Anomalophylla tristicula Reitter, 1887: 232
Fairmaire 1891: 197
Brenske 1897: 423
Melaserica thibetana Brenske, 1897: 421
Anomalophylla thibetana Reitter, 1902: 147
Anomalophylla moupinea
Balthasar, 1932: 43
Homaloplia discoidalis Fairmaire, 1897: 244
Taxonomic revision of the genus Anomalophylla Reitter, 1887 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)
Ahrens, Dirk
Zootaxa
2005
2005-11-04
1076
1
1
62
Reitter, 1887
Reitter
1887
[264,730,1564,1590]
Insecta
Melolonthidae
Anomalophylla
Animalia
Coleoptera
6
7
Arthropoda
species
tristicula
( Fig. 1A–C, 5)
Anomalophylla tristicula Reitter, 1887: 232; Fairmaire 1891: 197; Brenske 1897: 423. Melaserica thibetana Brenske, 1897: 421, synonym. Anomalophylla thibetana Reitter, 1902: 147, synonym. Anomalophylla moupineaab. unicolor Balthasar, 1932: 43(name not available). Homaloplia discoidalis Fairmaire, 1897: 244, syn. n.
Type material. Anomalophylla tristiculaReitter lectotypehere designated: ♂” Homaloplia tristicula n. sp.Przewalsk. As. centr. Coll. Reitter / [with additional labels of the curator:] Asia centr.: am oberen Laufe des blauen Flusses 12000’ [feet] Bythu VI 1884leg. Przewalsky / Holotypus1887 Anomalophylla tristicula” (HNHM). Melaserica thibetanaBrenske lectotypehere designated: ♂”Thibet bor. Kukunoor [Handschrift Kaszab] / coll. Reitter / Thibetana m. Kukunoor 1902 / Holotypus Anomalophylla( Xorema) thibetanaReitter[Handschrift Kaszab]” ( HNHM), Paralectotypes: 1 ♂”Kukunoor Thibet [Handschr. Reitter] / coll. Reitter / Paratypus Anomalophylla( Xorema) thibetanaReitter[Handschrift Kaszab]” ( HNHM), 1 ♂”Thibet KukuNor 3200 mF. Hauser 1898 / coll. Reitter / Paratypus Anomalophylla( Xorema) thibetanaReitter[Handschrift Kaszab]” ( HNHM), 2 ♂♂”Thibet bor. Kukunoor [Handschrift Kaszab] / coll. Reitter / Paratypus Anomalophylla( Xorema) thibetanaReitter[Handschrift Kaszab]” ( HNHM). Anomalophylla thibetanaReitter lectotypehere designated: ♂”Thibet Tatsienlou Mgt. F. Biet / Melaserica thibetanatype Brsk. / coll. Brenske” ( ZMHB). Homaloplia discoidalisFairmaire syntypes: 1 ♀”Tchou[n]gKing SzeTchouan / Type / Homaloplia discoidalisFairm. n. sp./ Museum Paris P. Guerry 1924” ( MNHN), 1 ♀”Museum Paris 1906 Coll. Leon Fairmaire / Tchou[n]gKing SzeTchouan / Type / Homaloplia discoidalisFairm. n. sp./ Museum Paris P. Guerry 1924” ( MNHN). Additional material examined.– 1 ♂“ TatsienluKiulung China Reitter/ ab. Unicolor/ Det. Dr. Balthasara. unicolorm.” ( ZSM), 5 specimens” Thibet KukuNor 3200 mF. Hauser 1898 / coll. Brenske” ( ZMHB), 2 ♂♂“ Thibet KukuNor 3200 mF. Hauser 1898 / Hauser Coll. 190463.” ( BMNH), 11 ♂♂, 3 ♀♀” Thibet KukuNor 3200 mColl. Hsr. 98” ( DEI, BMNH), 1 ♂“ Thibet KukuNor 3200 mF. Hauser1898 / Sammlung J. Daniel” ( ZSM), 4 ♂♂“ KukuNor TibetF. Hauser / Nevinson Coll. 191814.” ( BMNH), 1 ♂“ KukuNor1901 H. / SammlungJ. Daniel” ( ZSM). Gansu: 1 specimen”W. ChinaB. M. 1932203 / Kina S. Kansu/ Sven Hedins Exp. Ctr. Asien Dr. Hummel” ( BMNH), 6 specimens( ♂) ”China Gansu, Laji Shan Xiahe, 16.–20.VI.1994 3000–3600 mMer B. Zvari č lgt. ( CA, TICB), 5 specimens( ♀) “ CHINA Gansureg. Xiahe17–17.7. M. Nikodýmlgt. 1990 ( CK), 32 specimens China, Kansumer., Xiahe( Labrang), 3300–3700 m, 1.–15.vi. 1998 V. Majorleg. ( TICB), 21 specimens China, prov. Gansu, Labrang, 2900–3500 m, 30.vi.1991 A. Helialeg. ( TICB), 7 specimens China. S. Gansu Dengkagoin5.– 9.6.1996 Beneš& Štepař lgt. ( CA), 7 specimens China Gansureg. 4200 mM. Nikodýmlgt. 