Conostethus
pamparum Berg, 1883: 76
Lygus (Lygus) aeruginosus Berg, 1892: 92
Carvalho and Carpintero, 1992: 88
Orthotylus pamparum : Carvalho and Drake, 1943: 523
Carvalho, 1958: 106
Carvalho and Fontes, 1973: 499
Schuh, 1995: 166
Saileria chilena Carvalho and Carpintero, 1991: 778
Conostethus pamparum
Orthotylus
O. pamparum
Orthotylini
Chileria
Co. pamparum
Chileria
Chileria pamparum ( Berg, 1883 )
Hadronemisca argentina Carvalho and Wallerstein, 1978: 523
Schuh, 1995: 117
Hadronemella argentina : Carvalho, 1984: 51
Hadronemella argentina
Hadronemella
Hadronemella
H. argentina
Hadronemella
Hadronemella Carvalho, 1984
Hadronemella
Hadronemella
H. argentina
H. argentina
Mecomma ambulans ( Fallen, 1807 ) ( Pluot-Sigwalt and Matocq, 2006 )
Araucanocoris Carvalho
Hadronema
Orthotylus species
Orthotylus
O. chilensis
O. chullan
O. kakan
O. kikin
Lopidea
Orthotylini
Description of One New Species of Chileria and Three New Species of Orthotylus, with Nomenclatural and Distributional Notes on Neotropical Orthotylinae (Heteroptera: Miridae: Orthotylini)
Forero, Dimitri
American Museum Novitates
2009
2009-03-31
3642
1
50
TZT5
(Berg)
Forero
2009
Berg
1883
[715,1013,1472,1497]
Insecta
Miridae
Chileria
Animalia
Hemiptera
20
21
Arthropoda
species
pamparum
comb. nov.
Figures 1, 8, 9
Conostethus? pamparum Berg, 1883: 76[n. sp.]. Lygus (Lygus) aeruginosus Berg, 1892: 92[n. sp.]; Carvalho and Carpintero, 1992: 88[synonym]. Orthotylus pamparum: Carvalho and Drake, 1943: 523[list, n. comb.]; Carvalho, 1958: 106[catalog]; Carvalho and Fontes, 1973: 499[redescription, male genitalia]; Schuh, 1995: 166[catalog]. Saileria chilena Carvalho and Carpintero, 1991: 778[n. sp.]. NEW SYNONYM.
DIAGNOSIS: Recognized by its relatively small size (fig. 1); green-spotted dorsum (fig. 1); anteriorly directed ramus of vesica with a median dorsal flap, apex with two portions of subequal length (fig. 8); left (medial) ramus of posteriorly directed rami of vesica with a median, dorsally serrate flap, apex curved dorsally; right (lateral) ramus of posteriorly directed rami apically bifid, with medial portion expanded toward the apex; apical ramus of vesica with dorsal portion small and rounded, ventral portion simple, not serrate, about twice as long as dorsal portion; genital capsule with a single anterior tergal process on the right margin of aperture, close to dorsal margin, transversely elongate and strongly serrate; tergal processes on left side of genital capsule small, anterior one strongly curved laterally, hooklike, posterior one shorter and slightly curved laterally. Chileria pamparumis easily distinguished from other species of Chileriaby its relatively small size (fig. 1) and particular structure of the vesica. Chileria pamparumis similar to Ch. araucanain having only one tergal process on the right margin of the aperture of the genital capsule (figs. 7, 8). In Ch. pamparumthe process is transversely elongate and located anteriorly (fig. 8), whereas in Ch. araucanait is knoblike and located posteriorly (fig. 7). The right tergal process of Ch. pamparumis similar to the anterior tergal process on the right margin of the aperture of Ch. andina(fig. 6), but in the latter the process is less heavily sclerotized and not strongly serrated as in Ch. pamparum(fig. 6). In addition, the right margin of the aperture of the genital capsule of Ch. andinahas two processes (fig. 6), not one as in Ch. pamparum(fig. 8).
