Protubera beijingensis sp. nov. (Protophallaceae, Phallales) from China Li, Guojie Deng, Dejiang Wei, Jinkang Zhang, Chulong Zhao, Ruilin Lin, Fucheng Phytotaxa 2018 2018-04-27 348 2 133 140 9ZDTM G.J. Li & R.L. Zhao G. J. Li & R. L. Zhao 2018 [136,631,1684,1711] Agaricomycetes Phallogastraceae Protubera Fungi Hysterangiales 2 135 Basidiomycota species beijingensis sp. nov.   Etymology—refers to the locality of the typespecimen.    Holotype— China, Beijing City, Huairou District, Labagoumen Manchu Village, Labagoumen Natural Reserve, N 40°54′ E 116°36′, elv.  1200 m.,  19 September 2017, En-Yong Zhao, ZRL20171200 ( holotype HMAS! 279558). Short diagnosis—Basidiocarps gasteroid, 40–50 mmin diam., pale brown intermixed with light green. Peridium bistratal, outer layer composed of repent and interwoven hyphae, 0.5–1.0 mm thick. Glebal tissues divided into elongated plates arranged centripetally near center. Pale green, gelatinous matrix present in center. Basidiospores 3.4–5.2 × 1.5–2.4 μm. Temperate species.   FIGURE 1.  Protuberaphylogeny generated by the Bayesian analysis of the atp6, nLSU, and rpb2sequences. Bayesian posterior probability (PP) values and bootstrap support (BS) values>50% are given at the internodes (PP/BS).  Description—Basidiocarpgasteroid, with rhizomorph, spherical to irregularly shaped, 40–50 mmin diam., pale brownish tinged, intermixed with pale green color, first Ochraceous-Orange (XV13′m) to Old Gold (XVI19′i), turning Sarccardo’s Olive (XVI19′m), Cinnamon-Brown (XV15′k) to Prout’s Brown (XV15′m) when mature, smooth first, then wrinkled, slightly groovy towards base, cracked when mature, compressed and hard after drying. Peridiumbistratal, outer layer compact and thin, pale brown, inner layer rigid, white, 0.5–1.0 mm thick. Glebal tissues composed of centripetally arranged plates extending inward from inner peridium to center, pale green tinge of Mignonette Green (XXXI25′i), Asphodel Green (XLI29′′′) to Deep Grape Green (XII25′′′i), immersed in hyaline to translucent gelatinous matrix radially arranged from center. Rhizomorphssimple, rarely branched, same color as peridium, 20 mmlong. Odornot distinct. Tastemild. Spore printpale brownish green tinged of Olive-Yellow (XXX23′′).  Basidiospores[50/1/1] (3.4–) 3.7–4.4 (–5.2) × 1.5–2.4 μm, Q = (1.58–) 1.90–2.44 (–2.67), ( Q= 2.16 ± 0.23), hyaline to pale yellow, narrowly ellipsoid, rarely ellipsoid, truncate toward the base, smooth. Basidia20–33 × 3–4 μm, 8-spored, hyaline, ventricose, cylindrical to irregularly clavate. Sterigmatashort, up to 1 μm long. Peridium outer layer250–550 μm in width, composed of repent, interwoven, pale brownish to yellowish, septate, occasionally clamped hyphae, 2–4 μm in width, inner layer 400–800 μm wide, consisting of hyaline hyphae embedded in gelatinous matrix, septate, slightly thick-walled, 2–3 μm in width, crystals not observed. Gelatinous tissuecomposed of loosely interwoven, hyaline hyphae 2–3 μm wide intermixed with gelatinous matrix.  Habit and habitatsolitary on soil of broad-leaved forest dominated by  Quercus mongolicaFisch. ex Ledeb.and  Betula platyphyllaSuk. 2017-09-19 HMAS China City 1200 40.9 Labagoumen Manchu Village 1159 116.6 Huairou District 2 135 2 Beijing holotype