Quedius ( Sauridus ) boluensis KORGE, 1971: 49 Quedius ( Sauridus ) deuvei COIFFAIT, 1978a: 170 Quedius ( Sauridus ) boluensis Korge Quedius ( Sauridus ) boluensis Korge On some species of the Quedius obliqueseriatus group, with notes on Q. nivicola (Coleoptera: Staphylinidae: Staphylininae) Assing, Volker Linzer biologische Beiträge 2016 2016-12-19 48 2 1137 1148 4R32J KORGE, 1971 Korge 1971 [178,638,1235,1257] Insecta Staphylinidae Quedius Animalia Coleoptera 1 1138 Arthropoda species boluensis Raphirus      Quedius( Sauridus) boluensis KORGE, 1971: 49f.     Quedius( Sauridus) deuvei COIFFAIT, 1978a: 170f. T y p e m a t e r i a l e x a m i n e d: Holotype6 [aedeagus missing]: "leg. H. Korge, Anatoliabor. / Abant-Geb. 1100-1500 m/ 6 - Holotypus  Quedius( Sauridus) boluensisKorge" (MNB). Paratype ♀: " Anatoliabor., W. Heinz leg. / Boludagl. bei Mengen 900 m, 21.7.1963/ ♀- Paratypus  Quedius( Sauridus) boluensisKorge" (MNB).   A d d i t i o n a l m a t e r i a l e x a m i n e d: Turkey: B o l u 16, 7 kmS Bolu, 40°40'N, 31°38'E,  930 m, oak forest, sifted,  29.III.2010, leg. Assing(cAss); 16, Bolu- Yığılca, 40°51'N, 31°39'N,  1060 m,  29.IV.2012, leg. Brachat& Meybohm(cAss);  1♀, 11 kmS Bolu, 40°40'N, 31°38'N,  1080 m,  30.IV.2012, leg. Brachat& Meybohm(cAss); 366, Abant mountainsnear Bolu, pitfall trap,  29.IV.-20. V.1976, leg Korge( MNB);  1♀, forest between Boluand Yiğilca,  1100 m,  15.VII.-26.VIII.1972, leg. Heinz( MNB). Z o n g u l d a k 2♀♀, ca.  15 kmW Devrek, 41°14'N, 31°52'E,  700 m, beech forest with rhododendron undergrowth, litter sifted,  26.III.2010, leg. Assing(cAss). K a s t a m o n u: 16, pass N Çambaşı,  1700 m, leg. Korge( MNB); 16, 1♀, W-Yaraligöz, S Çatalzeytin,  1300-1700 m,  25.VII.1970( MNB). S i n o p: 1♀,  25 kmS Sinop, W Lala, 41°44'N, 35°01'E,  215 m, oak forest with undergrowth, sifted,  31.III.2009, leg. Assing(cAss); 16, 1♀, ca.  30 kmS Ayancık, Çangal Dağı, 41°47'N, 34°40'E,  820 m, beech forest, sifted,  1.IV.2009, leg. Assing(cAss);  1♀, ca.  30 kmS Ayancık, Çangal Dağı, 41°49'N, 34°39'E,  920 m, beech forest, sifted,  1.IV.2009, leg. Assing(cAss); 16, S Ayancık, Çangal Dağı,  1100 m,  26.VII.1970, leg. Heinz( MNB); 16, Dranaz Dağınear Sinop,  1250 m,  27.VII.1970, leg. Heinz( MNB). S a m s u n: 1♀, 20 kmS Samsun, NE Asarcık, 41°07'N, 36°17'E,  840 m, Quercus - Carpinusforest with rhododendron, litter sifted,  20.VII.2008, leg. Assing(cAss). O r d u: 266, 1♀, ca. 30 kmSW Ordu,  1 kmNE Gölköy, 40°43'N, 37°37'E,  1050 m, beech forest with rhododendron and  Ilexundergrowth, sifted,  13.VII.2008, leg. Assing& Schülke(cAss, MNB); 866, 10♀♀, road Ünye - Akkuş,  18 kmNE Akkuş, 40°56'N, 37°07'E,  920 m, mixed deciduous forest with dominant  Fagus, litter sifted,  15.VII.2008, leg. Assing& Schülke(cAss, MNB); 16, 1♀, Akkuş,  1400 m, leg. Heinz( MNB); 16, between Ulubeyand Gölköy,  900-1100 m,  5.-15.VII.1977, leg. Korge( MNB); 16, Ulubey,  5.VII.1977, leg. Korge( MNB);   16, 15 kmS Gölköy, Harçbeli Geçidi, 40°35'N, 37°38'E,  1610 m, beech forest with rhododendron, litter and mushrooms sifted,  22.VII.2008, leg. Assing(cAss). G i r e s u n: 1♀, ca. 30 kmS Giresun, 40°39'N, 38°27'E,  830 m, beech & hazelnut forest, sifted,  28.VII.2006, leg. Assing(cAss);  1♀, ca.  40 kmS Giresun, N Kümbet, 40°34'N, 38°26'E,  1520 m, spruce forest with rhododendron,  29.VII.2006, leg. Assing(cAss); 16, ca. 30 kmS Giresun, 40°36'N, 38°27'E,  1250 m, spruce forest with rhododendron, sifted,  29.VII.2006, leg. Schülke(cAss). G ü m ü ş h a n e: 1♀, ca.  50 kmSW Trabzon, NE Kürtün, 40°44'N, 39°13'E,  1430 m, spruce forest with rhododendron, near stream, sifted,  27.VII.2007, leg. Schülke( MNB). Tr a b z o n: 16 [slightly teneral], ca.  50 kmS Trabzon, 20 kmS Maçka, Altındere Milli Park, 40°40'N, 