Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae)
HUBER, BERNHARD A.
Zootaxa
2012
2012-09-07
3461
1
1
138
4XVHS
Huber, 2012
Huber
2012
[151,409,1405,1431]
Arachnida
Pholcidae
Smeringopus
Animalia
Araneae
50
51
Arthropoda
species
koppies
sp. nov.
Figs. 245–246, 258–259, 280–281, 305–309 Type. Male holotypefrom South Africa, Free State, Koppiesdam Nat. Res.( 27°13’S, 27°42’E), 1400 ma.s.l., 28.ix.1993( L.N. Lotz), in NMBA(6609 part). Etymology. The name is a noun in apposition, derived from the typelocality. Diagnosis. Distinguished from similar congeners (other species of the natalensisgroup, especially S. natalensis, S. florisbad, S. lesnei, S. blyde, S. harare, S. badplaas) by shapes of bulbal processes ( Figs. 307, 308); from other close relatives by absence of process near palpal tarsal organ ( Fig. 305), relatively straight procursus (ventrally), absence of prolateral process on procursus tip, and three black lines ventrally on abdomen (versus two; compare Fig. 246with Figs. 248 and 250). Male ( holotype). Total body length 6.4, carapace width 2.0. Leg 1: 42.4 (11.3 + 0.8 + 11.1 + 16.7 + 2.5), tibia 2: 7.7, tibia 3: 5.9, tibia 4: 8.3; tibia 1 L/d: 51. Habitus as in Figs. 245 and 246. Carapace ochre-yellow with distinct dark pattern (median, lateral, and submarginal marks), clypeus with pair of dark marks widening distally, sternum brown with light marks, legs with barely visible darker rings subdistally on femora and tibiae, abdomen dorsally with distinct dark pattern, ventrally with three dark lines in median part (median line narrow but distinct). Distance PME-PME 175 µm, diameter PME 175 µm, distance PME-ALE 70 µm, distance AME-AME 45 µm, diameter AME 150 µm. Ocular area slightly elevated, secondary eyes with very indistinct ‘pseudo-lenses’; deep thoracic pit. Chelicerae as in S. badplaas(cf. Figs. 314, 315; just slightly larger). Palps as in Figs. 258 and 259, coxa without retrolateral apophysis, trochanter barely modified, femur with retrolateral furrow with distinct rim proximally, cymbium without projection near tarsal organ, procursus ventrally almost straight, without prolateral process at tip ( Figs. 305, 306), bulb with three distinctively shaped processes ( Figs. 307, 308). Legs without spines, few vertical hairs, with curved hairs on tibiae and metatarsi 1 and 2, retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium present on tibia 1. Gonopore with two epiandrous spigots. Variation. Tibia 1 in5 other males: 11.1–12.4 (mean 11.6). Female. In general similar to male; tibia 1 in 8 females: 7.7–14.0 (mean 10.3). Epigynum a simple plate without pockets ( Fig. 280), variable in shape and coloration and thus possibly not clearly distinguishable from close relatives ( S. natalensis, S. florisbad, S. lesnei, S. blyde, S. harare, S. badplaas); internal genitalia as in Figs. 281and 309(also similar to close relatives). Distribution. Known from several localities in northern South Africaand southern Botswana( Fig. 299). FIGURES 300–309. Smeringopus natalensis(300–304) and S. koppies(305–309). 300, 305. Left cymbia and procursi, retrolateral views. 301, 306. Left procursi, prolateral views. 302–303, 307–308. Left bulbal processes, prolateral and dorsal views. 304, 309. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm. Materialexamined. SOUTH AFRICA: Free State: Koppiesdam Nat. Res.: 1♂ holotypeabove; same data, 1♂ 1♀in NMBA(6609 part). Farm Lusthof, Edenville[ 27°32’S, 27°39’E], 5.iv.1967( N. Genis), 1♂ 1♀ 1 juv.in NCP(76/2018). Northern Cape Province: Kimberley[ 28°44’S, 24°46’E], “Hebrou”, no date ( E. Simon), E. Simoncollection 15271, 1♂in MNHN( AR 10353). Northwest Province: Mafeking[=Mafikeng, 25°51’S, 25°38’E], ii.1905( Schultze), 1♂6 juvs in ZMB(1132).
BOTSWANA: South-East: Gaborone[ 24°40.0’S, 25°56.7’E], Museumyard, iii.1974( J.K. Ullberg), 5♂ 6♀+ juvs in ZMT(3532); Gaborone, 16.vii.1973(R. Hakanen), 1♀ 1 juv.in ZMT(3381); Gaborone, inside house, 4.vii.1973(R. Hakanen), 4♀2 juvs in ZMT(3382).
3747820397
1993-09-28
NMBA
L. N. Lotz
South Africa
1400
-27.216667
Koppiesdam Nat. Res.
1238
27.7
50
51
1
Free State
holotype
3747820780
NMBA
South Africa
Koppiesdam Nat. Res.
Material
52
53
3
1
2
Free State
holotype
3747820663
1967-04-05
N. Genis
South Africa
-27.533333
Edenville
1236
27.65
Farm Lusthof
52
53
3
1
1
Free State
holotype
3747820526
MNHN
E. Simon
South Africa
-28.733334
Kimberley
1230
24.766666
52
53
AR 10353
1
1
Northern Cape
holotype
3747820478
1905-02
ZMB
Schultze
South Africa
-25.85
Mafeking
1244
25.633333
52
53
1
1
Northwest Province
holotype
3747820947
1974-03
ZMT
Museum & J. K. Ullberg
Botswana
-24.666666
South-East
125
25.945
52
53
11
6
5
South East
3747820586
[403,1142,367,391]
1973-07-16
R, ZMT
Hakanen
Botswana
Gaborone
52
53
2
1
Gaborone
3747820891
1973-07-04
R, ZMT
Hakanen
Botswana
Gaborone
52
53
4
4
Gaborone