A Revision of the Australian Funnel-web Spiders (Hexathelidae: Atracinae) Gray, Michael R. Records of the Australian Museum 2010 2010-11-24 62 3 285 392 3J78K Gray, 2010 Gray 2010 [318,574,1740,1766] Arachnida Atracidae Hadronyche GBIF Animalia Araneae 53 338 Arthropoda species monaro sp. nov.   Etymology. The specific epithet refers to the Monaro region of New South Wales. Figure 47.  Hadronyche monaro, male: ( A) sternum, labium and Types. Holotypemale:AMS KS8744, Sawpit Creek, Kosciuszko National maxilla; ( B) cephalothorax and chelicerae, lateral; ( C) spinnerets; Park, New South Wales, 28°22'S 152°49'E, J. Gold, February 1982. ( D) cheliceral groove teeth; ( E) cephalothorax and chelicerae, Paratypes(all AM). New South Wales. Males:AMS KS6647, Sawpit Creek, dorsal; ( F) palp, prolateral; ( G) leg II, prolateral. Scale lines 1 mm. Male( holotype) — Size. Carapace length 6.23, width 6.08. inferior 1, 0. Trichobothria legs I, II: tarsus 11, 8; metatarsus Abdomen length 7.24, width 5.23. — Colour. Basic colour 13, 10; tibia p6 r6, p5 r5. — Leg spination. Leg I: femur d9, pattern. — Carapace. Slightly longer than wide, cephalic patella 2 (p1 v1), tibia 25, metatarsus 23, tarsus 9. Leg II: area moderately raised. Height 2.95. Frontal width 4.67. femur d6, patella p2, tibia 7, metatarsus 13, tarsus 9. Leg III: Fovea procurved, anterior margin weakly notched. Mid- femur d2, patella 12 (p10 r2), tibia 17 (p4 r4), metatarsus dorsal cephalic setae do not reach fovea. Anterior strial 25, tarsus 13. Leg IV: femur d4 (bristle-like), patella r1, tibia setae absent. Anterolateral carapace angle with small, weak 6 (p1 r3), metatarsus 23, tarsus 21. — Abdomen. Sparsely bristles. — Eyes. Central eye region slightly raised. Eye group covered with weak bristles and hairs. Posterior lateral width 1.76. Median ocular quadrangle length 0.66, anterior spinnerets with medium length apical segment. Lengths: total width 0.80, posterior width 1.14. Diameters: AME 0.20, 2.34; basal segment 1.00, middle 0.58; apical 0.76. Apical ALE 0.33, PLE 0.20, PME 0.16. — Chelicerae. Cheliceral segment width 0.32. groove short, of medium width, margins diverging distally. Groove length 1.85, middle width 0.41. Cheliceral teeth: Distribution. East of the Snowy Mountains in the 19 central, occupying full length of groove; 9 prolateral; Eucumbene/Jindabyne region of New South Wales(Fig. 49). 9 retrolateral. — Labium. Almost as long as wide, not indented apically. Length 1.28, width 1.29. Labiosternal   Hadronyche mascordi n.sp.sigilla narrowed toward midline. Cuspules c. 226, number moderate. — Sternum. Ovoid. Length 4.08, width 3.22. Figures 49–52; Tables 15, 34Posterior sigilla oval, broad. — Palp. Tegular area wider than long. Embolus of moderate length and width; shaft straight Etymology. The species is named for Ramon Mascord— to slightly curved, weakly tapered distally. Distal part of author, photographer and student of Australian spiders. embolus weakly twisted and flange rather short. Embolus  Figure 48.  Hadronyche monaro, male ( A,D–K, AMS KS1571, RHS; B,C, AMS KS6646): ( A) palp, prolateral; ( B) bulb; ( C) embolus, distal; ( D,E) distal patella and tibia I— D, ventral, E, prolateral; ( F,G) metatarsus I— F, ventral, G, prolateral; ( H,I) distal patella and tibia II— H, ventral, I, prolateral; ( J,K) metatarsus II— J, ventral, K, prolateral. Scale lines: 0.2 mm, except A0.3 mm and J0.1 mm.   Table 14. Male morphological data—  Hadronyche monaro(n = 3).    character range mean character range mean character ratio  CL* 6.23–6.43 6.33 Mt1S 23–30 27 CW/CL 0.97  CW 6.08–6.20 6.14 Ta1S* 7–9 8 CH/CW 0.49  CH 2.95–3.08 3.02 Ti2S* 7–11 9 CFW/CL 0.74  CFW 4.65–4.67 4.66 STC2teeth 10–11 11 CHGW/L 0.21  ChGL 1.85–1.94 1.90 BulbL 2.05–2.19 2.14 SW/SL 0.84  ChGW 0.37–0.41 0.39 EmbL 1.34–1.42 1.38 LL/LW* 0.97  ChGCT 19–26 23 BulbW 0.81–0.93 0.87 PLSAPW/L 0.43  LL 1.02–1.28 1.15 EmbmidW 0.18–0.20 0.19 BulbW/EmbL 0.63  LW 1.07–1.29 1.18 PalpTibL 2.50–3.00 2.85 EmbmidW/L 0.14  CUSP* 143–226 185 PalpTibW 1.46–1.50 1.48 PalpTibW/L 0.52  SL 3.86–4.08 3.96 PalpTibS 0 — BulbL/TibL 0.75  SW 3.22–3.42 3.32 PalpPatS 0 —  PLSAPW 0.32–0.46 0.39 PalpFemS 0–3 2  PLSAPL 0.76–1.06 0.91  Fe1S 2–9 6  Pa1S 2–6 (p1–2) 5   Types.  Holotypemale: AMS KS2065, Somersby, via Gosford, New Southslightly offset from tegulum. Bulb length 2.19, width 0.93.   Wales, 33°21'S 151°17'E, L. Abra,  October 1978. Paratypes(all AM). Emboluslength 1.42, midwidth 0.18. Lengthof femur 2.86, New South Wales. Males: AMS KS13393, Kulnura,  33°13'S 151°13'E, 25 patella 1.40, tibia 3.00. Widthof tibia 1.46. Spination: femur  October 1983; AMS KS13401, Kulnura,  33°13'S 151°13'E,  September 1983, 2, patella 0, tibia 0. A few sinuous bristles on distal femur. Nelsonand Rodgers; AMS KS4102, Somersby,  33°21'S 151°17'E, 3 October — Legs.4123. Leg Femur Patella Tibia Metatarsus Tarsus Total 1 5.04 2.50 4.12 3.77 2.26 17.69 2 4.76 2.38 3.76 3.47 2.26 16.63 3 4.22 1.81 2.61 3.33 2.31 14.28 4 5.03 2.38 3.69 4.22 2.85 18.17  Tibia Iwidth 0.94. Femora I and II with dorsal spines. Leg I unmodified. Tibia II without apophysis, weakly excavated distoventrally; ventral spines grouped in proximal half. Metatarsus II weakly sinuous with a small spined midventral apophyseal swelling. Metatarsus IV slightly enlarged distally, concavely arched dorsally. Anterior coxal hairs normal. Tarsal scopulae: legs I, II weak; legs III, IV moderate to strong. Distal metatarsal scopulae: legs I–III absent to weak; leg IV moderately developed, extending to midventral Figure 49. Collection records for  Hadronyche monaro(squares) metatarsus. Tarsal claw teeth legs I, II: superior 11, 10, and  H. mascordi(circles) 1979,  R. Mascord;  AMS KS8571, Somersby, 33°21'S 151°17'E, 12 Octoberwidth 1.22. Diameters: AME 0.25, ALE 0.42, PLE 0.33, 1981, L.Abra;  AMS KS4108, Somersby, 33°21'S 151°17'E,  3 October 1979,  PME0.30. —  Chelicerae. Anterodorsalpaturon bristles K. Mascord;  AMS KS991, Somersby, 33°21'S 151°17'E,  November 1956, J.  V. Brown.  AMS KS7335, Somersby, 33°21'S 151°17'E, 12 Septemberrather short. Cheliceralgroove moderately short and wide, 1956, J. Brown. Females:  AMS KS15749, Kariong, near Gosford, 33°26'Smargins diverging distally. Groovelength 1.69, middle 151°18'E, L. Abra,  October 1984;  AMS KS14381, Kulnura, 33°13'Swidth 0.42. Cheliceralteeth: 7 central, running full length 151°13'E,  23 March 1984, M. Gray;  AMS KS14757, Kariong, 33°26'Sof groove; 8 prolateral; 6 retrolateral. —  Labium. Short— 151°18'E,  4 June 1984;  AMS KS14234, Kulnura, 33°13'S 151°13'E, 10 length 0.98, width 1.39. Labiumfused posteromedially with  April 1984, B. Thomas;  AMS KS14233, Kulnura, 33°13'S 151°13'E, 10 April 1984, B. Thomas;  AMS KS14551, Kulnura, 33°13'S 151°13'E, 21 sternum. Labiosternalsigilla reduced to a weak, transverse  March 1984, E. Thompson;  AMS KS14226, Kulnura, 33°13'S 151°13'E, line suture flanked by two small lateral sigilla. Cuspulesc.  5 April 1984;  AMS KS14547, Kariong, 33°26'S 151°18'E,  13 April 1984. 