Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae) HUBER, BERNHARD A. Zootaxa 2012 2012-09-07 3461 1 1 138 4XVJ3 Huber, 2012 Huber 2012 [151,433,1118,1144] Arachnida Pholcidae Smeringopus Animalia Araneae 118 119 Arthropoda species mayombe sp. nov.  Type.  Male holotypefrom CongoD.R., Congo Centrale Prov.(“Bas-Congo”), Mayombe, Luki Forest Reserve( 5°37.3’S, 13°05.9’E), hand catch, edge of road near guest house,  4.xi.2006( D. de Bakker, J.P. Michiels), in MRAC( 221613part). Etymology. The name is a noun in apposition, derived from the typelocality. Diagnosis. Distinguished from similar congeners (other species of the  thomensisspecies group) by shape of embolus (one large and one tiny apophysis arising from embolus; Figs. 729, 730; similar  S. principe); from  S. thomensisalso by pair of sclerotized areas on posterior rim of epigynum ( Fig. 715) and wider pore plates ( Fig. 731); from  S. principealso by absence of dorsal projection from procursus tip (procursus as in  S. thomensis, cf. Figs. 719, 720). Easily distinguished from all other congeners by distinctive color pattern dorsally on abdomen (figure composed of three black elements on posterior half; Fig. 701). Male ( holotype). Total body length 6.1, carapace width 1.8. Leg 1: 53.3 (13.7 + 0.8 + 12.8 + 23.7 + 2.3), tibia 2: 8.7, tibia 3: 6.1, tibia 4: 8.5; tibia 1 L/d: 60. Habitus as in Fig. 701. Carapace ochre-yellow with light brown median mark and narrow lateral marks, clypeus distal half darker, sternum dark brown with small light spots, legs ochre-yellow, femora and tibiae with dark subdistal rings and light tips, abdomen grey with distinctive dorsal pattern, ventrally with dark anterior and posterior plates and three indistinct bands in median section. Distance PME-PME 125 µm, diameter PME 160 µm, distance PME-ALE 60 µm, distance AME-AME 25 µm, diameter AME 135 µm. Ocular area slightly elevated, secondary eyes with small dark ‘pseudo-lenses’; deep thoracic pit. Chelicerae with pair of small distal apophyses as in  S. thomensis(cf. Fig. 721). Palps in general as in  S. thomensis(cf. Figs. 705, 706), procursus not distinguishable (cf. Figs. 719, 720), only bulb different, with one large and one tiny dorsal processes ( Figs. 729, 730). Legs without spines, few vertical hairs, without curved hairs, retrolateral trichobothrium on tibia 1 at 3%; prolateral trichobothrium present on tibia 1. Variation. Tibia 1 in3 other males: 10.9, 11.3, 11.3. Posterior part of median mark on carapace sometimes barely visible.   FIGURES 732–744.  Smeringopus thomensis(732–740) and  S. principe(741–744). 732. Left male palpal femur, retrolateral view. 733. Left procursus tip, prolateral view. 734. Male cheliceral apophysis. 735. Left embolus, distal view. 736. Male gonopore. 737. Female ALS. 738. Epigynum. 739. Cleared female genitalia, dorsal view. 740. Detail of pore plate. 741. Left embolus, distal view. 742. Left procursus tip, prolateral view. 743. Cleared female genitalia, dorsal view. 744. Detail of pore plate. Scale lines: 10 µm (734, 737), 20 µm (740), 30 µm (744), 80 µm (733, 735–736), 100 µm (739, 741–742), 200 µm (732, 738, 743). Female. In general similar to male; tibia 1 in 11 females: 6.7–11.3 (mean 9.7). Epigynum a simple, roughly triangular plate with pair of pockets and pair of sclerotized areas on posterior rim, transversal internal structure variably well visible, posterior plate semicircular ( Figs. 715, 716); internal genitalia as in Figs. 717and 731. Distribution. Only known from typelocality in western CongoD.R. ( Fig. 718).  Materialexamined.  CONGOD.R.:  Congo Centrale Prov.: Mayombe, Luki Forest Reserve: typeabove, together with 2♀; same locality, fogging in primary rainforest,  4.–13.xi.2006and 26.ix.–4.x.2007(8 vials) 5♂ 13♀in MRAC(parts of 219850, 219852–53, 219855, 224312, 224315, 224318–19); fogging in old secondary rainforest,  18.–25.ix.2007(5 vials), 9♀in MRAC(parts of 224304–07, 224311); beating in old secondary rainforest,  17.ix.2007and 22.ix.2007(2 vials), 3♂ 2♀ 1 juv.in MRAC(parts of 223120, 224308); beating in primary rainforest,  29.ix.2007, 1♀in MRAC(223842 part); beating along trails near guest house,  9.xi.2006and 19.ix.2007(2 vials), 3♀in MRAC(parts of 221618, 223761); central zone of Luki F.R.,  26.–27.ix.2007( W. Hubau), 1♀in MRAC(222155 part). 3747820607 2006-11-04 MRAC D. de Bakker & J. P. Michiels Republic of the Congo Centrale Prov. -5.621667 Luki Forest Reserve 130 13.098333 Mayombe 118 119 221613 1 Bas-Congo holotype 3747820903 2007-09-18 2007-09-29 2007-09-18 R, MRAC Luki F. & W. Hubau Democratic Republic of the Congo Material Mayombe Centrale Prov. 120 121 40 31 8 Bas-Congo 3747820422 2007-09-18 2007-09-29 2007-09-18 R, MRAC Luki F. & W. Hubau Democratic Republic of the Congo Material Mayombe Centrale Prov. 120 121 40 31 8 Bas-Congo 3747820659 2007-09-18 2007-09-29 2007-09-18 R, MRAC Luki F. & W. Hubau Democratic Republic of the Congo Material Mayombe Centrale Prov. 120 121 40 31 8 Bas-Congo 3747820429 2007-09-18 2007-09-29 2007-09-18 R, MRAC Luki F. & W. Hubau Democratic Republic of the Congo Material Mayombe Centrale Prov. 120 121 40 31 8 Bas-Congo 3747820582 2007-09-18 2007-09-29 2007-09-18 R, MRAC Luki F. & W. Hubau Democratic Republic of the Congo Material Mayombe Centrale Prov. 120 121 40 31 8 Bas-Congo 3747820661 2007-09-18 2007-09-29 2007-09-18 R, MRAC Luki F. & W. Hubau Democratic Republic of the Congo Material Mayombe Centrale Prov. 120 121 40 31 8 Bas-Congo