Comstock, 1881 : 333 Review of the family Coccidae (Hemiptera: Coccomorpha) in Laos Choi, Jinyeong Soysouvanh, Pheophanh Lee, Seunghwan Hong, Ki-Jeong Zootaxa 2018 2018-08-17 4460 1 1 62 T3Z2 Comstock, 1881 COmstOck 1881 [151,685,151,178] Insecta Coccidae Ceroplastes GBIF Animalia Hemiptera 13 14 Arthropoda species cirripediformis     Coccus cirripediformis  Comstock, 1881: 333.   Diagnosis.Body covered with grayish white wax without a distinct dorsal horn at maturity ( Fig. 8A). Dorsum with  Ceroplastes- typepores of 4 typespresent: mono-, bi-, tri- and quadrilocular pores ( Fig. 9A); bi- or trilocular pores frequent; anal plates each with about 4 apical setae; and dorsal setae short with swollen tips, sparsely present except for clear areas ( Fig. 9C). Marginal setae numbering only 2 or 3 between anterior and posterior stigmatic clefts on each side ( Fig. 9G). Stigmatic clefts shallow, each with conical or bullet-shaped stigmatic spines arranged in 2 or 3 rows ( Figs 8C, 9B). Venter with multilocular disc-pores present on all abdominal segments, with a few pores present laterad of meta- and mesocoxa ( Fig. 9D); tubular ducts each with a long filamentous inner ductule, present on submarginal area of posterior abdomen and head ( Fig. 9E); antenna 6 to 8 segmented, usually with 7 segments ( Figs 8D, 9K); and legs each with a tibio-tarsal articulatory sclerosis ( Figs 8E, 9F) (partially adopted from Hodgson & Peronti 2012).   Material examined.15 ♀♀, LAOS, Kham Dist., Xiangkhoang Prov., 3.v.2015, coll. J.Y. Choi, on  Alternanthera bettzickiana(Regel) (Amaranthaceae); 1 ♀, Paksong Dist., Champasak Prov., 11.vii.2015, coll. P.P. Soysouvanh, on  Plumeria rubraL. ( Apocynaceae).   Hosts.Polyphagous. According to García Morales et al. (2016),  C. cirripediformishas been recorded from plants belonging to 118 genera in 62 families.   Distribution.Most zoogeographical regions except for the Australian Region; in the  OrientalRegion, recorded from Indonesiaand Philippines( García Morales et al. 2016); Laos(new country record).  Economic importance. Hamon & Williams (1984)and Gill (1988)said that  C. cirripediformisis an occasional pest of citrus and diverse ornamental plants in California and Florida, and Bakr et al. (2010)considered it to be a serious pest of guava ( Psidium guajava) in Egypt.   Remarks.  Ceroplastes cirripediformisis similar to  C. sinensis DelGuercio, but can be separated from it by the following morphological differences (character states of  C. sinensisin parenthesis): (i) dorsal setae usually with swollen apices (cylindrical with blunt or pointed apices), (ii) filamentous ducts absent (present on ventral submargin), and (iii) multilocular disc-pores present on all abdominal and thoracic segments (restricted to posterior abdominal segments) (partially taken from Gimpel et al. 1974).