirk-Spriggs & Marais & Mann Dussi 67 68 3 3 Okavango River 1999-01-01 MT Kirk-Spriggs & Marais & Mann Divuju 67 68 7 5 2 Okavango River [360,1272,1787,1811] 1998-12-26 1998-12-27 1998-12-26 MT Kirk-Spriggs & Marais & Mann Xawasha 67 68 1 1 Okavango River 1998-12-26 Marais & Kirk-Spriggs Xawasha 67 68 1 1 Okavango River 1997-02-14 1997-02-26 1997-02-14 MT Kapofi & Irwin Namib Desert Res Kuiseb Riv. Stn. 67 68 3 3 Okavango River 2003-03-30 2003-04-01 2003-03-30 MT Kirk-Spriggs 68 69 Varianto 67 68 1 1 Okavango River HT 68 69 5 2 3 Okavango River [631,1263,315,339] 2003-03-29 2003-03-30 2003-03-29 HT Kirk-Spriggs 68 69 14 9 5 Okavango River [312,866,347,371] 2003-03-25 2003-03-27 2003-03-25 MT Kirk-Spriggs Mile 68 69 2 2 Okavango River [878,1171,347,371] HT 68 69 2 1 1 Okavango River 2003-03-28 HT Kirk-Spriggs Lodge Rundu 68 69 2 1 1 Kavango West 2001-02-06 2001-02-08 2001-02-06 MT Spriggs & Marais & Wheeler Somerkoms 68 69 4 4 Kavango West 2001-02-26 2001-02-27 2001-02-26 MT Kirk-Spriggs & Marais Luchando village 68 69 3 3 Kavango West [469,1100,475,499] 1998-12-27 MT Kirk-Spriggs Xawasha 68 69 5 4 1 Kavango West 2000-01-31 2000-02-07 2000-01-31 MT Kirk-Spriggs Mannheim Research Station 68 69 4 3 1 Kavango West 2001-03-01 2001-03-04 2001-03-01 PT Kirk-Spriggs Salambala 68 69 1 1 Kavango West 2004-02-06 2004-02-07 2004-02-06 HT Kirk-Spriggs Hippo Lodge 68 69 3 3 Zambezi 2004-02-18 2004-02-20 2004-02-18 HT Kirk-Spriggs Toggenburg 68 69 7 5 2 Zambezi [777,1034,603,627] MT 68 69 1 1 Zambezi 2004-02-13 2004-02-16 2004-02-13 MT Kirk-Spriggs Popa Falls 68 69 24 18 6 Zambezi 2004-02-10 2004-02-13 2004-02-10 MT Kirk-Spriggs Kwando Bum Hill 68 69 3 3 Rivers [824,1158,667,691] HT 68 69 31 17 14 Rivers 2004-02-08 2004-02-10 2004-02-08 MT Kirk-Spriggs Salambala 68 69 5 4 1 Rivers [933,1219,699,723] HT 68 69 9 6 3 Rivers 2004-02-07 2004-02-22 2004-02-07 Shihepo Omatjenne Agricultural Research Station 68 69 22 10 12 Rivers [548,960,891,915] KM Veterinary Services 68 69 KM1 1 1 Rivers [971,1319,892,915] 2004-03-05 2004-03-12 2004-03-05 NU-L Mudenga 68 69 1 Rivers [312,1192,923,947] 2000-04-25 UV Meakin & Raleigh Int. & Bberg Hungorob 68 69 UV 31 1 1 Rivers 2000-04-07 MT Meakin & Raleigh Int Upper Hungorob 68 69 1 1 Rivers [931,1083,956,979] Bberg Mal 68 69 1 Rivers 2000-04-17 MT Meakin & Raleigh Int. Falls 68 69 1 1 Rivers [757,908,988,1011] Bberg Mal 68 69 1 Rivers LOCALITY Somalia Democratic Republic Widespread in East Africa southern Africa 68 69 1 RECORD Uganda -21.083334 Uganda 1265 15.083333 68 69 1 [389,1275,347,371] 1998-01-23 1998-01-24 1998-01-23 MT Kirk-Spriggs & Marais Simanya 69 70 2 1 1 Okavango River [494,954,379,403] 1997-03-14 MT Koch Ruacana area 69 70 1 1 Okavango River 1998-12-20 1998-12-22 1998-12-20 MT Spriggs & Marais & Mann Nama 69 70 1 1 Okavango River [688,1092,411,435] 1994-02-25 Marais Persianer 69 70 3 3 Okavango River 2004-02-18 2004-02-20 2004-02-18 HT Kirk-Spriggs Toggenburg 69 70 3 3 Okavango River