Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae) HUBER, BERNHARD A. Zootaxa 2012 2012-09-07 3461 1 1 138 4XVHS Huber, 2012 Huber 2012 [151,409,1405,1431] Arachnida Pholcidae Smeringopus Animalia Araneae 50 51 Arthropoda species koppies sp. nov.   Figs. 245–246, 258–259, 280–281, 305–309 Type.  Male holotypefrom South Africa, Free State, Koppiesdam Nat. Res.( 27°13’S, 27°42’E),  1400 ma.s.l.,  28.ix.1993( L.N. Lotz), in NMBA(6609 part). Etymology. The name is a noun in apposition, derived from the typelocality. Diagnosis. Distinguished from similar congeners (other species of the  natalensisgroup, especially  S. natalensis, S. florisbad, S. lesnei, S. blyde, S. harare, S. badplaas) by shapes of bulbal processes ( Figs. 307, 308); from other close relatives by absence of process near palpal tarsal organ ( Fig. 305), relatively straight procursus (ventrally), absence of prolateral process on procursus tip, and three black lines ventrally on abdomen (versus two; compare Fig. 246with Figs. 248 and 250). Male ( holotype). Total body length 6.4, carapace width 2.0. Leg 1: 42.4 (11.3 + 0.8 + 11.1 + 16.7 + 2.5), tibia 2: 7.7, tibia 3: 5.9, tibia 4: 8.3; tibia 1 L/d: 51. Habitus as in Figs. 245 and 246. Carapace ochre-yellow with distinct dark pattern (median, lateral, and submarginal marks), clypeus with pair of dark marks widening distally, sternum brown with light marks, legs with barely visible darker rings subdistally on femora and tibiae, abdomen dorsally with distinct dark pattern, ventrally with three dark lines in median part (median line narrow but distinct). Distance PME-PME 175 µm, diameter PME 175 µm, distance PME-ALE 70 µm, distance AME-AME 45 µm, diameter AME 150 µm. Ocular area slightly elevated, secondary eyes with very indistinct ‘pseudo-lenses’; deep thoracic pit. Chelicerae as in  S. badplaas(cf. Figs. 314, 315; just slightly larger). Palps as in Figs. 258 and 259, coxa without retrolateral apophysis, trochanter barely modified, femur with retrolateral furrow with distinct rim proximally, cymbium without projection near tarsal organ, procursus ventrally almost straight, without prolateral process at tip ( Figs. 305, 306), bulb with three distinctively shaped processes ( Figs. 307, 308). Legs without spines, few vertical hairs, with curved hairs on tibiae and metatarsi 1 and 2, retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium present on tibia 1. Gonopore with two epiandrous spigots. Variation. Tibia 1 in5 other males: 11.1–12.4 (mean 11.6). Female. In general similar to male; tibia 1 in 8 females: 7.7–14.0 (mean 10.3). Epigynum a simple plate without pockets ( Fig. 280), variable in shape and coloration and thus possibly not clearly distinguishable from close relatives (  S. natalensis, S. florisbad, S. lesnei, S. blyde, S. harare, S. badplaas); internal genitalia as in Figs. 281and 309(also similar to close relatives). Distribution. Known from several localities in northern South Africaand southern Botswana( Fig. 299).   FIGURES 300–309.  Smeringopus natalensis(300–304) and  S. koppies(305–309). 300, 305. Left cymbia and procursi, retrolateral views. 301, 306. Left procursi, prolateral views. 302–303, 307–308. Left bulbal processes, prolateral and dorsal views. 304, 309. Cleared female genitalia, dorsal views. Scale lines: 0.3 mm.  Materialexamined.  SOUTH AFRICA:  Free State: Koppiesdam Nat. Res.: 1♂ holotypeabove; same data, 1♂ 1♀in NMBA(6609 part).  Farm Lusthof, Edenville[ 27°32’S, 27°39’E],  5.iv.1967( N. Genis), 1♂ 1♀ 1 juv.in NCP(76/2018).   Northern Cape Province: Kimberley[ 28°44’S, 24°46’E], “Hebrou”, no date ( E. Simon), E. Simoncollection 15271, 1♂in MNHN( AR 10353).   Northwest Province: Mafeking[=Mafikeng, 25°51’S, 25°38’E],  ii.1905( Schultze), 1♂6 juvs in ZMB(1132).    BOTSWANA:  South-East: Gaborone[ 24°40.0’S, 25°56.7’E], Museumyard,  iii.1974( J.K. Ullberg), 5♂ 6♀+ juvs in ZMT(3532);  Gaborone,  16.vii.1973(R. Hakanen), 1♀ 1 juv.in ZMT(3381);  Gaborone, inside house,  4.vii.1973(R. Hakanen), 4♀2 juvs in ZMT(3382). 3747820397 1993-09-28 NMBA L. N. Lotz South Africa 1400 -27.216667 Koppiesdam Nat. Res. 1238 27.7 50 51 1 Free State holotype 3747820780 NMBA South Africa Koppiesdam Nat. Res. Material 52 53 3 1 2 Free State holotype 3747820663 1967-04-05 N. Genis South Africa -27.533333 Edenville 1236 27.65 Farm Lusthof 52 53 3 1 1 Free State holotype 3747820526 MNHN E. Simon South Africa -28.733334 Kimberley 1230 24.766666 52 53 AR 10353 1 1 Northern Cape holotype 3747820478 1905-02 ZMB Schultze South Africa -25.85 Mafeking 1244 25.633333 52 53 1 1 Northwest Province holotype 3747820947 1974-03 ZMT Museum & J. K. Ullberg Botswana -24.666666 South-East 125 25.945 52 53 11 6 5 South East 3747820586 [403,1142,367,391] 1973-07-16 R, ZMT Hakanen Botswana Gaborone 52 53 2 1 Gaborone 3747820891 1973-07-04 R, ZMT Hakanen Botswana Gaborone 52 53 4 4 Gaborone