DISTRIBUTION: Ch. pamparum, widespread in Argentina( Carpintero and Carvalho, 1993; Melo et al., 2004), is also present in Chile, and here newly recorded from Bolivia(fig. 9). Carvalho and Fontes (1973)recorded Ch. pamparumfrom Paraguay. I have not examined specimens from that country.
HOSTS: Solanum tuberosumL. ( Solanaceae), Medicago sativaL. ( Fabaceae) ( Carpintero and Carvalho, 1993), and an unidentified Malvaceae. Both records given by Carpintero and Carvalho (1993)—potato and alfalfa—are not likely to be natural hosts of Ch. pamparum. Accurate host-plant documentation is needed for Ch. pamparum.
DISCUSSION: Saileria chilenawas described from two femalesfrom Coquimbo, Chile, without discussing its generic placement ( Carvalho and Carpintero 1991). I was able to associate males and females that agree in size, color pattern, and shape of head and pronotum with the figure provided by Carvalho and Carpintero (1991: 778, fig. 46), and from the same area where S. chilenawas described, allowing for examination of male genitalic structures. Male genitalia indicate that what Carvalho and Carpintero (1991)designated as ‘‘ S. chilena’’ is, in fact, the same as Ch. pamparum(e.g., Carvalho and Fontes, 1973: 500[as Orthotylus pamparum], their figs. 15–17). Saileria bella( Van Duzee, 1916), the type species of Saileria, has a simple sclerotized vesical spicule (fig. 20). Other Saileriaspeciesalso have simple vesical spicules (e.g., S. irrorata, S. mexicana, S. compsus, Carvalho, 1985; Henry, 1980, 1985). In the examined males of ‘‘ S. chilena’’, the vesica is heavily sclerotized ( Ch. pamparum, fig. 8). Furthermore, Saileriaincludes species with delicate hemelytra and eyes removed from the anterior margin of the pronotum ( Hsiao, 1945) (see Saileria bellabelow, fig. 2), which is not the situation in ‘‘ S. chilena’’ ( Ch. pamparum, fig. 1). The hemelytral coloration of Ch. pamparummay vary from nearly uniform bright or pale green, to green-spotted. All species of Saileriaalso have a greenspotted dorsum (e.g., S. bella, fig. 2), which is probably why Carvalho and Carpintero (1991)described their species in Saileria. The habitus illustration of the female holotypeof S. chilena( Carvalho and Carpintero, 1991: 778)is consistent with the hemelytral color pattern of males and females of Ch. pamparumfrom Argentina, Bolivia, and Chile. After examining large series of specimens across the distributional range of this species, I consider S. chilenaa junior synonym of Ch. pamparum. Saileriaincludes 12 species ( Schuh, 1995; Carvalho and Costa, 1994). Some of these species may well not be in Saileria(e.g., S. fluminensisCarvalho, 1990, probably in the Orthotylusspeciesgroup; and S. sulina Carvalho, 1989, in the Zanchiusspeciesgroup, but not congeneric with Saileria) following the concept of Hsiao (1945), but this issue needs further study.
Conostethus pamparumwas transferred to Orthotylusby Carvalho and Drake (1943). Carvalho and Fontes (1973)provided drawings of the parameres and vesica for O. pamparum, but some of the structures were misinterpreted (e.g., the vesica) or not illustrated (e.g., the genital capsule). Detailed drawings of the male genitalia (fig. 8) are provided in this paper to facilitate its recognition and for future analysis of additional Neotropical Orthotylini. The shape of the left and right parameres, the presence of a dorsal left protuberance on the genital capsule, and the particular structure of the vesica, are also shared with other species of Chileria(figs. 5– 7). Therefore, I regard Co. pamparumas belonging to Chileria, with the consequent new combination Chileria pamparum( Berg, 1883).