1990 Dogcanglhamo12–15.7. ( CN), 4 specimens China Kansu Ponggartang 30. Jun 1992M. Boklgt.” ( CA). Sichuan: 6 specimens“ Tibet Chasseursde Tatsienlou 1895” ( MNHN), 15 specimens“ Thibet Tatsienlou Mgt.F. Biet” ( MNHN), 3 specimens( ♀) “ TatsienLou Chasseurs Thibetains1896” ( MNHN), 2 specimens“ Tatsienlu Szetschwan/ 12398 / ex. Orig. Samlg.J. Breit Wien” ( NHMB), 1 specimen“ Omei Szetschwan” ( NHMB), 1 specimen( ♂) ” China7.92 Sichuan prov.Tagu” ( CA), 1 specimen( ♂) ” China, N Sichuan prov.Barkam, ca 2300 m22.– 26.6.1991 J. Kalábleg.” ( CN), 13 specimens” China. N. Sichuan Songpan17.–18.1996 Beneš& Štepař lgt. ( CA), 7 specimens China North Sichuan Songopanenv. 30.5.–3.6.95 Benes( CA), 5 specimens China, North Sichuan, Zhangiaenv. 7.–10.6.95 Benes( CA), 2 specimens( ♂) China Sichuan, Kangding(formerly Tatsienlu) 2900 m, 2.VII.1996 30°03’N 102°02’EC48 / collected by A. Smetana, J. Farkacand P. Kabátek” ( NHMB, HAHC), 3 specimens” China, Sichuan6.– 9.VII. Kandingenv., 2500–3000 m 30°05’N 101°55’E1995 M. Trýznaet O. Safráneklgt.” ( TICB), 8 specimens” China, Sichuan1997 Daliang Shan mts.27.VII. road Meigu Leibopass 15 kmNE Meigu 28°25’N 103°17’EM. Trýznaet O. Safráneklgt.” ( TICB), 9 specimens”C. China, W Sichuan Kanding, 30.vi.–8.vii.1999 V. Benešleg.” ( TICB), 14 specimens” China: W Sichuan, Maniganggoenv. cca 4000 m 31°55’N 99°10’E, 2– 7.VII.98 A. K ř í ž ova & J. Mikšovská( TICB), 2 specimensC. China, W. Sichuan Litangenv. 3.–6.vii.1999 V. Benešleg. ( TICB), 9 specimens“ CHINA SichuanXiangcheng 2700 m 29.VII.1996 29°00’N 99°46’E/ collected by J. Farkač, P. Kabátek& A. Smetana( NHMB), 2 specimens“China prov. Sichuan Maniganggovill. env. 15.–28.VII.2002 31°54’N 99°13’Elgt. M. Kalabza 3800–5000 m( TICB), 2 specimens“ Szetschwan Sunpanting Exp. Stötzner( ZSM). Yunnan: 4 specimens( ♂, ♀) ” China, N. Yunnan Lijiang 27.–28.vi.1996local collector” ( CA), 10 specimens“ China: N. Yunnan 30 kmN of Lijiang 3000 m, 3.VII.1990L.& M. Bocák lgt.” ( NHMB), 1 specimen( ♂) ” China: Yunnan, Baoshanenv. h= 2000 m, 15.vii.2000” ( CA), 4 specimens” Yunnan3200– 2700 m 27°00’N 100°12’E1993 Yulongshan mts.23–24/6. Vit Kubanleg.” ( TICB), 1 specimen” Yunnan 3600–4100 m 27°02’N 100°11’E Yulongshan mts.27/5. Vit Kubanleg. 1993“ ( TICB), 1 specimen“ CHINANYunnan 27°08’N 100°14’E Yulongshants. 2900–3500 mBeishuivill. leg. D. Kral7.–12.VII.90” ( CK), 7 specimens“ Yunnan 3000 m 27°05’N 100°15’E Yulongshan mts.Ganhaizi pass 4.7.92 David Kralleg.” ( CK), 2 specimens“ CHINANYunnan 27°06’N 100°15’EYulongshan mts. 3000–3500 mGanhaizipass leg. D. Kral18.–23.VII.90” ( CK), 1 specimen” Yunnan24–26. May Yulong Mts.1995 27°01’N 100°21’E Bolmlgt. 3200 m” ( TICB), 8 specimens” China, Yunnan, 2,5–3,8 km 27°20’N; 100°11’E Habashan Mts.SE slope, 3.–6.6. lgt. S. Becvar, 1995” ( TICB), 2 specimens“ChinaS Sichuan, 1997 Daliang Shan mts.27.VII. road MeiguLeibopass 15 kmNE Meigu 28°25’ N, 103°17’EM. Trýzna& O. Safráneklgt.” ( TICB), 1 specimen“ China, NYunnan Deqen SW env., 3000–3500 m28°28’ / 29°51’–55’ 1.7. 92 L.+R. Businskylgt.” ( NHMW). Qinghai: 14 specimens” China Qinghaireg. 2500 mvalley 40 kmS of Huangyuan6– 8.7.1990M. Nikodým lgt.” ( CN, CA, CK), 7 specimens( ♀) “ CHINA, Qinghai prov. Xinning 2300 m25.6.93 Th. Deuvelgt.” ( CK), 1 specimen“ CHINA, E Qinghai 20 kmW Tongren, h= 3400–3600 m, 2–3.VII.1999 S. Murzinleg.” ( CDKC). Tibet: 1 specimen” East Tibet Poshö 12,000 ft. VI.1936/ R. J. H. Kaulback B. M. 1937–547” ( BMNH), 1 specimen” East Tibet Du Chu Valley Poshö 12,000 ft. 7.VI.1936” ( BMNH), 5 specimens”E Tibet, Tuntala Shan mts.road MarkamZogang 40 kmE Zogang, 4500 m 29°40’N, 98°08’Epicea forest, 29.VI.