209, covering most of ventral labium, number moderate. — Sternum. Ovoid. Raised anteriorly and continuous with Other material examined. New South Wales(males): labium. Length 3.81, width 2.82. Posterior sigilla very small,  AMS KS13394, Kulnura,  25 October 1983;  AMS KS13399, ovoid to elongate. —  Palp. Tegular areaslightly wider than Kulnura,  September 1983;  AMS KS13400, Kulnura, long. Embolusshaft short, tapering, weakly curved. Distal  September 1983;  AMS KS13377, Somersby, 14 Septemberembolus weakly to moderately twisted, often with a subdistal 1983, K. Gould;  AMS KS1929, Somersby,  October 1978, L. “notch”. Embolusmoderately offset from tegulum. Bulb Abra;  AMS KS23333, Kulnura–Somersby area. length 2.10, width 0.84. Emboluslength 1.34, midwidth 0.15. Length of femur 2.70, patella 1.40, tibia 2.68. Width Diagnosis. CL 5.54–6.68 (male). Small Atracinae. Differs of tibia 1.22. Spination: femur 1; few weak sinuous bristles from other atracine species in having a short labium (LW/ on distal femur. — Legs. 4123. LL 0.64) that is fused with the sternum posteromedially Leg Femur Patella Tibia Metatarsus Tarsus Total (Figs. 50A, 51A). 1 5.11 2.38 3.81 3.03 1.97 16.30 Male( holotype) — Size. Carapace length 6.19, width 5.68. 2 4.88 2.42 3.90 2.69 1.89 15.78 3 4.10 1.91 2.95 3.03 2.13 14.12 Abdomen length 6.07, width 4.51. — Colour. Basic colour 4 5.08 2.15 4.04 3.81 2.50 17.58 pattern, dark brown to black. Abdomen with 4–5 pairs of small distinct chevron markings on lateral dorsal abdomen.  Tibia Iwidth 0.88. All femora with dorsal spines. Metatarsus — Carapace. Slightly longer than wide, strongly raised. I normal to slightly thickened proximally. Tibia II lacking Height 3.01. Frontal width 4.63. Fovea procurved. Mid- apophysis, spines centrally-proximally placed. Metatarsus dorsal cephalic setae numerous, reach fovea. Anterior strial II weakly sinuous, with a small mid-ventral swelling. setae present. Anterolateral angle of carapace with strong Coxal hairs normal. Tarsal and distal metatarsal scopulae: bristles. — Eyes. Sessile. Eye group width 1.74. Median weak to absent on legs I, II; well developed on legs III, IV. ocular quadrangle length 0.62, anterior width 0.79, posterior Tarsal claw teeth legs I, II: superior 13, 13; inferior 6, 5. Trichobothria legs I, II: tarsus 8, 9; metatarsus 12, 10; tibia p6 r6, p6 r6. — Leg spination. Leg I: femur d4, patella 0,  Figure 51.  Hadronyche mascordi, female: ( A) sternum, labium Figure 50.  Hadronyche mascordi, male: ( A) sternum, labium and and maxilla; ( B) cephalothorax and chelicerae, lateral; ( C) maxilla; ( B) cephalothorax and chelicerae, lateral; ( C) cheliceral spermathecae; ( D) spinnerets; ( E) cephalothorax and chelicerae, groove teeth; ( D) cephalothorax and chelicerae, dorsal; ( E) leg II, dorsal; ( F) leg III, prolateral, patella, tibia, metatarsus; ( G) prolateral; ( F) palp, prolateral; ( G) spinnerets. Scale lines 1 mm. cheliceral groove teeth. Scale lines 1 mm.  Figure 52.  Hadronyche mascordi, male ( A,D–K, AMS KS23333; B,C, AMS KS7335): ( A) palp, prolateral; ( B) bulb; ( C) embolus, distal; ( D,E) distal patella and tibia I— D, ventral, E, prolateral; ( F,G) metatarsus I— F, ventral, G, prolateral; ( H,I) distal patella and tibia II— H, ventral, I, prolateral; ( J,K) metatarsus II— J, ventral, K, prolateral. Scale lines: A,E,F,I, 0.3 mm, B1.0 mm, D,G,H0.2 mm and J,K0.1 mm.   Table 15. Male morphological data—  Hadronyche mascordi(n = 8).    character range mean character range mean character ratio SD  CL* 5.54–6.68 6.27 Mt1S 22–43 33 CW/CL 0.97 0.061  CW 5.60–6.26 5.96 Ta1S 9–24 14 CH/CW* 0.50 0.018  CH 2.83–3.22 3.00 Ti2S* 8–13 10 CFW/CL 0.77 0.036  CFW 4.63–5.33 4.81 STC2teeth 13–14 14 CHGW/L 0.22 0.022  ChGL 1.69–1.86 1.80 BulbL 1.95–2.13 2.06 SW/SL 0.75 0.026  ChGW 0.33–0.46 0.40 EmbL 1.25–1.39 1.32 LL/LW* 0.64 0.051  ChGCT 7–18 12 BulbW 0.84–0.95 0.89 PLSAPW/L* 0.51 0.066  LL 0.8–1.03 0.94 EmbmidW 0.12–0.16 0.14 BulbW/EmbL* 0.68 0.033  LW 1.39–1.58 1.47 PalpTibL 2.68–3.00 2.88 EmbmidW/L 0.11 0.008  CUSP 194–260 223 PalpTibW 1.22–1.41 1.31 PalpTibW/L 0.45 0.01  SL 3.73–4.35 3.95 PalpTibS 0 — BulbL/TibL 0.71 0.031  SW 2.70–3.05 2.94 PalpPatS 0 —  PLSAPW 0.33–0.41 0.37 PalpFemS 1–3 2  PLSAPL 0.60–0.87 0.75  Fe1S 2–5 4  Pa1S* 0 0 tibia 10, metatarsus 30, tarsus 11. Leg II: femur d5, patella patella p4, tibia 8 (p4 r2), metatarsus 23 (p6 r3), tarsus 13. p1, tibia 10, metatarsus 13, tarsus 15. Leg III: femur d2, Leg IV: femur 0, patella 0, tibia 3, metatarsus 21, tarsus patella p4, tibia 10 (p5 r2), metatarsus 19, tarsus 18. Leg 21. — Abdomen. Sparse cover of weak hairs and bristles. IV: femur d5, patella 0, tibia 6, metatarsus 20, tarsus 16. Posterior lateral spinnerets with short apical segment. — Abdomen. Posterior lateral spinnerets with short apical Lengths: total 2.58, basal segment 1.53, middle 0.82, apical segment. Lengths: total 1.99; basal segment 0.76, middle 0.82.Apical segment width 0.55. — Genitalia. Spermathecae 0.50, apical 0.73. Apical width 0.38. short, bluntly conical, lateral margins sloping medially; wide basally, separated by about half basal width. Length 0.75,  Female( paratypeAMS KS15749) —   Size. Carapacelength width 0.68. 7.14, width 7.04. Abdomenlength 12.50, width 8.78. —  Colour. Basiccolour pattern. —  Carapace. Almostas  Distribution. Kulnura/ Somersby region, Central Coast, Newwide as long; strongly raised, broad frontally. Height3.81; South Wales(Fig. 49). frontal width 6.05. Cephaliclength 5.44. Foveaprocurved. Mid-dorsalcephalic setae reach fovea. Anteriorstrial setae  Comments. Afemale spider from Kulnura, NSW was present. Anterolateralcarapace angle with a few bristles. collected (apparently dead) in her burrow with 14 young —  Eyes. Centraleye region slightly raised. Eyegroup width (KS14551). These“spiderlings” were remarkable because 2.11. Diameters: AME 0.22, ALE 0.46, PLE 0.34, PMEof their large size—their mean carapace length was 4.0 mm, 0.28. Interdistances: AME–AME 0.37, AME–ALE 0.20, compared with 8.0 mm for the female. ALE–PLE 0.26, PLE– PME0.14, PME– PME0.95. Medianocular quadrangle length 0.73, anterior width 0.91, posterior width 1.45. —  Chelicerae. Cheliceralgroove margins diverge distally. Groovelength 2.31, middle width 0.65. Cheliceral   Hadronyche nimoola n.sp.teeth: 22 central in a single row occupying full length of groove; 8 prolateral; 6 retrolateral. —  Labium. Short(length Figures 53–56; Tables 16, 341.09, width 1.67), fused with sternum as in male; anterior margin not indented. Cuspulesdistributed over most of  Etymology. Thespecific epithet is taken from an aboriginal ventral labium. —  Sternum. Ovoid, fused anteriorly with word for a steep slope, a reference to the caput profile of labium, surface strongly convex. Length4.62, width 3.81. this species. Posteriorsternal sigilla small, narrow and elongate: length 0.61, width 0.21. —  Palp. Spination: tibia 0 (strong bristles),   Types. Holotypemale: AMS KS867, Batemans Bay, New South Wales, 35°43'S 150°13'E, J. R. Mylott,  3 March 1973. Paratypes(all AM). New  tarsus 6. Tarsalclaw with 8 teeth. Trichobothria: tibia p5 r6, South Wales Males: AMS KS2276 Benandarah State Forest, 35°39'S tarsus 10. — Legs. 4123. Legs I and  IVsubequal. 