39°40'E,  1560 m, spruce forest with rhododendron, sifted,  26.VII.2006, leg. Assing(cAss). C o m m e n t: The original description of  Q. boluensisis based on a male holotypefrom "Abant-Gebirbe bei Bolu" and two female paratypesfrom "Bolu-Gebirge bei Mengen" ( KORGE 1971), that of  Q. deuveion a male holotypeand a female paratypefrom "Drannaz Dag" ( COIFFAIT 1978a). The latter was synonymized with the former by SOLODOVNIKOV (2004). SOLODOVNIKOV (2004, 2005) provided additional records from Bolu, Zonguldak, Kastamonu, Sinop, Ordu, Gümüşhane, and Trabzonprovinces. He also reported one femalefrom Borçka in Artvin province, but this record is zoogeographically implausible and most likely refers to  Q. smetanaior  Q. walteri. The confirmed distribution of  Q. boluensisis confined to the Pontic Mountains from Boluin the west to Trabzonin the east ( Map 1) and parapatrically borders on that of  Q. smetanai. In one locality in Trabzon(Altındere Milli Park), one maleof  Q. boluensiswas collected together with two malesand one femaleof  Q. smetanai(see grey circle in Map 1). For characters distinguishing  Q. boluensisand  Q. smetanaisee the comment in the section on  Q. smetanai. Size, coloration, and other external characters of  Q. boluensisare subject to remarkable intraspecific variation. The pronotum may be uniformly reddish, reddish with a diffuse and oblong darker spot on either side of the midline, or blackish-brown with reddish margins. The colour of the abdomen ranges from reddish to blackish-brown with narrowly reddish posterior margins. The palisade fringe at the posterior margin of tergite VII may be complete (though very narrow), reduced to indistinct rudiments, or completely absent. Considering that, like the typematerial (see also KORGE 1971), all the examined specimens are micropterous and have a blackish head, it appears noteworthy that SOLODOVNIKOV (2004)states on p. 229 that "in  Q. boluensishead and pronotum [are] never black" and that SOLODOVNIKOV (2005: 7)seems to suggest that the specimens examined by him earlier are not micropterous: "Unlike other examined material on  Q. boluensis..., these two specimens... have shortened wings". 2010-03-29 Assing Turkey 930 40.666668 Bolu 1161 31.633333 3 1140 1 Bolu 2012-04-29 Brachat & Meybohm Turkey 1060 31.65 Yigilca 3 1140 1 Bolu 2012-04-30 Brachat & Meybohm Turkey 1080 31.633333 Bolu 3 1140 1 1 Bolu 1976-04-29 1976-05-20 1976-04-29 V, MNB Korge Turkey Abant mountains 3 1140 1 Bolu 1972-07-15 2010-03-26 1972-07-15 MNB Heinz & Assing & Cambasi & Korge Turkey 1100 41.233334 Yigilca 1158 31.866667 3 1140 3 3 Bolu 1970-07-25 2009-03-31 1970-07-25 MNB Assing Turkey 1500 41.733334 Lala 1154 35.016666 Catalzeytin 3 1140 2 2 Sinop 2009-04-01 Assing Turkey 820 41.783333 Cangal Dagi 1154 34.666668 3 1140 1 1 Sinop 2009-04-01 Assing Turkey 920 41.816666 Cangal Dagi 1153 34.65 3 1140 1 1 Sinop 1970-07-26 MNB S Ayancik & Heinz Turkey 1100 Cangal Dagi 3 1140 1 Sinop 1970-07-27 2008-07-20 1970-07-27 MNB Heinz & NE Asarcik & Assing & Ilex & Schulke Turkey 1250 40.716667 Carpinus forest 1161 37.616665 Dranaz Dagi 3 1140 2 2 Sinop 2008-07-15 MNB Fagus & Assing & Schulke Turkey 920 40.933334 Unye - Akkus 1159 37.116665 3 1140 10 10 Sinop [437,845,694,721] MNB Heinz Turkey 1400 Akkus 3 1140 1 1 Sinop 1977-07-05 1977-07-15 1977-07-05 MNB Korge Turkey 1000 Golkoy Ulubey 3 1140 1 Sinop 1977-07-05 MNB Korge Turkey Ulubey 3 1140 1 Sinop 2006-07-28 2008-07-22 2006-07-28 Assing Turkey 1610 40.65 Harcbeli Gecidi 1161 38.45 3 1140 1 1 Giresun 2006-07-29 Assing Turkey 1520 40.566666 Kumbet 1162 38.433334 3 1140 1 1 Giresun 2006-07-26 2007-07-27 2006-07-26 MNB, T Schulke & Assing Turkey Kurtun 1250 40.666668 Altindere Milli Park 1161 39.666668 Macka 3 1140 1 1 Giresun