–3.VII. Jaroslav Turnaleg. 1997” ( TICB), 4 specimens”E Tibet, ca. 4400 malt. Bamdaenv., grassland 30°15’N, 97°16’E, 5.VII.1997 M. Trýzna& O. Safráneklgt.” ( TICB), 1 specimen” ChinaSW Kangding5.2001 lgt. M. Häckel” ( Coll. M. Múcka), 10 specimens“SE Tibet, 24.vi.1997YanjingN env., 2600 malt. 28°51’N, 98°41’EM. Trýzna& O. Šafráneklgt.” ( TICB). Form A. tristiculaB (This form has a tubercle at the medial base of the right paramere, basal of its basal lobe, Fig. 1D–F): Yunnan: 26 ♂♂”China pr. Yunnanb. occ. Degen env. 4200 m, 8.6.1993 R. Cervenkalgt.” ( DEI), 1 ♂“ CHINA N.W. Yunna San Ba, Haba mts. 2500 m, 30.VI.1998leg. S. Murzin” ( CA), 1 ♂“ CHINA, Yunnan Zhongdian, 3200 m17.– 19.6.1995 T. Spevárleg.” ( TICB), 6 ♂♂, 3 ♀♀“CHINANW Yunnan, 3000 m, road DequenYenYing 30 kmW of Dequen, 1997 28°32’N, 98°49’E, 23.VI. M. Trýzna& O. Safráneklgt.” ( TICB), 3 ♂♂, 2 ♀♀“ChinaNW Yunnan, 3600 m, road DequenYenYing 10 kmW Dequen, 28°28’N, 98°53’E, 23.VI. Jaroslav Turna 1997” ( TICB, a few specimens showing intermediate features to typical form of A. tristicula), 1 ♂“ YUNNAN1995 25 kmE of ZHOGDIAN 3300–4000 m Bolmlgt., 12.–14.Jul” ( NHMB), 24 ♂♂“N YUNNAN 19–25.VI.1994 27°49’N 99°43’Ecca 3600 mZHONGDIAN lgt. D. Král& J. Farkač ( NHMB, CK), 3 ♂♂“China: Yunnan Prov.Na Pa Hai, 20 kmNW Zhongdian, 2002/6/16, alt. 3365 m HansBŠnzinger leg feeding on Rosa omeiensis” (CA). Lectotypedescription( Anomalophylla tristicula). Length: 6.1 mm, length of elytra: 3.9 mm, width: 3.5 mm. Body oblong, black, dorsal surface dull with long, dense, erect setae; pilosity black, setae on elytra and sometimes those on pronotum posteriorly white. Head:Labroclypeus transverse, widest medially; lateral margins strongly convex and slightly convergent anteriorly and toward base; anterior angles strongly rounded; lateral border and ocular canthus producing a distinct blunt angle; anterior and lateral margin strongly reflexed and anteriorly sinuate medially; surface almost flat medially and moderately shiny, coarsely and densely punctate, with numerous long, erect setae. Frontoclypeal suture distinct, weakly curved and slightly elevated; smooth area anterior to eye as wide as long. Ocular canthus moderately long and slender, finely and densely punctate, densely setose. Frons with coarse, dense punctures; punctures bearing a long and erect seta; basal punctation less dense. Eyes small, ratio of diameter / interocular width: 0.47. Antenna black with ten antennomeres; club with five equal in length antennomeres; club 2.5 times as long as the remaining antennomeres combined, reflexed. Prementum almost flat. Pronotum:widest adjacent to base; lateral margins in posterior half subparallel, in anterior third strongly convex and convergent; anterior angles weakly produced and moderately rounded; posterior angles blunt and moderately rounded; anterior margin medially weakly convex with distinct, fine marginal line; basal margin with a fine marginal line. Pronotal surface with dense punctation with fine glabrous punctures mixed with large punctures, bearing a long erect seta that is slightly directed anteriorly and weakly curved posteriorly. Anterior and lateral borders setose; basal margin of hypomeron not produced ventrally, not transversely sulcate anterior to base. Scutellum:moderately long; apex weakly rounded with fine, moderately dense