150°15'E,  30 November 1978–  4 January 1979, C. Horseman;  AMS KS863, Bermagui, 36°25'S 150°04'E, K. Walker;  AMS KS12124, Bondi State Leg Femur Patella Tibia Metatarsus Tarsus Total Forest, nr.Bombala, 37°06'S 149°15'E, 26 January 1981, G. Gowing et al.; 1 5.66 2.99 4.11 3.32 2.21 18.29  AMS KS2665, Bowral, 34°28'S 150°25'E,  7 February 1979, G. Goodfellow; 2 5.10 2.79 3.88 3.18 2.18 17.13  AMS KS2081, Crookwell, 34°27'S 149°28'E,  28 November 1978, L.E. 3 4.22 1.97 2.48 2.89 2.24 13.80 Willis;  AMS KS862, Narooma, 36°12'S 150°08'E,  14 February 1971, 4 5.58 2.58 3.89 3.77 2.48 18.30 B. Plunkett-Cole;  AMS KS88199, Exeter, 34°36'S 150°19'E,  21 January 2004;  AMS KS3876, Narooma, 36°13'S 150°08'E,  January 1979, B.H.  Plunkett-Cole. Females: AMS KS4664, Bundanoon, 34°39'S 150°18'E,  Tibia I width 1.20. MetatarsusI proximal width 0.86. Coxal February, 1980; AMS KS16446, Bermagui, 36°25'S 150°04'E, K. Walker; setae unmodified. Tarsalclaw teeth legs I, II: superior 10, AMS KS12139, Bondi State Forest, nr. Bombala, 37°06'S 149°15'E, 26 10; inferior 5, 5. Trichobothrialegs I, II: tarsus 11, 11;  January 1981, G. Gowing et al.; AMS KS6163, Bowral, 34°28'S 150°25'E,   October 1980, H.Styles; AMS KS6825, Grabben Gullen, 34°32'S 149°24'E,  metatarsus 13, 10; tibia p6 r6. —  Legspination. Leg I: femur  16 December 1980, R. Chown; AMS KS4506, Moruya, 35°54'S 150°06'E, 0, patella 0, tibia 1, metatarsus 6, tarsus 5. Leg  II: femur 0,  January 1980;  AMS KS3625, Narooma, 36°12'S 150°08'E,  15 June 1971, patella 0, tibia 2, metatarsus 11, tarsus 7. Leg  III: femur 0, J. Cobcroft.  Other material examined.   New South Wales(males): AMS KS2032, pattern. —  Carapace. Aboutas wide as long, strongly raised. Benandarah State Forest,  5 October 1978–  2 November 1978, C.Horseman; Height 3.61. Frontal width 5.99. Fovea broad, procurved, AMS KS867, Bateman’s Bay,  3 March 1973; AMS KS12150, Bondi State Forest, near Bombala,  26 January 1981, G. Gowing et al.; AMS KS3460, anterior margin weakly indented. Mid-dorsal cephalic Yass River,  30 July 1966, A.B.Thompson; AMS KS6163, Bowral, October setae small, closely spaced, reach, or almost reach, fovea. 1980, H. Styles; AMS KS13806, Bundanoon,  5 February 1984, W. Nooijen; Anterior strial setae absent. Anterolateral angle of carapace AMS KS6825, Grabben Gullen,  16 December 1980, R. Chown; AMSwith weak bristles. — Eyes. Sessile to slightly raised in KS7567, Grabben Gullen,  23 October 1980, R. Chown; AMS KS2117, central eye region. Eye group width 2.32. Median ocular Kioloa State Forest,  2 November 1978–  30 November 1978, C. Horseman; AMS KS2293, Kioloa State Forest,  30 November 1978–  4 January 1979, quadrangle length 0.78, anterior width 1.05, posterior C. Horseman; AMS KS1163, Mittagong,  13 January 1978, M. Johnson; width 1.60. Diameters: AME 0.33, ALE 0.47, PLE 0.37, AMS KS6275, Moss  Vale,  27 December 1980, J. Kelly; AMS KS868& PME0.25. — Chelicerae. Cheliceral groove broad, margins 869, Nerrigundah,  December 1945, V. Haskell; AMS KS861, Narooma, diverging distally. Groovelength 2.28, middle width 0.52.  16 November 1974, B. Plunkett-Cole.   New South Wales(females): AMS KS1865, Bega,  24 January 1976, H.W. Kemp; AMS KS1867, Bermagui, Cheliceral teeth: 16 central, running full length of groove;  May 1964, K.Walker; AMS KS11996, Bondi State Forest, near Bombala, 14 9 prolateral; 11 retrolateral. — Labium. Wider than long.  November 1980, G. Gowing et al.; AMS KS1872, Bowral, H.H. Florence; Length 1.44, width 1.66. Labiosternal sigilla narrowed in AMS KS1873, Bundanoon,  8 January 1952, N.C. de Meyrick; Merimbula, midline. Cuspules c. 234, number moderate. — Sternum.  5 June 1964( MV); AMS KS4777, Mittagong,  June 1937, A. Livingstone; Ovoid, broad. Setae weak. Length 4.35, width 3.65. Posterior AMS KS2961, Narooma,  29 April 1979, A. Brown.   Australian Capital Territory(male): AMS KS13824, Tidbinbilla Nature Reserve, 9 Marchsigilla elongate, ovoid. —  Palp. Tegular areawider than 1978, P. Ormay.long. Embolusrather short, strongly tapered distally, weakly curved. Distalpart moderately twisted, embolic groove   Diagnosis. CL 5.85–7.22 (male). Small atracine spiders sometimes rather wide. Embolus weakly to moderately without leg II apophyses. Differs from most  Hadronycheoffset from tegulum. Bulb length 2.43, width 0.99. Embolus spp. by caput being high and wide, almost bulbous ( CH/CW: length 1.61, midwidth 0.18. Length of femur 3.50, patella male 0.53 [ Fig. 53A], female 0.61 [Fig. 54B]), and having 1.45, tibia 3.49. Width of tibia 1.64. Spination: femur with relatively few STC teeth (7–9) on tarsi I, II; from  H. mascordi7 sinuous bristle-like spines, tibia with 2 bristle-like spines. by having an entire labiosternal sigilla; from  lamingtonensis—Legs. 4123. group species by having a long central cheliceral tooth row. Leg Femur Patella Tibia Metatarsus Tarsus Total 1 6.23 3.01 4.80 4.26 2.65 20.95  Male( holotype) — Size. Carapace length 7.01, width 7.30. 2 5.66 2.87 4.43 3.90 2.46 19.32 Abdomen length 8.44, width 6.03. — Colour. Basic colour 3 4.82 2.34 3.28 3.62 2.65 16.71 4 6.10 2.88 4.92 4.74 3.15 21.79  Tibia Iwidth 1.15. Femur I with few or no dorsal spines; femur II usually with spines, occasionally none; femora III and IV with spines. Legs I and II unmodified, spines small, scattered. Metatarsus IV thickened distally, concavely arched dorsally.Anteromedial hairs on coxae I, II usually thickened Leg Femur Patella Tibia Metatarsus Tarsus Total basally. Tarsal and distal metatarsal scopulae moderately 1 6.11 3.36 4.18 3.28 2.12 19.05 developed on legs I, II; strongly developed on legs III, IV. 2 5.45 3.20 3.62 3.03 2.26 17.56 Metatarsi IV scopulae extend to mid ventral area. Tarsal 3 4.58 2.36 2.59 2.83 2.35 14.71 claw teeth legs I, II: superior 7, 8: inferior 0, 0. Trichobothria 4 6.15 3.77 4.63 3.98 2.74 21.27 legs I, II: tarsus 11, 10, metatarsus 13, 13; tibia p6 r6, p6 r6. — Leg spination. Leg I: femur 0, patella p1, tibia 11 (p3), Tibia Iwidth 1.48. Metatarsus I proximal width 1.03. Coxae metatarsus 25, tarsus 22. Leg II: femur d5, patella p1, tibia I, II with basally thickened and thorn-like setae anteriorly. 17, metatarsus 32, tarsus 23. Leg III: femur d6, patella 33 Tarsal claw teeth legs I, II: superior 7, 7; inferior 2, 2. (p26 r7), tibia 22 (p7, r2, d2), metatarsus 40, tarsus 27. Leg Trichobothria legs I, II: tarsus 11, 11, metatarsus 12, 11; tibia IV: femur d7, patella r5, tibia 8 (r1 d2), metatarsus 22, tarsus p6 r5, p6 r5. — Leg spination. Leg I: femur 0, patella 0, tibia 35. — Abdomen. Sparse, fine hair cover, bristles lacking or 1, metatarsus 10, tarsus 7. Leg II: femur 0, patella 0, tibia 1, very weak. Posterior lateral spinnerets with moderately metatarsus 14, tarsus 7. Leg III; femur 0, patella pd 20, tibia short apical segment. Lengths: total 2.88; basal segment 12 (p6 r3), metatarsus 33 (p14 r6), tarsus 15. Leg IV: femur 1.32; middle 0.58; apical 0.98. Apical segment width 0.48. 0, patella 0, tibia 1, metatarsus 14, tarsus 7. — Abdomen.  Figure 53.  Hadronyche nimoola, male: ( A) cephalothorax Figure 54.  Hadronyche nimoola, female: ( A) sternum, labium and chelicerae, lateral; ( B) sternum, labium and maxilla; ( C) and maxilla; ( B) cephalothorax and chelicerae, lateral; ( C) cephalothorax and chelicerae, dorsal; ( D) palp, prolateral; ( E) leg spermathecae; ( D) cephalothorax and chelicerae, dorsal; ( E) II, prolateral (NB, dorsal femoral spines present or absent); ( F) cheliceral groove teeth; ( F) spinnerets; ( G) leg III, prolateral, cheliceral groove teeth; ( G) spinnerets. Scale lines 1 mm. patella, tibia, metatarsus. Scale lines 1 mm.  