Walker, 1852 : 1079 Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos Choi, Jinyeong Soysouvanh, Pheophanh Lee, Seunghwan Hong, Ki-Jeong Zootaxa 2018 2018-08-17 4460 1 1 62 4TZNC (Walker, 1852) Walker 1852 [151,541,685,711] Insecta Coccidae Saissetia GBIF Animalia Hemiptera 51 52 Arthropoda species coffeae     Lecanium coffeae  Walker, 1852: 1079.   Diagnosis.Marginal setae variable in size, mostly with fimbriate apices ( Fig. 48F); 11–18 setae present between anterior and posterior stigmatic clefts on each side. Venter with multilocular disc-pores mainly present around vulvar area, plus a few pores present laterad of meta- and mesocoxa ( Fig. 48I); tubular ducts of 3 types: type I each with a broad inner ductule, present on medial submarginal area, and few ducts present around meso- and procoxa; type II each with a narrow inner ductule, present on inner submarginal area and medial area of thorax and abdomen; and type III each with a filamentous inner ductule, present on outer submarginal area ( Figs 47C, 48L) (partially adopted from Williams & Watson 1990; Choi & Lee 2017b).    Material examined. 5 ♀♀, LAOS, Kham Dist., Xiangkhoang Prov.,  2.v.2015, coll. J.Y. Choi, on  Citrussp. ( Rutaceae).   Hosts.Polyphagous. According to García Morales et al. (2016),  S. coffeaehas been recorded from plants belonging to 290 genera in 106 families.   Distribution.All zoogeographical regions;  OrientalRegion( India, Indonesia, Laos, Malaysia, Philippines,  SriLanka, Taiwan, Thailand, and Vietnam) ( Waterhouse 1993; García Morales et al. 2016).  Economic importance. Hamon & Williams (1984)and Alford (2012)considered  S. coffeaeto be a pest on a variety of ornamental plants; it was also listed as a pest of coffee in Ethiopia( Abebe 1987) and of citrus in Mediterranean countries ( Franco et al. 2006). In addition, Rosen et al. (1971)and Ricalde et al. (2015)noted that  S. coffeaeis one of the main pests on olives (  Olea europaea) in Israeland Brazil.   Remarks.  Saissetia coffeaediffers from other  Saissetiaspecies in having 3 typesof ventral tubular ducts in the submarginal areas. It is similar to  S. bobuaeTakahashiin having ventral tubular ducts with broad inner ductules, but differs in having the following combination of character states (character states of  S. bobuaein parenthesis): (i) dorsal tubercles present (absent), (ii) marginal setae mostly with fimbriate apices (with pointed apices), and (iii) antenna 8 segmented (7 segmented) ( Tao et al. 1983; Williams & Watson 1990). 1918155440 2015-05-02 J. Y. Choi & Citrus Laos Kham Dist. Xiangkhoang Prov. 51 52 5 5