punctures; medially smooth; minute setae present in the punctures. Elytra:Shape oblong, widest medially; striae indistinctly impressed and finely densely punctate; intervals flat with fine, moderately dense punctures; punctures bearing long, erect setae, which are longer basally. Epipleural edge fine, ending at the strongly convex external apical angle of elytra, epipleura densely setose, strongly curved in anterior third of elytra; apical border chitinous, without short microtrichomes. Venter:Ventral surface dull with fine, dense punctures. Metasternum densely setose, setae long, partially appressed, partially erect. Metacoxa glabrous adjacent to articulation of the leg; elsewhere with fine, long appressed setae. Abdominal sternites with an indistinct transverse row of coarse punctures bearing thick setae between fine, dense punctation; all abdominal sternites with fine, long setae; tegument of abdominal sternites (60x magnification) with fine polygonal mesh pattern formed by microtrichomes; ultimate three abdominal sternites at midline with longitudinally impressed line. Mesosternum between mesocoxae narrow, narrower than mesofemur with irregularly scattered, thick setae. Ratio of length of metepisternum / metacoxa: 1 / 1.45. Pygidium moderately convex, posteriorly shiny; with fine, dense punctures bearing fine, long setae; without smooth midline. Legs:slender with shiny surface. Femora with two longitudinal rows of setae, finely densely punctate and setose; anterior edge of metafemur acute, lacking adjacent serrated line; posterior margin weakly convex with a few fine setae medially, ventral posterior margin weakly widened in apical half; posterior margin smooth ventrally and dorsally. Metatibia moderately slender and long, widest at apex; ratio width / length: 1 / 3.9; dorsal margin sharply carinate and smooth with two groups of spines, the basal group at one third, apical one at two thirds of metatibial length, basally with a few single spines in punctures; lateral face longitudinally convex with dense, fine punctures (some of them longitudinally impressed); ventral edge serrated, with five thick and long, equally spaced spines; medial face finely punctate, apex sharply truncate interiorly near tarsal articulation. Tarsomeres dorsally glabrous, coarsely punctate; ventrally with sparse, short setae; metatarsomeres dorsally without longitudinal impressions, ventrally with strongly serrated ridge, laterally without strong longitudinal carina; first metatarsomere slightly shorter than the following two tarsomeres combined and one third of its length longer than the upper tibial spur. Protibia short, bidentate. Protarsal claws symmetrical, basal tooth of inner protarsal claw evenly pointed. Aedeagus: Fig. 1A–C. Intraspecific variation.Length: 5.2–6.5 mm, length of elytra: 3.1–4.2 mm, width: 2.8– 3.6 mm. In some specimens elytra uniformly reddish brown without dark lateral margins. Density of punctation on dorsal surface variable. Anomalophylla tristiculashows in the southern part of its range a considerable infraspecific variation in shape of parameres, which seems to be neither geographically nor morphologically well defined (with sympatrically occurring different intermediate stages; see form B, in additional material examined above, Fig. 1B, E). ♀: Antennal club with three antennomeres and distinctly shorter than the remaining antennomeres combined; eyes equal in size to that of male; pygidium apically less convex. FIGURE 1.