Figure 55.  Hadronyche nimoola, male ( A,D–K, AMS KS88199; B,C, AMS KS3876): ( A) palp, prolateral; ( B) bulb; ( C) embolus, distal; ( D,E) distal patella and tibia I— D, ventral, E, prolateral; ( F,G) metatarsus I— F, ventral, G, prolateral; ( H,I) distal patella and tibia II— H, ventral, I, prolateral; ( J,K) metatarsus II— J, ventral, K, prolateral. Scale lines: 0.2 mm, except B0.5 mm and E,G0.3 mm.   Table 16. Male morphological data—  Hadronyche nimoola(n = 17).    character range mean character range mean character ratio SD  CL* 5.85–7.22 6.62 Mt1S 6–25 17 CW/CL* 1.01 0.032  CW 6.07–7.46 6.72 Ta1S 6–22 13 CH/CW* 0.53 0.029  CH* 3.20–4.03 3.56 Ti2S* 4–18 10 CFW/CL* 0.83 0.042  CFW 4.81–6.23 5.5 STC2*teeth 7–9 8 CHGW/L 0.24 0.041  ChGL 1.97–2.56 2.17 BulbL 2.14–2.70 2.38 SW/SL* 0.88 0.034  ChGW 0.42–0.78 0.51 EmbL 1.30–1.75 1.54 LL/LW 0.85 0.053  ChGCT 9–26 17 BulbW 0.90–1.40 0.99 PLSAPW/L 0.45 0.029  LL 1.06–1.60 1.28 EmbmidW 0.12–0.18 0.15 BulbW/EmbL 0.64 0.053  LW 1.34–1.68 1.49 PalpTibL 2.65–3.77 3.21 EmbmidW/L 0.10 0.012  CUSP 160–281 219 PalpTibW 1.44–1.74 1.58 PalpTibW/L 0.49 0.031  SL 3.82–4.67 4.12 PalpTibS 0–2 <1 BulbL/TibL 0.74 0.029  SW 3.25–4.14 3.63 PalpPatS 0 —  PLSAPW 0.38–0.49 0.42 PalpFemS 0–4 1  PLSAPL 0.82–1.20 0.95  Fe1S 0–5 <1  Pa1S 0–3 <1 Posterior lateral spinnerets with short apical segment. Female( paratypeAMS KS4664) — Size. Carapace length Lengths: total 3.95; basal segment 1.79, middle 0.87, apical 8.74, width 8.65. Abdomen length 12.24, width 8.30. — 1.29. Apical segment width 0.69. — Genitalia. Spermathecae Colour. Basic colour pattern. — Carapace. About as long as short, digitiform, well separated basally (by slightly more wide, cephalic area broad and very strongly raised. Height than basal spermatheca width); diverge toward rounded and 5.08; frontal width 7.84. Cephalic length 6.90. Fovea wide, slightly enlarged apices. Length 0.87, width 0.44. deeply procurved, anterior margin indented. Mid-dorsal cephalic setae variable, may or may not reach fovea. Anterior Distribution. Southern highland and south coastal regions strial setae absent. Anterolateral carapace angle with several of New South Wales(Fig. 56). weak bristles. — Eyes. Eye region sessile. Eye group width 2.76. Diameters: AME 0.28, ALE 0.47, PLE 0.38, PME 0.24. Interdistances: AME–AME 0.55, AME–ALE 0.43, A.L.E–PLE 0.40, PLE–PME 0.26, PME–PME 1.36. Median ocular quadrangle length 0.88, anterior width 1.17, posterior width 1.88. — Chelicerae. Cheliceral groove wide, margins diverging distally. Groove length 2.83, middle width 0.74. Cheliceral teeth: 36 central, occupying full length of groove in irregular double row; 11 prolateral; 9 retrolateral. — Labium. Slightly wider than long, anterior margin weakly or not indented. Length 2.10, width 2.46. Cuspules occupy anterior two-thirds of labium. Labiosternal sigilla entire, narrow. — Sternum. Broad, subcircular. Length 5.78, width 5.08. Anterolateral angles with a few short, basally thickened, bristle-like setae. Posterior sigilla long, ovoid: length 1.31, width 0.49. — Palp. Spination: tibia 1, tarsus 8. Trichobothria: tibia p6 r5, tarsus 14. Tarsal claws with 6 teeth. — Legs. 4123. Figure 56. Collection records for  Hadronyche nimoola(squares) and  H. modesta(circles).    Hadronyche modesta(Simon)Male(AMS KS8475, Lilydale, 14 May 1980). — Size. Carapace length 6.40, width 6.10. Abdomen length 5.85, Figures 56–59; Tables 17, 34width 6.12. — Colour. Basic colour pattern. — Carapace. Slightly longer than wide, cephalic area moderately raised.  Atrax modestaSimon, 1891: 302. Hogg, 1901: 272. Main, Height 2.68, frontal width 4.16. Fovea slightly procurved. 1985: 41. Mid-dorsal cephalic setae few, do not reach fovea. Anterior  Hadronyche modesta.–Gray, 1988: 114 (transferred from strial setae absent. Anterolateral carapace angle with a  Atrax). few very weak bristles. — Eyes. Central eye region raised.  Types.  Holotypefemale: MNHN, Paris(not seen). Melbourne, Victoria. Eyegroup width 1.82. Medianocular quadrangle length  0.64, anterior width 0.86, posterior width 1.14. Diameters:  Othermaterial examined. Victoria(males): Ellinbankvia Warragul, 11 AME 0.28, ALE 0.32, PLE 0.22, PME 0.14. —  Chelicerae.  May 1962, J. Copley( MV); AMS KS8471,  Emerald,  3 April 1978; AMS  Cheliceralgroove margins subparallel to weakly divergent. KS8472,  Emerald,  20 April 1979; Leongatha,  6 June 1963, I. Bissett( MV);   AMS KS7292, Back Creek Rd., Gembrook State Forest, near Gembrook, Groovelength 2.06, middle width 0.34. Cheliceralteeth: Victoria, L. Gibson,  10 March 1981);  Mirboo North  15 April 1966, G.B. 23 central, occupying full length of groove; 8 prolateral, Burr( MV); 6 kmNE. of Mirboo North,  26.iii.1967,  J. Seebeck( MV); 9 retrolateral. Centraltooth row occasionally shorter. —  AMS KS8473, Mount Eliza,  23 April 1979;  AMS KS8930, Mount Tassie,  Labium. Widerthan long, apically indented. Length1.00, Strezelecki Ranges,  25 March 1982,  R. Waters;  AMS KS8933, Tanjil South, near Moe, February 1982,  N. Barton; AMS KS8474, Thorpdale, 10 Aprilwidth 1.38. Labiosternalsigilla narrowed toward midline. 1980; Wright, SE of Emerald,  September 1951,  C. Oke( MV); Yallourn, Slighttransverse groove behind cuspules. Cuspulesc. 123,  6 February 1963,  J. Irving( MV); Yarra Junction,  4 May 1964,  J. Mooneynumber low-moderate. —  Sternum. Ovoid. Length4.01, ( MV); Yarra Junction  20 April 1964,  J. Mooney( MV).  Victoria(females): width 3.28. Posteriorsigilla elongate, narrow. —  Palp. AMS KS10775, Cockatoo,  10 October 1982,?  McKelvey; AMS KS8477, Tegular areawider than long. Embolusshort and broad, Ferny Creek,  6 October 1979;  AMS KS12618, Hawthorn,  23 January 1983,   M. Harvey; AMS KS8476, Milford,  26 October 1976;  AMS KS8339, almost straight, base moderately offset from tegulum; distal Mirboo North,  Apr 1980;  AMS KS8932, Moe area,  October 1981,  N.Barton; embolus straight, hardly twisted. Bulblength 2.26, width AMS KS8470, Mount Evelyn, Lilydale Shire,  5 June 1979;  AMS KS8931, 1.03. Emboluslength 1.37, midwidth 0.20. Lengthof femur Newborough, near Moe,  March 1982,  N. Barton; AMS KS8926, Sassafras, 3.06, patella 1.36, tibia 3.18. Widthof tibia 1.71. Spination:  14 February 1982,  N.Wentworth; AMS KS10560, Warragul,  January 1983, N. Barton; AMS KS3386,Yinnar, 9 July 1979, N. Barton. femur 3, patella 4, tibia 0. Distal femur spined, sinuous bristles few to absent. — Legs. 4123.  Diagnosis. CL 5.48–7.30 (male). Small atracine spiders with Leg Femur Patella Tibia Metatarsus Tarsus Total leg II apophyseal swellings (Figs. 57G, 59H–K). Differs from 1 6.16 2.69 4.82 4.45 2.75 20.87  H. meridianaand  H. jensenaeby having a short, straight 2 5.91 2.61 4.82 4.20 2.75 20.29 embolus but with the distal part neither twisted nor angled, 3 5.22 2.23 3.76 4.20 2.92 17.33 respectively ( Fig. 59B,C). Differs from other  Hadronyche 4 6.32 2.72 5.24 5.44 3.48 23.20 spp. with leg II apophyses/swellings by having a relatively short labium (LL/LW 0.71) (Fig. 57E).  Figure 58.  Hadronyche modesta, female: ( A) sternum, labium and Figure 57.  