HOLOTYPE( MALE): [ ARGENTINA]: Buenos Aires: Chascomús[ 35.5667 °S 58.0167 °W], Spegazzinicol., ‘‘ Typus’’ male, MLP No.1615, 13 ( MLP). HOLOTYPEOF SYNONYM: [ CHILE]: Coquimbo[ – Region IV: Elqui Province]: S. Vicuña, Oct 1983, L. E. Peña, Holotypus Saileria chilenadet. Carvalho, MLP No.2637, Col. Carv.no. 238 ( MLP). OTHER SPECIMENS EXAMINED: ARGEN- TINA: Buenos Aires: La Colina, 37.33333 °S 61.53333 °W, 12 Sep 1938, C. J. Drake, 13 ( AMNH_ PBI 00190677), 1♀( AMNH_ PBI 00190682) ( USNM). Catamarca: Arroyo El Pintado, 28.05 °S 65.58333 °W, 1968, L. E. Peña, 2♀( AMNH_ PBI 00190680, AMNH_ PBI 00190681) ( USNM). Chaco: Pa[mpa] d’[el] Infierno, 26.51911 °S 61.16075 °W, 27 Aug 1988, L. E. Peña, 63 ( AMNH_ PBI 00190666– AMNH_ PBI 00190671) ( USNM). La Rioja: Guandacol, 42 kmSW of Villa Union, 29.5674 °S 68.5071 °W, 28 Nov 1993, J. G. Rozen, 2♀( AMNH_ PBI 00180623, AMNH_ PBI 00180624) ( AMNH). LosRobles, 28.41666 °S 67.11667 °W, 30 Nov 1983, L. E. Peña, 163 ( AMNH_ PBI 00191008– AMNH_ PBI 00191023), 38♀( AMNH_ PBI 00191024– AMNH_ PBI 00191061) ( USNM). San Blas, 28.4 °S 67.08333 °W, 12 Nov 1991, L. E. Peña, 13 ( AMNH_ PBI 00107591), 2♀( AMNH_PBI 00107592, AMNH_ PBI 00107593) ( USNM). Santa Clara, 29.55 °S 68.5167 °W, 07 Nov 1991, L. E. Peña, 63 ( AMNH_ PBI 00107586– AMNH_ PBI 00107590, AMNH_ PBI 00190674), 2♀( AMNH_ PBI 00107594, AMNH_ PBI 00107595) ( USNM). Mendoza: Cochico, 37.26666 °S 67.38333 °W, 09 Sep 1987, L. E. Peña, 23 ( AMNH_ PBI 00190675, AMNH_ PBI 00190676), 2♀( AMNH_ PBI 00190678, AMNH_ PBI 00190679) ( USNM). N of San Rafael, 34.49031 °S 68.55632 °W, 06 Dec 1983, L. E. Peña, 13 ( AMNH_ PBI 00190967) ( USNM). San Rafael, 34.6 °S 68.3333 °W, 06 Dec 1983, L. E. Peña, 13 ( AMNH_ PBI 00190672), 1♀( AMNH_ PBI 00190673) ( USNM). Neuquen:EL Huecu, 37.6167 °S 70.6 °W, 04 Feb 1991, L. E. Peña, 123 ( AMNH_ PBI 00106739– AMNH_ PBI 00106750), 7♀( AMNH_ PBI 00106751– AMNH_ PBI 00106757) ( USNM); 16 Feb 1995– 17 Feb 1995, L. E. Peña, 63 ( AMNH_ PBI 00190698– AMNH_ PBI 00190703), 10♀( AMNH_ PBI 00190704– AMNH_ PBI 00190713) ( USNM). Rio Agrio, N of Zapala, 38.5208 °S 70.19227 °W, 09 Dec 1983, L. E. Peña, 33 ( AMNH_ PBI 00190968– AMNH_ PBI 00190970), 2♀( AMNH_ PBI 00190993, AMNH_ PBI 00190994) ( USNM). Salta: 6 kmSW of Pichanal, 23.3549 °S 64.2584 °W, 09 Nov 1993, J. G. and B. L. Rozen, 13 ( AMNH_ PBI 00180627) ( AMNH). San Juan: El Huaco, 30.15 °S 68.51667 °W, 22 Jan 1995, L. E. Peña, 23 ( AMNH_ PBI 00107605, AMNH_ PBI 00107606) ( USNM). Santiago del Estero: Beltran, 28.58333 °S 64.41667 °W, 28 Feb 1992, L. E. Peña, 1♀( AMNH_ PBI 00190488) ( USNM). Tucuman: 35 kmSE of Quilmes, 26.68 °S 65.81348 °W, 2958 m, 01 Mar 2006, T. Henryand D. Forero, Malvaceae, 93 ( AMNH_ PBI 00195176– AMNH_ PBI 00195184), 10♀( AMNH_ PBI 00195185– AMNH_ PBI 00195194) ( AMNH). Cadillal, 26.6833 °S 65.2666 °W, 07 Nov 1991, L. E. Peña, 63 ( AMNH_ PBI 00190683– AMNH_ PBI 00190688), 9♀( AMNH_ PBI 00190689– AMNH_ PBI 00190697) ( USNM). BOLIVIA: Santa Cruz: Comarapa, 17.9 °S 64.48333 °W, 1800 m, 14 Dec 1984, L. E. Peña, 23 ( AMNH_ PBI 00190648, AMNH_ PBI 00190649), 6♀( AMNH_ PBI 00107628– AMNH_ PBI 00107629, AMNH_ PBI 00190650– AMNH_ PBI 00190653) ( USNM). CHILE: Antofagasta– Region II: Antofagasta: N of Paposo, 24.964 °S 70.451 °W, 30 Oct 1983, L. E. Peña, 2♀( AMNH_ PBI 00190995, AMNH_ PBI 00190996) ( USNM). Atacama– Region III: Pinte, W [sic] of Vallenar, 28.98335 °S 70.27644 °W, 25 Oct 1980, L. E. Peña, 13 ( AMNH_ PBI 00190962), 1♀( AMNH_ PBI 00190976) ( USNM). Bio-Bio– RegionVIII: Nuble: Las Trancas, Chillan, 36.92136 °S 71.49416 °W, 17 Mar 1983– 20 Mar 1983, L. E. Peña, 23 ( AMNH_ PBI 00190965, AMNH_ PBI 00190966), 1♀( AMNH_ PBI 00190979) ( USNM). Shangri-La, 36.98099 °S 71.50932 °W, Mar 1984, D. Veas, 13 ( AMNH_ PBI 00190963) ( USNM). Coquimbo– Region IV: Choapa Province:N of los Vilos, 31.89036 °S 71.48757 °W, 24 Sep 1982, N.Z.E., 13 ( AMNH_ PBI 00190964), 2♀( AMNH_ PBI 00190977, AMNH_ PBI 00190978) ( USNM). Elqui Province: 36 kmS of Coquimbo, 30.24979 °S 71.3877 °W, 04 Oct 1994, Rozen, Quinter, Ascher, 23 ( AMNH_ PBI 00180625, AMNH_ PBI 00180626) ( AMNH). El Pangue, 30.2 °S 70.65 °W, 1900 m, 20 Oct 1991, L. E. Peña, 1♀( AMNH_ PBI 00190980) ( USNM). NE of Choros Bajos, 29.24628 °S 71.2382 °W, 20 Oct 1983, L. E. Peña, 53 ( AMNH_ PBI 00190971– AMNH_ PBI 00190975), 11♀( AMNH_ PBI 00190997– AMNH_ PBI 00191007) ( USNM). Vicuña, 30.03194 °S 70.70806 °W, 17 Oct 1991, L. E. Peña, 12♀( AMNH_ PBI 00190981– AMNH_ PBI 00190992) ( USNM). 14 kmN of Los Vilos, 31.7801 °S 71.48934 °W, 150 m, 06 Nov 1981, R. T. Schuhand N. I. Platnick, 2♀( AMNH_ PBI 00180628, AMNH_ PBI 00180629) ( AMNH). Hadronemella argentina(Carvalho and Wallerstein) Figures 1, 10
Hadronemisca argentina Carvalho and Wallerstein, 1978: 523[n. sp.]; Schuh, 1995: 117[catalog]. Hadronemella argentina: Carvalho, 1984: 51[new combination].