(A–C) Anomalophylla tristiculaReitter(Tibet: KukuNor); (D–F) A. tristiculaReitter(China, Yunnan, Dequen env., “population B”); (G–J) A. tsangpoana sp. n.(holotype); (K–M) A. moupinea(Fairmaire)(lectotype “Chine: Moupin”); (N–P) A. subcarinata sp. n.(holotype); (Q–S) A. obscuripennis sp. n.(holotype). (A, D, G, K, N, Q) aedeagus, left side lateral view; (B, E, H, L, O, R) parameres, dorsal view; (C, F, J, M, P, S) aedeagus, right side lateral view. Scale bar = 0.5 mm. Remarks.The original description based on an uncertain number of syntypes(”Am oberen Laufe des Blauen Flusses, ByTchu genannt (12,000 ü. d. M.), im Juni 1884einige Exemplare erbeutet.” Reitter 1887: p. 232). Therefore it was necessary to designate the only available syntype( NHHM) as lectotypefor A. tristiculato stabilize the identity and taxonomy of the taxon. According to the statement of Nikolaev (1982)[...in the collection of the ZIN are preserved the types of A. tristiculaRtt.and A. kozloviMedv...”] there might be other paralectotypes(number unknown) in the ZIN, however these specimens could not be examined. The original description of Melaserica thibetanaBrenskebased on an uncertain number of syntypes. Therefore I designate the only available syntype( ZMHB) as lectotypefor Melaserica thibetana. The paralectotypes, which should be preserved in the MSNG according to Brenske (1897), were not found. Balthasars specimen of A.“ unicolor” is virtually identical to the lectotypeof A. tristicula. Homaloplia discoidalisFairmaireis known from female specimens only. At MNHN no additional male specimens could be found among the syntypeseries or from the same locality. Since it is presently impossible to distinguish female specimens without males of the same sample (or at least locality), the identity of this taxon remains unclear. So far recognized, the external morphology is identical with the females of A. tristicula, and its validity cannot be proved based on presently known morphological characters. The species considered by Miyake & Yamaya (2001)as Anomalophylla tristiculaand A. moupinea, are not conspecific with this species. According to their drawings of parameres ( Figs 6, 7), it is not clear whether these taxa even belong in the genus Anomalophylla. Diagnosis. Anomalophylla tristiculais in genital shape and habitus similar to A. moupinea Fairmaire, 1891. It is differentiated from A. moupineaby the basal lobe of right paramere directed medially ( Fig. 1B) and by the parameres with a sharp lateral carina in basal half ( Fig. 1A,C).
ZSM
China
Det. Dr. Balthasar
Reitter
Tatsienlu-Kiulung
7
8
1
1
ZMHB
Brenske
China
3200
Thibet Kuku-Nor
7
8
5
BMNH
China
3200
Thibet Kuku-Nor
7
8
2
2
DEI, BMNH
China
3200
Thibet Kuku-Nor
7
8
14
3
11
ZSM
China
3200
Hauser
Thibet Kuku-Nor
7
8
1
1
[364,1257,1203,1230]
BMNH
China
Kuku-Nor
7
8
4
4
Xizang
ZSM
China
Sammlung
Kuku-Nor
7
8
1
1
Xizang
BMNH
China
Gansu
7
8
1
Gansu
1994-06-16
1994-06-20
1994-06-16
CA, TICB, CK
Nikodym
China
3300
Xiahe
7
8
13
1
1
Gansu
1998-06-01
1998-06-15
1998-06-01
TICB
V. Major
China
3500
Labrang
Xiahe
7
8
32
Gansu
1991-06-30
TICB
A. Helia
China
3200
Labrang
7
8
21
Gansu
1996-06-09
CA
Benes & Stepa
China
Dengkagoin
7
8
7
Gansu
CN
Nikodym & Dogcanglhamo
China
4200
Gansu
7
8
7
Gansu
1992-06-30
CA, MNHN
Bok & Chasseurs
China
Ponggartang
7
8
6
Gansu
[388,1058,1644,1670]
MNHN
China
Thibet Tatsienlou Mgt.
7
8
15
Gansu
MNHN
China
Tatsien-Lou Chasseurs Thibetains
7
8
4
1
Gansu
NHMB
China
Orig. Samlg.