Hadronyche modesta, male: ( A) cephalothorax and maxilla; ( B) cephalothorax and chelicerae, lateral; ( C) spinnerets; chelicerae, dorsal; ( B) cephalothorax and chelicerae, lateral; ( C) ( D) cheliceral groove teeth; ( E) cephalothorax and chelicerae, cheliceral groove teeth; ( D) spinnerets; ( E) sternum, labium and dorsal; ( F) spermathecae; ( G) leg III, prolateral, patella, tibia, maxilla; ( F) palp, prolateral; ( G) leg II, prolateral. Scale lines 1 mm. metatarsus. Scale lines 1 mm.  Figure 59.  Hadronyche modesta, male (KS7292): ( A) palp, prolateral; ( B) bulb; ( C) embolus, distal; ( D,E) distal patella and tibia I— D, ventral, E, prolateral; ( F,G) metatarsus I— F, ventral, G, prolateral; ( H,I) distal patella and tibia II— H, ventral, I, prolateral; ( J,K) metatarsus II— J, ventral, K, prolateral. Scale lines: 0.3 mm, except B0.5 mm and F,I,J0.2 mm.   Table 17. Male morphological data—  Hadronyche modesta(n = 26).    character range mean character range mean character ratio SD  CL* 5.48–7.30 6.24 Mt1S 20–49 36 CW/CL 0.96 0.026  CW 5.28–6.85 5.97 Ta1S* 15–41 28 CH/CW 0.44 0.025  CH 2.36–3.03 2.64 Ti2S 5–19 13 CFW/CL 0.67 0.015  CFW 3.73–4.96 4.20 STC2* 7–10 8 CHGW/L 0.17 0.011  ChGL 1.76–2.24 2.01 BulbL 1.89–2.38 2.13 SW/SL 0.81 0.027  ChGW 0.30–0.40 0.31 EmbL 1.12–1.41 1.28 LL/LW* 0.71 0.055  ChGCT 8–23 19 BulbW 0.91–1.15 1.03 PLSAPW/L* 0.51 0.053  LL 0.76–1.15 0.95 EmbmidW 0.15–0.24 0.17 BulbW/EmbL* 0.80 0.048  LW 1.13–1.52 1.37 PalpTibL 2.53–3.59 3.06 EmbmidW/L* 0.15 0.014  CUSP* 104–160 129* PalpTibW 1.39–1.83 1.55 PalpTibW/L* 0.52 0.023  SL 3.50–4.70 3.93 PalpTibS 0–5 1 BulbL/TibL 0.70 0.024  SW 2.84–3.79 3.19 PalpPatS* 3–9 5  PLSAPW 0.27–0.52 0.36 PalpFemS 3–7 5  PLSAPL 0.53–92 0.70  Fe1S 3–8 5  Pa1S 0–4 (p0) 2  Tibia Iwidth 1.14. Femora I and II with dorsal spines. Leg Length 5.08, width 4.38. Posterior sigilla elongate, length I unmodified, tibial and metatarsal ventral spines numerous. 1.02, width 0.39. — Palp. Spination: tibia 1–3 ventral bristle- Patella I and II lack prolateral spines. Ventral tibia II weakly like spines, tarsus 9. Trichobothria: tibia p5 r4, tarsus 7. swollen and spinose proximally-centrally; with a few Tarsal claw with 5 teeth. — Legs. 1423. enlarged central spines; distal tibia II concave ventrally and Leg Femur Patella Tibia Metatarsus Tarsus Total lacking spines (other than apical pair). Metatarsus II sinuous, 1 7.08 4.04 5.29 4.11 3.01 23.53 with a small, spined, midventral apophysis. Coxal setae 2 6.29 3.13 4.74 3.64 2.41 20.21 unmodified. Scopulae legs I–IV: tarsus, weak to moderate; 3 4.83 2.72 3.01 3.11 2.62 16.29 distal metatarsus, all weak. Tarsal claw teeth legs I, II: 4 6.30 3.70 4.82 4.32 3.05 22.19 superior 9, 8; inferior 0, 2. Trichobothria legs I, II: tarsus 9, 8; metatarsus 9, 9; tibia p5 r5, p6 r5. — Leg spination. Leg  Tibia Iwidth 1.40. Metatarsus I proximal width 0.88. Coxal I: femur d4, patella 1, tibia 30, metatarsus 41, tarsus 32. Leg setae unmodified. Tibia I, II with no to few spines. Tarsal II: femur d4, patella 2, tibia 12, metatarsus 32, tarsus 32. claw teeth legs I, II: superior 7, 8; inferior 0, 0. Trichobothria Leg III: femur d2 (plus strong bristles), patella p9, tibia 15 legs I, II: tarsus 10, 9; metatarsus 10, 9; tibia p6 r6, p6 r5. (p1 r4), metatarsus 24, tarsus 31. Leg IV: femur 0 (strong — Leg spination. Leg I: femur 0, patella 0, tibia 0, metatarsus bristles present), patella 0, tibia 12 (r3), metatarsus 21, tarsus 6, tarsus 12. Leg II: femur 0, patella 0, tibia 0, metatarsus 32. — Abdomen. Posterior lateral spinnerets with short apical 7, tarsus 16. Leg III: femur 0, patella pd13, tibia 5(p2 r1), segment. Lengths: total 2.01; basal segment 0.94; middle metatarsus 17(pd4 rd2) tarsus 19. Leg IV: femur 0, patella 0.40; apical 0.67. Apical segment width 0.31.  0, tibia 0, metatarsus 12, tarsus 24. — Abdomen. Sparse cover of weak hairs and bristles. Posterior lateral spinnerets Female( MV K11082), Hoddles Creek, Victoria, 37°50'Swith short apical segment. Length: total 3.16; basal segment 145°36'E, S. Johnson,  18 October 1981). —  Size. Carapace1.50, middle 0.67, apical 0.99. Apicalsegment width 0.56. length 7.48, width 7.11. Abdomenlength 10.74, width 7.21. —  Genitalia. Spermathecaedigitiform, short, apical third —  Colour. Basiccolour pattern. Anterodorsalabdominal separated from basal two thirds by a slight constriction. sigilla prominent. —  Carapace. Slightlylonger than wide, Length0.85, width. 0.41. strongly raised. Height 3.60. Frontal width 6.02. Cephalic length 5.57. Fovea strongly procurved. Mid-dorsal cephalic Distribution. Melbourne/Dandenong Range region to setae few, reach only two-thirds way to fovea. Anterior eastern Gippsland, Victoria(Fig. 56). strial setae absent. Anterolateral carapace angle with a few hairs, no bristles. — Eyes. Central eye region weakly   Hadronyche meridianaHogg raised. Eye group width 1.97. Diameters: AME 0.25, ALE 0.43, PLE 0.27, PME 0.18. Interdistances: AME–AME Figures 60–63; Tables 18, 34 0.37, AME–ALE 0.26, ALE–PLE 0.28, PLE–PME 0.30, PME–PME 0.84. Median ocular quadrangle length 0.71,  Hadronyche meridianaHogg, 1902: 122. Original descripanterior width 0.81, posterior width 1.24. — Chelicerae. tion by Hogg (1901, 274) as mis-identified male of H. Paturon with sparse anterodorsal bristle cover. Cheliceral  cerberea. groove moderately wide, margins diverging distally. Groove  Types.  Holotypemale: BMNH 1903.2.10.6 ( NHM). Mt. Macedon, Victoria, length 2.65, middle width 0.68. Cheliceralteeth: 18 central, 37°23'S 144°35'E, H. R. Hogg.  in single row occupying full length of groove, occasionally row shorter; 11 prolateral; 9 retrolateral. — Labium. Wider Other material examined. New South Wales(males): AMS KS4503, than long, apically shallowly indented. Length 1.43, width Adelong,  22 July 1980,  M. Pearce; AMS KS917, Kunama,  10 January 1971;   AMS KS916, “Loch Haven”, Wantagong, 20 June971,  R. Flynn;  AMS 2.01. Cuspulesgrouped on anterior half. Labiosternalsigilla  KS915, Wantagong,  1 April 1971,  R. Hunter;  AMS KS913, Wondalga, 23 slightly narrowed toward midline. —  Sternum. Broad, ovoid.  September 1948,  C.H. Jagoe.  New South Wales(females):  AMS KS4973, Adelong,  10 January 1980, M. Pearce;  AMS KS6053, Rosewood, 20 varies from basal area only to full length of groove. —  October 1980,  D.M.Fordham.  Victoria(males):  AMS KS8488, Baranduda,  Labium. Widerthan long, surface flattened. Length1.20,  13 October 1978, A. McDonald;  AMS KS10771, Lake Bolac,  June 1982,   H. Parnaby; Avenel,  6 September 1963( MV); Benalla, 20 October 1968 width 1.58. Labiosternalsigilla narrowed centrally. Cuspules( MV); Euroa,  19 June 1965( MV);  Pranjip, Longwood area,  4 July 1965 c. 134, number low to moderate. —  Sternum. Ovoid. Length( MV); AMS KS8484, Trawool, near Seymour,  15 August 1980,  P May4.20, width 3.24. Posterior sigilla ovoid. —  Palp. Tegular area Williams; Violet Town,  5 February 1980,  A. Kube( MV); AMS KS8485, slightly wider than long. Embolusof moderate length, rather Willowmavin,near Kilmore,  29 April 1974.   