DIAGNOSIS: Recognized by the black dorsal coloration with claval suture pale or whitish, and a pale spot on the anterior portion of cuneus (fig. 1); male genitalia with a single, simple spicule ( Carvalho and Wallerstein, 1978: fig. 2).
Hadronemella argentinacan be recognized among the species included in Hadronemellaby the coloration and the simple structure of the male genitalia. Other species of Hadronemellafrom Argentinaare dark, but not as dark as H. argentina(personal obs.). The vesica, with a simple spicule ( Carvalho and Wallerstein, 1978: fig. 2), is shared with just a few species of Hadronemella(see Carvalho, 1984; Carvalho and Carpintero, 1991; Carvalho and Costa, 1992b).
DESCRIPTION: FEMALE: STRUCTURE: Genitalia(fig. 10): Subgenital plate longer than wide, apex rounded; base of ovipositor located anteriorly to longitudinal midpoint of abdomen; posterior wall with interramal sclerite triangular, well sclerotized, apex strongly curved medially; dorsal margin of posterior wall with scattered microtrichia; dorsal lobe of interramal sclerite elongate, gently curved medially, narrowing apically, covered with very small microtrichia, arranged in a scalelike pattern; medial sclerite indistinct; prolongation of medial margins of second gonapophyses produced as a rounded anterior sclerotized process ventral to interramal sclerite (fig. 10, arrow); dorsal labiate plate not sclerotized; sclerotized rings elongate, subrectangular, anterior portion with microtrichia, lateral recurved portion well sclerotized; ventral labiate plate not sclerotized, without conspicuous tubercles or processes; anterior wall with inner margins of first gonapophyses symmetrical, not sclerotized.
DISTRIBUTION: Hadronemella argentinawas originally described from San Luis Province( Carvalho and Wallerstein, 1978). Carpintero and Carvalho (1993)listed this species additionally from Buenos Aires Provincewithout specific localities. The distribution is extended in this paper to Catamarca.
HOSTS: No host plants have been documented yet for this species.
DISCUSSION: Schuh (1995)listed this taxon under HadronemiscaCarvalho, 1973, although Carvalho (1984)had transferred it to Hadronemella. I propose to include Hadronemellain the Orthotylusspeciesgroup of Schuh (1974).
Hadronemella Carvalho, 1984, is an illdefined genus. All of the species included ( Schuh, 1995, 2006a) are morphologically diverse (see Carvalho, 1984; Carvalho and Carpintero, 1991; Carvalho and Costa, 1992b), in particular regarding male genitalic structure. Hadronemellais probably not monophyletic, an issue that a revision of the genus should address. Nonetheless, species of Hadronemellaare not common in collections, and series of males and associated females are rare. Because of this paucity of material and because females have not been illustrated to date, I decided to illustrate the female of H. argentinagiven the abundance of specimens associated with males from a single locality. The female of H. argentinahas symmetrical medial margins on the first gonapophyses. Such condition is also found in Mecomma ambulans( Fallen, 1807) ( Pluot-Sigwalt and Matocq, 2006), some species of AraucanocorisCarvalho(personal obs.), and in members of the Hadronemagroup ( Forero, 2008). In other members of the Orthotylusspeciesgroup, like some species of Orthotylus(e.g., O. chilensis, O. chullan, O. kakan, O. kikin, fig. 16) and species of Lopidea( Davis, 1955; personal obs.), the medial margins of the first gonapophyses are highly asymmetrical. It is not known if the symmetrical condition is a symplesiomorphic character in Orthotylini, which extent in the nominotypical tribe has been poorly documented.
SPECIMENS EXAMINED: ARGENTINA: Catamarca: Punta Balasto, 26.96666 °S 66.13333 °W, 2155 m, 10 Feb 1983, L. E. Peña, 33 ( AMNH_ PBI 00190577– AMNH_ PBI 00190578, AMNH_ PBI 00190583), 4♀( AMNH_ PBI 00190641– AMNH_ PBI 00190644) ( AMNH); 173 ( AMNH_ PBI 00190564– AMNH_ PBI 00190576, AMNH_ PBI 00190579– AMNH_ PBI 00190582), 57♀( AMNH_ PBI 00190584– AMNH_ PBI 00190640) ( USNM).
