Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae) HUBER, BERNHARD A. Zootaxa 2012 2012-09-07 3461 1 1 138 6YS5Q Huber, 2012 Huber 2012 [151,424,1297,1323] Arachnida Pholcidae Smeringopus Animalia Araneae 30 31 Arthropoda species ngangao sp. nov.   Figs. 6–7, 138–154, 165–168 Type. Male holotypefrom Kenya, Coast Province, Taita Hills, Ngangao Forest( 3°22.2’S, 38°20.4’E),  1810 ma.s.l.,  19.i.2010( B.A. Huber), in ZFMK( Ar 8546). Etymology. The name is a noun in apposition, derived from the typelocality. Diagnosis. Small species, easily distinguished from known congeners by shapes of procursus and embolus ( Figs. 149–152) and by cheliceral armature (one pair of relatively large apophyses in unusual frontal and proximal position; Figs. 144, 153). Male ( holotype). Total body length 4.5, carapace width 1.7. Leg 1: 39.6 (10.6 + 0.6 + 9.6 + 17.5 + 1.3), tibia 2: 6.9, tibia 3: 5.5, tibia 4: 7.1; tibia 1 L/d: 62. Habitus as in Figs. 138 and 139. Carapace brown with lighter areas beside ocular area, clypeus with indistinct pair of brown lines and brown rim, sternum monochromous brown, legs brown, tips of femora and tibiae lighter, abdomen with distinct pattern dorsally and ventrally. Distance PME-PME 115 µm, diameter PME 170 µm, distance PME-ALE 45 µm, distance AME-AME 45 µm, diameter AME 140 µm. Ocular area slightly elevated, secondary eyes apparently without ‘pseudo-lenses’; deep thoracic pit. Chelicerae as in Fig. 153, with one pair of relatively large apophyses in unusual frontal and proximal position; each with one modified hair at tip ( Fig. 166). Palps as in Figs. 142 and 143, coxa unmodified, trochanter with small retrolateral projection, femur with retrolateral furrow, proximal rim with apophysis, cymbium without projection near tarsal organ, procursus with large whitish protrusion prolaterally ( Figs. 149, 150), bulb with simple hooked embolus ( Figs. 151, 152). Legs without spines, few vertical hairs, without curved hairs, retrolateral trichobothrium on tibia 1 at 1.5%; prolateral trichobothrium present on tibia 1. Gonopore with two epiandrous spigots ( Fig. 165).   FIGURES 155–168.  Smeringopus chogoria(155–164) and  S. ngangao(165–168). 155–156. Right palp, retrolatero-dorsal and prolatero-dorsal views. 157–158. Left palp, dorsal and prolatero-dorsal views. 159. Male gonopore. 160–161. Male chelicerae and cheliceral apophysis (arrow points to modified hair). 162–163. Male and female ALS. 164. Epigynum. 165. Male gonopore. 166. Modified hair on male cheliceral apophysis. 167. Female ALS. 168. Epigynum. Scale lines: 10 µm (166), 20 µm (161, 167), 30 µm (162, 163), 50 µm (165), 80 µm (159), 100 µm (160), 200 µm (155–158, 168), 300 µm (164). Variation. Tibia 1 in9 other males: 8.0–9.6 (mean 9.1). In males from Amani ( Figs. 140, 141), the tip of the procursus is minimally different and the embolus is slightly more slender. Female. In general similar to male; tibia 1 in 10 females: 7.6–8.4 (mean 7.9). Epigynum simple anterior plate with pair of dark dots, without pockets, simple arc-shaped posterior plate ( Figs. 145, 146, 168); internal genitalia as in Figs. 147and 154. ALS with eight spigots each ( Fig. 167). Females from Amani with distinct x-shaped structure in internal female genitalia ( Fig. 148). Distribution. Known from TaitaHills in southern Kenyaand Usambara Mountains in northeastern Tanzania( Fig. 58). Material examined.  KENYA: Coast Prov.: TaitaHills, Ngangao Forest: 1♂ holotypeabove ; same data, 3♂ 7♀in ZFMK( 2♂ 6♀, Ar8547-48) and NMKE( 1♂ 1♀) ; same data, 3♀2 juvs in pure ethanol in ZFMK( Ken90).  Taita Hills, roadside near Ngangao Forest(~ 3°22.2’S, 38°20.4’E),  1810 ma.s.l.,  19.i.2010( B.A. Huber), 1♂in ZFMK( Ar8549).  Taita Hills, Chavia Forest( 3°28.8’S, 38°20.4’E),  1590 ma.s.l.,  20.i.2010( B.A. Huber), 4♂ 1♀in ZFMK( Ar8550); same data, 1♂ 2♀in pure ethanol in ZFMK( Ken96).    TANZANIA:  Tanga Region: East Usambara Mountains, Amani( 5°05.7’S, 38°38’E),  950 ma.s.l.,  27.x.–9.xi.1995( C.E. Griswold, N. Scharff, D. Ubick), 2♂ 4♀in CAS. 3747820412 2010-01-19 ZFMK B. A. Huber Kenya Coast Province 1810 -3.37 Taita Hills 130 38.34 Hills 30 31 Ar 8546 1 Taita/Taveta holotype 3747820744 [425,1227,295,320] 2010-01-19 ZFMK, NMKE B. A. Huber Ken 1810 -3.37 Ar 130 38.34 Hills 32 33 10 7 3 Taita/Taveta holotype 3747820357 2010-01-19 ZFMK, NMKE B. A. Huber Ken 1810 -3.37 Ar 130 38.34 Hills 32 33 10 7 3 Taita/Taveta 3747820779 [728,1374,332,356] 2010-01-19 ZFMK, NMKE B. A. Huber Ken 1810 -3.37 Ar 130 38.34 Hills 32 33 10 7 3 Taita/Taveta 3747820840 2010-01-19 ZFMK, NMKE B. A. Huber Ken 1810 -3.37 Ar 130 38.34 Hills 32 33 10 7 3 Taita/Taveta 3747820738 2010-01-19 ZFMK, NMKE B. A. Huber Ken 1810 -3.37 Ar 130 38.34 Hills 32 33 10 7 3 Taita/Taveta 3747820742 [361,956,439,463] 2010-01-19 ZFMK, NMKE B. A. Huber Ken 1810 -3.37 Ar 130 38.34 Hills 32 33 10 7 3 Taita/Taveta 3747820345 1995-10-27 1995-11-09 1995-10-27 CAS C. E. Griswold & N. Scharff & D. Ubick Tanzania 950 -5.095 Amani 926 38.633335 East Usambara Mountains 32 33 6 4 2 Tanga