Tatsienlu Szetschwan
7
8
2
Gansu
NHMB
China
Omei Szetschwan
7
8
1
Gansu
[387,1033,1763,1790]
CA
China
Sichuan prov.
7
8
2
1
Sichuan
1991-06-26
CN, CA
J. Kalab & Benes & Stepa
China
2300
Songpan
7
8
15
1
Sichuan
CA
Songopan & Benes
China
8
9
North
7
8
7
Sichuan
CA
Zhangia & Benes
China
North
8
9
5
Sichuan
1996-07-02
NHMB, HAHC
A. Smetana & J. Farkac & P. Kabatek
China
2900
30.05
Tatsienlu
1224
102.03333
Kangding
8
9
3
Sichuan
TICB
M. Tryzna & O. Safranek
China
2750
30.083334
Kanding
1223
101.916664
8
9
8
Sichuan
Leibo
China
Daliang Shan mts.
8
9
1
Sichuan
TICB
Tryzna & Safranek
China
28.416666
15 km NE Meigu
1232
103.28333
8
9
9
Sichuan
1999-06-30
1999-07-08
1999-06-30
TICB
V. Benes
China
Kanding
8
9
14
Sichuan
TICB
Miksovska
China
4000
31.916666
Maniganggo
1213
99.166664
8
9
1
Sichuan
1999-07-03
1999-07-06
1999-07-03
TICB
V. Benes
China
Litang
8
9
2
Sichuan
1996-07-29
2002-07-28
1996-07-29
NHMB, TICB
J. Farka & P. Kabatek & A. Smetana
China
2700
31.9
Maniganggo
1213
99.21667
8
9
11
Sichuan
ZSM
China
Szetschwan Sunpanting Exp. Stotzner
8
9
2
Sichuan
1996-06-27
1996-06-28
1996-06-27
CA
China
Lijiang
8
9
6
1
1
Yunnan
1990-07-03
NHMB
China
3000
30 km N of Lijiang
8
9
10
Yunnan
2000-07-15
CA
China
2000
Baoshan
8
9
2
1
Yunnan
TICB
Vit Kuban
China
2700
27.0
Yulongshan mts.
1239
100.2
8
9
4
Yunnan
TICB
Vit Kuban
China
3850
27.033333
Yulongshan mts.
1239
100.183334
8
9
1
Yunnan
[366,1148,1084,1110]
Yulongshan
China
27.133333
CHINA
1238
100.23333
8
9
1
CK
D. Kral & David Kral
China
3000
27.083334
Yulongshan mts.
1239
100.25
8
9
9
Yunnan
[274,926,1204,1230]
China
27.1
CHINA
1238
100.25
8
9
1
CK, TICB
D. Kral & Bolm
China
3200
27.016666
May Yulong Mts.
1239
100.35
8
9
9
Yunnan
TICB
S. Becvar
China
Habashan Mts.
27.333334
Meigu-Leibo
1237
100.183334
Daliang Shan mts.
8
9
2
Yunnan
TICB
Tryzna & Safranek
China
28.416666
15 km NE Meigu
1232
103.28333
8
9
1
Yunnan
NHMW
China
3250
Businsky
8
9
93
1990-07-08
CN, CA, CK
China
2500
40 km S of Huangyuan
8
9
14
Qinghai
CK
Th. Deuve
China
2300
Xinning
8
9
8
1
Qinghai
1999-07-02
1999-07-03
1999-07-02
CDKC
S. Murzin
China
3500
20 km W Tongren
8
9
1
Qinghai
1936-06
BMNH
China
3658
Posho
East
8
9
1
Xizang
1936-06-07
BMNH
China
East
3658
Tuntala Shan mts.
Du Chu Valley Posho
8
9
6
Xizang
1997-07-05
TICB
Jaroslav Turna & M. Tryzna & Safranek
China
4500
30.25
Bamda
1222
97.26667
8
9
5
Xizang
1997-06-24
TICB
Kangding & M. Hackel & Coll. M. Mucka & Tryzna & Safranek
China
2600
9
10
28.85
Tibet
1230
98.683334
8
9
10
Xizang
1993-06-08
2002-06-16
1993-06-08
DEI, CA, TICB, NHMB, CK
R. Cervenka & N. W. Yunna San Ba & S. Murzin & T. Spevar & M. Tryzna & O. Safranek & D. Kral & J. Farka
China
Hans
4200
28.533333
Haba mts.
1232
98.816666
Bolm
9
10
70
5
65
Yunnan