MALE, MLP
Spegazzini & Typus' & No.
Argentina
-35.5667
Chascomus
6
-58.0167
22
23
1
Ciudad Autonoma de Buenos Aires
holotype
1983-10
IV
S. Vicuna & L. E. Pena
Chile
Elqui Province
Region
22
23
1
Coquimbo
holotype
MLP
Chile
Carvalho
22
23
1
Coquimbo
holotype
MLP
No. & Col. Carv.
Chile
22
23
1
Coquimbo
holotype
1938-09-12
AMNH
C. J. Drake
-37.33333
La Colina
1
-61.53333
22
23
PBI 00190677
1
Ciudad Autonoma de Buenos Aires
AMNH, USNM
22
23
PBI 00190682
1
1
Ciudad Autonoma de Buenos Aires
AMNH, USNM
L. E. Pena
-28.05
Arroyo El Pintado
555
-65.58333
22
23
PBI 00190680, PBI 00190681
2
2
Catamarca
1988-08-27
AMNH, USNM
L. E. Pena
-26.51911
Infierno
1
-61.16075
Pa
22
23
PBI 00190666, PBI 00190671
1
Chaco
1993-11-28
AMNH
J. G. Rozen
-29.5674
Guandacol
6
-68.5071
22
23
PBI 00180623, PBI 00180624
2
2
La Rioja
1983-11-30
AMNH
L. E. Pena
-28.41666
Los
1
-67.11667
22
23
PBI 00191008, PBI 00191023
1
La Rioja
AMNH, USNM
22
23
PBI 00191024, PBI 00191061
38
38
La Rioja
1991-11-12
AMNH
L. E. Pena
-28.4
San Blas
5555
-67.08333
22
23
PBI 00107591
1
La Rioja
[445,654,1727,1755]
AMNH
22
23
2
2
La Rioja
[830,1224,211,235]
AMNH, USNM
22
23
PBI 00107593
1
1991-11-07
AMNH
L. E. Pena
-29.55
Santa Clara
555
-68.5167
22
23
PBI 00107586, PBI 00107590, PBI 00190674
1
AMNH, USNM
22
23
PBI 00107594, PBI 00107595
2
2
1987-09-09
AMNH
L. E. Pena
-37.26666
Cochico
1
-67.38333
Mendoza
22
23
PBI 00190675, PBI 00190676
1
AMNH, USNM
22
23
PBI 00190678, PBI 00190679
2
2
1983-12-06
AMNH, USNM
L. E. Pena
-34.49031
San Rafael
1
-68.55632
22
23
PBI 00190967
1
1983-12-06
AMNH
L. E. Pena
-34.6
San Rafael
5555
-68.3333
22
23
PBI 00190672
1
AMNH, USNM
22
23
PBI 00190673
1
1
1991-02-04
AMNH
L. E. Pena
-37.6167
Huecu
4400
-70.6
Neuquen
22
23
PBI 00106739, PBI 00106750
1
AMNH, USNM
22
23
PBI 00106751, PBI 00106757
7
7
1995-02-16
1995-02-17
1995-02-16
AMNH
Pena
22
23
PBI 00190698, PBI 00190703
1
AMNH, USNM
22
23
PBI 00190704, PBI 00190713
10
10
1983-12-09
AMNH
L. E. Pena
-38.5208
Rio Agrio
5
-70.19227
22
23
PBI 00190968, PBI 00190970
1
AMNH, USNM
22
23
PBI 00190993, PBI 00190994
2
2
1993-11-09
AMNH
J. G. and B. L. Rozen
-23.3549
Salta
7
-64.2584
22
23
PBI 00180627
1
1995-01-22
AMNH, USNM
L. E. Pena
-30.15
El Huaco
555
-68.51667
San Juan
22
23
PBI 00107605, PBI 00107606
1
1992-02-28
AMNH, USNM
L. E. Pena
-28.58333
Beltran
1
-64.41667
Santiago del Estero
22
23
PBI 00190488
1
1