Victoria(females): Rutherglenbroad, weakly curved and tapered, with distal part twisted. Research Institute; AMS KS13651–3, Shepparton,  15 January 1984, M. Gardener; Violet Town, 5 February 1980, A. Kube (MV); AMS KS3399, Embolus base weakly to moderately offset from tegulum. Hall’s Gap, 26 March 1974, M. Gray. Bulb length 2.51, width 0.95. Embolus length 1.67, midwidth 0.30. Length of femur 3.24, patella 1.27, tibia 3.30. Width  Diagnosis. CL 5.44–7.04. Small atracine spiders with weak of tibia 1.48. Spination: femur 5, patella 1, tibia 3. Sinuous apophyseal swellings on leg II (Figs. 60F, 62H–J). Differs bristles and spines present on distal femur. — Legs. 4123. from  H. modestaand  H. jensenaeby having embolus apically Legs I, 2 subequal. twisted ( Fig. 62B,C). Differs from other  Hadronychespp. Leg Femur Patella Tibia Metatarsus Tarsus Total with leg II apophyses/swellings by having a relatively short 1 6.15 2.87 4.59 4.48 2.99 21.08 labium (LL/LW 0.78) (Fig. 60D). 2 5.82 2.81 4.84 4.43 3.05 20.95 3 5.01 2.40 3.52 4.16 3.09 18.18  Male(AMS KS8490),  Avenel, Victoria, 36°54'S 144°12'E, 4 6.03 2.68 4.89 5.09 3.35 22.04  May 1981). — Size. Carapace length 6.92, width 6.64. Abdomen length 7.17, width 5.23. — Colour. Basic colour  Tibia Iwidth 1.07. Femora I and II with dorsal spines; bristle- pattern. — Carapace. Slightly longer than wide, cephalic area like spines on femora II, IV. Metatarsus I slightly thickened moderately raised. Height 3.12. Frontal width 5.45. Fovea near middle, large ventral spines grouped here. Metatarsus straight to slightly procurved. Mid-dorsal cephalic setae do II sinuous with spined midventral apophysis. Tibia II with not reach fovea. Anterior strial setae present. Anterolateral weak, spined apophyseal swelling mid ventrally, ventral angle of carapace with weak bristles. — Eyes. Central eye spines grouped with 2–4 central spines enlarged. Tibia II region slightly raised. Eye group width 1.86. Median ocular weakly excavated distally. Anterior coxal hairs normal. quadrangle length 0.75, anterior width 0.96, posterior Tarsal scopulae: leg I weak; legs II–IV moderate to strong. width 1.24. Diameters: AME 0.32, ALE 0.46, PLE 0.43, Distal metatarsal scopulae: legs I, II weak to absent, legs III, PME 0.20. — Chelicerae. Cheliceral groove rather narrow, IV moderate to strong. Tarsal claw teeth legs I, II: superior margins subparallel. Groove length 2.35, middle width 0.38. 12, 11; inferior 2, 2. Trichobothria legs I, II: tarsus 9, 9; Cheliceral teeth: 7 central ( 6 inbasal half, 1 indistal half); metatarsus 11, 8; tibia p5 r5, p6 r5. — Leg spination. Leg I: 12 prolateral; 12 retrolateral. Central tooth row distribution femur d5, patella v1, tibia 26, metatarsus 33, tarsus 13. Leg II: femur d4, patella p1, tibia 9, metatarsus 20, tarsus 13. Leg III: femur d4 (bristle like), patella p7, metatarsus 17, tarsus 20. Leg IV: femur d5 (bristle like), patella 0, tibia 2 (d1), metatarsus 10, tarsus 19. — Abdomen. Sparsely covered with weak bristles and hairs with a pair of larger bristles above pedicel. Posterior lateral spinnerets with a rather short apical segment. Lengths: total 2.84; basal segment 1.24; middle 0.74; apical 0.86. Apical segment width 0.46.  Figure 61.  Hadronyche meridiana, female: ( A) sternum, labium and Figure 60.  Hadronyche meridiana, male: ( A) spinnerets; ( B) maxilla; ( B) cephalothorax and chelicerae, lateral; ( C) cheliceral cephalothorax and chelicerae, lateral; ( C) cheliceral groove teeth; groove teeth; ( D) spinnerets; ( E) cephalothorax and chelicerae, ( D) sternum, labium and maxilla; ( E) cephalothorax and chelicerae, dorsal; ( F) leg III, prolateral, patella, tibia, metatarsus; ( G) dorsal; ( F) leg II, prolateral; ( G) palp, prolateral. Scale lines 1 mm. spermathecae. Scale lines 1 mm.  Figure 62.  Hadronyche meridiana, male ( A,D–K, AMS KS8485; B,C, Pranjip,Victoria, [MV]): ( A) palp, prolateral; ( B) bulb; ( C) embolus, distal; ( D,E) distal patella and tibia I— D, ventral, E, prolateral; ( F,G) metatarsus I— F, ventral, G, prolateral; ( H,I) distal patella and tibia II— H, ventral, I, prolateral; ( J) metatarsus II, prolateral; ( K) femur II, prolateral. Scale lines: 0.2 mm, except B0.5 mm and D,I,J0.3 mm.   Table 18. Male morphological data—  Hadronyche meridiana(n = 12).    character range mean character range mean character ratio SD  CL* 5.44–7.04 6.37 Mt1S 18–37 27 CW/CL 0.98 0.035  CW 5.51–6.64 6.26 Ta1S 7–22 14 CH/CW 0.50 0.026  CH 2.68–3.53 3.10 Ti2S 8–20 12 CFW/CL 0.73 0.047  CFW 4.15–5.45 4.68 STC2teeth 9–12 10 CHGW/L 0.16 0.011  ChGL 1.96–2.48 2.20 BulbL 2.11–2.54 2.68 SW/SL 0.78 0.028  ChGW 0.27–0.42 0.35 EmbL 1.36–1.67 1.54 LL/LW* 0.78 0.02  ChGCT 3–16 10 BulbW 0.83–1.00 0.93 PLSAPW/L 0.47 0.055  LL 0.96–1.24 1.09 EmbmidW 0.15–0.30 0.20 BulbW/EmbL* 0.60 0.033  LW 1.19–1.58 1.38 PalpTibL 2.74–3.30 3.07 EmbmidW/L 0.14 0.019  CUSP* 117–191 157 PalpTibW 1.26–1.53 1.41 PalpTibW/L 0.46 0.024  SL 3.52–4.40 4.02 PalpTibS 2–7 5 BulbL/TibL 0.77 0.031  SW 2.89–3.40 3.15 PalpPatS 1–2 1  PLSAPW 0.31–0.46 0.35 PalpFemS 5–7* 6  PLSAPL 0.68–0.98 0.70  Fe1S 3–9 6  Pa1S 0–7 (p0–3) 4  Female( K11083MV),  Strathbogie Range, Victoria, 36°56'Sapical segment short. Lengths. total 3.07; basal segment 145°38'E,  7 March 1982) — Size. Carapace length 7.83, 1.43; middle 0.69; apical 0.95. Apical segment width 0.56. width 7.07. Abdomen length 11.02, width 7.48. — Colour. — Genitalia. Spermathecae very short and broad. Apical Basic colour pattern. Abdominal chevrons distinct, well area narrower than expanded central region. Spermatheca separated. — Carapace. Slightly longer than wide, strongly length 0.82, width 0.68. raised. Height 4.01; frontal width 6.60. Cephalic length 5.60. Fovea wide, procurved anterior margin indented. Mid- Distribution. Western side of Great Dividing Range in dorsal cephalic setae do not reach fovea.Anterior strial setae Southern New South Walesand northern Victoria(Fig. 63). absent basally. Anterolateral carapace angle with a few weak This species is provisionally recorded from the Grampian bristles. — Eyes. Central eye region raised. Eye group width Ranges on the basis of a single female from the Hall’s Gap 2.11. Diameters: AME 0.27, ALE 0.41, PLE 0.34, PME area—male records are needed to confirm this. 0.23. Interdistances: AME–AME 0.36, AME–ALE 0.25, ALE–PLE 0.21, PLE–PME 0.17, PME–PME 0.96. Median Comments. In some males a few mid row or distal teeth ocular quadrangle length 0.70, anterior width 0.89, posterior may be missing, making keying such individuals difficult. width 1.43. — Chelicerae. Cheliceral groove margins diverge weakly distally. Groove length 2.65, middle width 0.54.   Hadronyche jensenae n.sp. Cheliceral teeth: 26 central, in single row occupying full length of groove; 13 prolateral; 11 retrolateral. Central tooth Figures 63–65; Table 19 row length variable. — Labium. Wider than long, anterior margin slightly concave, sides rounded. Length 1.55, width Etymology. The species is named for my wife, Greta Jensen 2.11. Cuspules occupying anterior two-thirds of labium. Gray. Labiosternal sigilla narrowed at midline. — Sternum. Ovoid.   Length5.10, width 4.01. Posteriorsternal sigilla elongate,  Types. Holotypemale: K11085( MV), Woodside, Victoria, 38°32'Sremoved from margins: length 0.99, width 0.34. — Palp. 146°53'E, 15 August 1966.  Paratypes. Victoria.  Males: K11086( MV), Spination: tibia 0–1 (strong bristles present), tarsus 8. Woodside, data as for holotype; K11087( MV), Morwell, 38°14'S 146°24'E.,  28 August 1967. Trichobothria: tibia p6 r6, tarsus 8. Tarsal claw with 4 teeth. — Legs. 4123. Legs I, IV subequal. Leg Femur Patella Tibia Metatarsus Tarsus Total 1 5.92 2.82 4.59 3.60 2.37 19.30 2 5.10 2.62 3.94 3.47 2.38 17.51 3 4.60 2.55 2.84 3.12 2.52 15.91 4 5.71 2.96 4.08 4.09 2.92 19.76  Tibia Iwidth 1.29. Metatarsus I proximal width 0.95. Coxae I, II with a few basally thickened setae frontally. Tibia I, II with few to no spines. Tarsal claw teeth legs I, II: superior 7, 7; inferior 3, 3. Trichobothria legs I, II: tarsus 9, 10; metatarsus 14, 12; tibia p6 r6, p6 r5. — Leg spination. Leg I; femur 0, patella 0, tibia 0, metatarsus 5, tarsus 4. Leg II: femur 0, patella 0, tibia 0, metatarsus 7, tarsus 5. Leg III: femur 0, patella p7, tibia 9 (p5 r2), metatarsus 15 (p5 rd2), tarsus 12. Leg IV: femur 0, patella 0, tibia 0, metatarsus 10, Figure 63. Collection records for  Hadronyche meridiana(squares) tarsus 14. — Abdomen. Posterior lateral spinnerets stout, and  H. jensenae(circles).   Table 19. Male morphological data—  Hadronyche jensenae(n = 4).    character range mean character range mean character ratio SD  CL* 4.72–6.60 5.77 Mt1S 14–27 (p0) 19 CW/CL 0.99 0.029  CW 4.84–6.60 5.72 Ta1S 6–10 8 CH/CW 0.45 0.03  CH 2.0–3.16 2.60 Ti2S* 5–7 (p0) 6 CFW/CL 0.70 0.043  CFW 3.36–4.85 4.06 STC2teeth 10–11 11 CHGW/L 0.16 0.013  ChGL 1.66–2.18 1.94 BulbL 1.25–2.34 2.17 SW/SL 0.81 0.022  ChGW 0.27–0.34 0.31 EmbL 1.15–1.36 1.28 LL/LW* 0.79 0.075  ChGCT 15–19 17 BulbW 0.86–1.09 0.99 PLSAPW/L 0.47 0.055  LL 0.78–1.29 1.02 EmbmidW 0.18–0.21 0.19 BulbW/EmbL* 0.77 0.022  LW 1.06–1.43 1.28 PalpTibL 2.71–3.09 2.82 EmbmidW/L 0.16 0.006  CUSP* 112–169 145 PalpTibW 1.26–1.52 1.40 PalpTibW/L* 0.50 0.003  SL 3.11–4.15 3.64 PalpTibS 1–2 2 BulbL/TibL* 0.77 0.053  SW 2.43–3.41 2.95 PalpPatS 1–3 2  PLSAPW 0.29–0.35 0.34 PalpFemS 0–3 1  PLSAPL 0.64–0.82 0.73  Fe1S 2–5 4  Pa1S* 0 0  Other material.  AMS KS112636, Vic. Dept. Agric., Bairnsdale (locality pattern; abdomen partly discoloured or damaged in all unknown),  20 June 1978.specimens. — Carapace. About as wide as long, cephalic area moderately raised. Height 2.65; frontal width 4.21. Fovea  Diagnosis. CL 4.72–6.60 (male). Small atracine spiders. procurved. Mid-dorsal cephalic setae few, do not reach fovea. Differ from other atracine species by having a short, broad Anterior strial setae absent.Anterolateral carapace angle with embolus with distal part set at distinct angle to shaft (Fig. weak hairs only. — Eyes. Central eye region slightly raised. 65B,C). Eye group width 1.57. Median ocular quadrangle length 0.60, anterior width 0.74, posterior width 1.10. Diameters:  Male( holotype) — Size. Carapace length 5.74, width 5.70. AME 0.20; ALE 0.35; PLE 0.29; PME 0.23. — Chelicerae. Abdomen length 6.5, width 4.7. — Colour. Basic colour Groove margins diverge weakly distally. Groove length 2.00, middle width 0.30. Cheliceral teeth. 15 central, occupying full length of groove; 9 prolateral; 12 retrolateral. — Labium. Much wider than long, rectangular, apically weakly indented. Length 0.92, width 1.25. Labiosternal sigilla broad. Cuspules c. 160, number moderate to low. — Sternum. Ovoid. Length 3.59, width 2.98. Posterior sternal sigilla ovoid to elongate; middle sigilla small. — Palp. Tegular area slightly wider than long. Embolus very short and broad, base strongly offset from tegulum. Distal part of embolus not twisted, but angled strongly at about 45° to shaft. Bulb length 2.23, width 1.01. Embolus length 1.31, midwidth 0.20. Tibia short, strongly incrassate proximally. Length of femur 2.71, patella 1.12, tibia 2.71. Width of tibia 2.01. Spination: femur 6; patella 1; tibia d2. Distal femur with spines, without sinuous bristles. — Legs. 4123. Leg Femur Patella Tibia Metatarsus Tarsus Total 1 5.34 2.52 3.94 3.46 2.38 17.64 2 4.79 2.30 3.74 3.19 2.38 16.40 3 4.08 2.14 2.89 3.03 2.24 14.38 4 5.17 2.28 4.08 3.90 2.82 18.25  Tibia Iwidth 0.94. Femora I, II with dorsal spines. Metatarsus I slightly thickened in proximal half, spines strong, scattered. Tibia II without apophysis, ventral spines few, grouped proximally with c. two larger spines toward middle; distoventral tibia weakly concave. Metatarsus II weakly to moderately sinuous with small, apophysis midventrally. Tarsi III, IV slightly swollen. Anterior coxal hairs unmodified. Tarsal and distal metatarsal scopulae: legs I, Figure 64.  Hadronyche jensenae, male: ( A) cephalothorax and II weak; legs III, IV moderate. Tarsal claw teeth legs I, II: chelicerae, dorsal; ( B) cephalothorax and chelicerae, lateral; ( C) superior 11, 11; inferior 0, 0. Trichobothria legs I, II: tarsus cheliceral groove teeth; ( D) sternum, labium and maxilla; ( E) leg 8, 7; metatarsus 6, 6; tibia p5 r5, p5 r5. — Leg spination. Leg II, prolateral; ( F) spinnerets; ( G) palp, prolateral. Scale lines 1 mm. I: femur d5, patella 0, tibia 18, metatarsus 19, tarsus 10. Leg  Figure 65.  Hadronyche jensenae, male ( A,D–K, AMS KS112636; B,C, 11086 [MV]): ( A) palp, prolateral; ( B) bulb; ( C) embolus, distal; ( D,E) distal patella and tibia I— D, ventral, E, prolateral; ( F,G) metatarsus I— F, ventral, G, prolateral; ( H,I) distal patella and tibia II— H, ventral, I, prolateral; ( J,K) metatarsus II— J, ventral, K, prolateral. Scale lines: 0.2 mm, except B0.5 mm and D,F,G0.3 mm. II: femur d4, patella 0, tibia 5, metatarsus 11, tarsus 9. Leg III: femur d1, patella pd10, tibia 10 (r2), metatarsus 17 (p5 r2), tarsus 8. Leg IV: femur d1–2 (strong bristles present), patella 0, tibia 3 (rd1), metatarsus 10, tarsus 11. — Abdomen. Bristles stronger anteriorly with a prominent frontal pair. Posterior lateral spinnerets with rather short apical segment. Lengths: total 1.90; basal segment 0.84; middle 0.42; apical 0.64. Apical segment width 0.34.   Distribution. Central Gippsland, Victoria(Fig. 63). [903,1467,1371,1390] AMS 55 340 KS2065 1 holotype 1978-10 AM, AMS L. Abra United Kingdom Kulnura -33.35 Embolus 1205 151.28334 South Wales. Males 55 340 KS13393 1 paratype 1983-10 AMS Width & Spination Wales -33.216667 Kulnura 1206 151.21666 55 340 KS13401 1 paratype 1983-09 AMS Wales Somersby -33.216667 Nelson 1206 151.21666 Rodgers 55 340 KS4102 1 paratype Wales -33.35 October - Legs. 1205 151.28334 55 340 1 paratype [172,273,153,172] R Mascord 56 341 1 [278,648,153,172] AMS -33.35 Somersby 1205 151.28334 56 341 KS8571 1 L. Abra October Diameters 56 341 1 [242,745,177,196] 1979-10-03 AMS -33.35 Somersby 1205 151.28334 56 341 KS4108 1 PME K. Mascord Chelicerae. Anterodorsal 56 341 1 [231,745,201,220] 1956-11 AMS -33.35 Somersby 1205 151.28334 56 341 KS991 1 [133,220,225,244] V Brown 56 341 1 AMS September & Cheliceral & J. Brown. Females -33.35 Somersby 1205 151.28334 56 341 KS7335 1 1984-10 AMS L. Abra Groove -33.433334 Kariong 1205 151.3 Gosford 56 341 KS15749 1 1984-03-23 AMS -33.216667 Kulnura 1206 151.21666 Cheliceral 56 341 KS14381 1 1984-06-04 AMS Labium. Short -33.433334 Kariong 1205 151.3 56 341 KS14757 1 1984-04 AMS Labium -33.216667 Kulnura 1206 151.21666 56 341 KS14234 1 [334,748,345,364] AMS -33.216667 Kulnura 1206 151.21666 56 341 KS14233 