2006-03-01
AMNH
T. Henry & D. Forero
2958
-26.68
Tucuman
555
-65.81348
22
23
PBI 00195176, PBI 00195184
1
AMNH
22
23
PBI 00195185, PBI 00195194
10
10
1991-11-07
AMNH
L. E. Pena
-26.6833
Cadillal
6
-65.2666
22
23
PBI 00190683, PBI 00190688
1
AMNH, USNM
22
23
PBI 00190689, PBI 00190697
9
9
1984-12-14
AMNH
L. E. Pena
Bolivia
1800
-17.9
Comarapa
5555
-64.48333
22
23
PBI 00190648, PBI 00190649
1
Santa Cruz
AMNH, USNM
Bolivia
22
23
PBI 00107628, PBI 00107629, PBI 00190650, PBI 00190653
6
6
Santa Cruz
[701,1023,1639,1663]
Region
Chile
Antofagasta
22
23
1
Antofagasta
1983-10-30
II, AMNH, USNM
L. E. Pena
Chile
-24.964
N of Paposo
74
-70.451
22
23
PBI 00190995, PBI 00190996
2
2
Antofagasta
[924,1117,1728,1752]
Region
Chile
Atacama
22
23
1
Atacama
1980-10-25
III, AMNH
L. E. Pena
Chile
23
24
-28.98335
Vallenar
1
-70.27644
Pinte
22
23
PBI 00190962
1
Atacama
AMNH, USNM
Chile
23
24
PBI 00190976
1
1
Atacama
1983-03-17
1983-03-20
1983-03-17
AMNH
L. E. Pena
Chile
Region
-36.92136
Las Trancas
1
-71.49416
Nuble
23
24
PBI 00190965, PBI 00190966
1
Biobio
AMNH, USNM
Chile
23
24
PBI 00190979
1
1
Biobio
1984-03
AMNH, USNM
Veas
Chile
-36.98099
1
-71.50932
23
24
PBI 00190963
1
Biobio
[363,570,476,500]
Region
Chile
Coquimbo
23
24
1
Coquimbo
1982-09-24
IV, AMNH
Chile
-31.89036
Vilos
1
-71.48757
Choapa Province
23
24
PBI 00190964
1
Coquimbo
AMNH, USNM
Chile
23
24
PBI 00190977, PBI 00190978
2
2
Coquimbo
1994-10-04
AMNH
Rozen & Quinter & Ascher
Chile
-30.24979
Elqui Province
4
-71.3877
23
24
PBI 00180625, PBI 00180626
1
Coquimbo
1991-10-20
AMNH, USNM
L. E. Pena
Chile
1900
-30.2
El Pangue
5576
-70.65
23
24
PBI 00190980
1
1
Coquimbo
1983-10-20
AMNH
L. E. Pena
Chile
-29.24628
Choros Bajos
4
-71.2382
23
24
PBI 00190971, PBI 00190975
1
Coquimbo
AMNH, USNM
Chile
23
24
PBI 00190997, PBI 00191007
11
11
Coquimbo
1991-10-17
AMNH, USNM
L. E. Pena
Chile
-30.03194
Vicuna
1
-70.70806
23
24
PBI 00190981, PBI 00190992
12
12
Coquimbo
1981-11-06
Schuh
Chile
150
-31.7801
14 km N of Los Vilos
5
-71.48934
23
24
1
Coquimbo
I, AMNH
Chile
Platnick
23
24
PBI 00180628, PBI 00180629
2
2
Coquimbo
1983-02-10
AMNH
L. E. Pena
Argentina
2155
-26.96666
Punta Balasto
1
-66.13333
24
25
PBI 00190577, PBI 00190578, PBI 00190583, PBI 00190641, PBI 00190644
4
4
Catamarca
AMNH, USNM
Argentina
24
25
PBI 00190564, PBI 00190576, PBI 00190579, PBI 00190582, PBI 00190584, PBI 00190640
57
57
Catamarca