1 1984-03 AMS Kulnura -33.216667 Labiosternal 1206 151.21666 56 341 KS14551 1 1984-04-05 AMS Kulnura -33.216667 Cuspules 1206 151.21666 56 341 KS14226 1 [239,745,417,436] 1984-04-13 AMS Kariong -33.433334 1205 151.3 56 341 KS14547 1 [121,571,502,525] 1983-10-25 AMS Kulnura 56 341 KS13394 1 1983-09 AMS Kulnura Palp. Tegular area 56 341 KS13399 1 1983-09 AMS Distal Kulnura Embolus 56 341 KS13400 1 AMS K. Gould Somersby September 56 341 KS13377 1 [294,716,590,613] 1978-10 AMS Somersby 56 341 KS1929 1 Bulb Abra Embolus 56 341 1 [187,1421,619,646] AMS Embolus Kulnura - Somersby area 56 341 KS23333 1 1973-03-03 PME, AMS, R, AM Size. Carapace & Abdomen & Colour. Basic & Carapace. Almost & Distribution. Kulnura & Central & New & Height & South Wales & Cephalic & Fovea & Mid-dorsal & Anterior & Comments. A & Kulnura & Anterolateral & Eyes. Central & Eye & These & Diameters & Interdistances & Median & Chelicerae. Cheliceral & Groove & Cheliceral & Labium. Short & Cuspules & Etymology. The & Sternum. Ovoid & Length & Posterior & Palp. Spination & Types. -35.716667 Batemans Bay 1191 150.21666 Somersby region 58 343 KS867 2 West Coast holotype [121,1421,1598,1631] AMS Benandarah State Forest -35.65 Tarsal South Wales Males 58 343 KS2276 1 [505,1301,1627,1655] 1978-11-30 1979-01-04 1978-11-30 IV C. Horseman 150.25 58 343 1 AMS Bermagui & K. Walker -36.416668 1187 150.06667 58 343 KS863 1 [1167,1421,1660,1679] AMS Bondi State 58 343 KS12124 1 [794,1284,1708,1727] 1979-02-07 AMS Bowral -34.466667 1199 150.41667 58 343 KS2665 1 [794,1373,1732,1751] 1978-11-28 AMS Crookwell -34.45 1199 149.46666 58 343 KS2081 1 Willis 58 343 1 [860,1417,1756,1775] 1971-02-14 AMS Narooma -36.2 1188 150.13333 58 343 KS862 1 2004-01-21 AMS Exeter -34.6 1198 150.31667 58 343 KS88199 1 [850,1369,1804,1823] 1979-01 AMS Narooma -36.216667 1188 150.13333 58 343 KS3876 1 [794,1417,1828,1847] AMS -34.65 1198 150.3 58 343 KS4664 1 [196,1321,1846,1871] AMS Metatarsus & Coxal February -36.416668 Bermagui 1187 150.06667 58 343 KS16446 1 II, AMS Walker & Tarsal -37.1 Bondi State Forest 1183 149.25 Bombala 58 343 KS12139 1 1981-01 II, AMS Trichobothria & Gowing -34.466667 Bowral 1199 150.41667 58 343 KS6163 1 [794,1418,1924,1943] 1980-10 AMS H. Styles -34.533333 Grabben Gullen 1198 149.4 58 343 KS6825 1 [121,1417,1934,1967] 1980-12-16 R, AMS Leg & Leg I & Chown -35.9 1190 150.1 58 343 KS4506 1 [626,910,1963,1991] 1980-01 II 58 343 1 [918,1418,1972,1991] 1971-06-15 AMS -36.2 Narooma 1188 150.13333 58 343 KS3625 1 [616,892,1993,2016] III Cobcroft 58 343 1 1966-07-30 1984-02-05 1966-07-30 AMS, R C. Horseman & G. Gowing & A. B. Thompson & H. Styles & W. Nooijen & M. Johnson & Diameters & Moss Carapace. About Bateman's Bay Benandarah State Forest 59 344 KS2032, KS867, KS12150, KS3460, KS6163, KS13806, KS6825, KS7567, KS2117, KS2293, KS1163, KS6275 1 New South Wales 1945-12 1980-12-27 1945-12 AMS, PME, V J. Kelly & Narooma & Groove & B. Plunkett-Cole Vale 59 344 KS868, KS861 1 Valais 1937-06 1980-11 1937-06 AMS, MV H. W. Kemp & K. Walker & G. Gowing & Bowral, H. H & N. C. de Meyrick & A. Livingstone & Narooma & A. Brown Bondi State Forest Bega 59 344 KS1865, KS1867, KS11996, KS1872, KS1873, KS4777, KS2961 1 New South Wales AMS March & P. Ormay. & Embolus & Distal Palp. Tegular area Tidbinbilla Nature Reserve 59 344 KS13824 1 Australian Capital Territory [185,1421,389,421] MNHN Paris Eye Melbourne 62 347 1 Victoria holotype PME Ellinbank & Warragul Other Diameters 62 347 1 Victoria 1962-05 MV, AMS J. Copley Chelicerae. 62 347 KS8471 1 Victoria [499,748,469,488] 1978-04-03 AMS Emerald 62 347 1 Victoria KS Cheliceral 62 347 KS8472 1 Victoria [200,745,493,512] 1963-06-06 1979-04-20 1963-06-06 MV I. Bissett Leongatha Emerald 62 347 1 Victoria 1981-03-10 AMS Cheliceral & L. Gibson Back Creek Rd. Gembrook Gembrook State Forest 62 347 KS7292 1 Victoria [450,699,541,560] 1966-04-15 Mirboo North 62 347 1 Victoria 1967-03-26 MV Mirboo North Burr 62 347 1 Victoria [593,1381,565,588] MV J. Seebeck & Central 62 347 1 Victoria [121,489,589,608] 1979-04-23 AMS Mount Eliza 62 347 KS8473 1 Victoria 1982-03-25 AMS Strezelecki Ranges Mount Tassie Length Labium. Wider 62 347 KS8930 1 Victoria [424,508,613,632] R Waters 62 347 1 Victoria [513,744,613,632] AMS Tanjil South 62 347 KS8933 1 Victoria 1951-09 AMS N. Barton Thorpdale Labiosternal April 62 347 KS8474 1 1963-02-06 MV C. Oke Yallourn Cuspules Slight 62 347 1 [273,650,685,704] 1964-05-04 MV J. Irving Yarra Junction 62 347 1 1964-04-20 MV J. Mooney & Sternum. Ovoid. Length Yarra Junction 62 347 1 1982-10-10 MV, AMS J. Mooney & Palp. Cockatoo Posterior 62 347 KS10775 1 Victoria 1979-10-06 AMS McKelvey Tegular area Ferny Creek Embolus 62 347 KS8477 1 Victoria [372,745,757,776] 1983-01-23 AMS Hawthorn 62 347 KS12618 1 Victoria [121,606,781,800] 1976-10-26 AMS M. Harvey Milford 62 347 KS8476 1 1980-04 AMS Mirboo North 62 347 KS8339 1 [329,655,805,824] 1981-10 AMS Moe area 62 347 KS8932 1 1979-06-05 AMS N. Barton & Bulb Mount Evelyn Lilydale Shire 62 347 KS8470 1 1982-03 AMS Moe Embolus Newborough 62 347 KS8931, 1.03 1 1982-02-14 AMS N. Barton Spination Sassafras Width 62 347 KS8926 1 [278,744,877,896] 1983-01 AMS N. Wentworth Warragul 62 347 KS10560 1 K11082 1981-10-18 MV S. Johnson & Size. Carapace & Apical & Abdomen & Genitalia. Spermathecae & Colour. Basic & Anterodorsal & Carapace. Slightly & Length -37.833332 Hoddles Creek 1179 145.6 64 349 1 Victoria NHM, R -37.383335 Cheliceral 1181 144.58333 Mt. Macedon 64 349 BMNH 1903.2 1 Victoria holotype 1980-07-22 AMS New South Wales 64 349 KS4503 1 New South Wales [990,1418,1911,1930] 1971-01-10 AMS M. Pearce 64 349 KS917 1 New South Wales [794,1275,1935,1954] AMS June Wantagong 64 349 KS916 1 [1286,1365,1935,1954] R Flynn 64 349 1 AMS Cuspules Labiosternal 64 349 AMS 2.01 1 [794,1080,1959,1978] 1971-04-01 KS Wantagong 64 349 KS915 1 [1089,1175,1959,1978] R Hunter 64 349 1 1948-09 AMS Wondalga Sternum. Broad 64 349 KS913 1 [941,1287,1982,2002] C. H. Jagoe. New South Wales 64 349 1 New South Wales 1980-01-10 AMS Adelong 65 350 64 349 KS4973 1 New South Wales 1980-10 AMS Rosewood 65 350 KS6053 1 New South Wales [288,562,177,196] D. M. Fordham. Victoria 65 350 1 Victoria 1978-10-13 AMS Baranduda & Labium. Wider Length 65 350 KS8488 1 Victoria [448,791,201,220] 1982-06 AMS Lake Bolac 65 350 KS10771 1 Victoria [166,561,225,244] 1963-09-06 MV H. Parnaby Avenel 65 350 1 1965-06-19 MV Labiosternal Euroa Cuspules 65 350 1 [473,794,249,268] 1965-07-04 Pranjip Longwood area 65 350 1 1980-08-15 MV, AMS Sternum. Ovoid. Length Seymour Trawool 65 350 KS8484 1 1980-02-05 P May Palp. Tegular area Violet Town Williams 65 350 1 1974-04-29 MV, AMS A. Kube Embolus Kilmore Willowmavin 65 350 KS8485 1 1984-01-15 AMS Rutherglen Shepparton Research Institute 65 350 KS13651-3 1 Victoria 1981-05 -36.9 Avenel 1185 144.2 65 350 1 Victoria 1982-03-07 -36.933334 Strathbogie Range 1184 145.63333 67 352 1 Victoria K11085 [166,1467,1377,1400] MV Length & Posterior & Types. -38.533333 Woodside 67 352 1 Victoria holotype K11086 MV Males Woodside Spination 67 352 1 Victoria paratype K11087 1967-08-28 MV Length & Posterior & Types. -38.233334 Morwell 1177 146.4 67 352 1 Victoria paratype KS112636 1978-06-20 AMS, KS 68 353 1