Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda) Wesener, Thomas Enghoff, Henrik Sierwald, Petra ZooKeys 2009 2009-09-04 19 19 1 128 6MN9Q Wesener Wesener & Enghoff & Sierwald 2009 [140,601,1187,1214] Diplopoda Spirobolellidae Hylekobolus Animalia Spirobolida 101 102 Arthropoda species brachiosauroides sp. nov.   urn:lsid:zoobank.org:act: 40FD10B2-63FB-4627-AAAC-6961BE9102E5    Materialexamined: 26 ♂, 21 ♀, 6 imm.  Holotype:1 ♂ ( 55 mmlong), FMMC W089, Vohimena Mountains, Westslope, Ivorona,  114 m, rainforest, 24°50.026’ S, 46°56.943’ E, leg. Weseneret al.,  23.V.2007.   Paratypes:3 ♂, 4 ♀, FMMC W089, same data as holotype; 1 ♂, 1 ♀, ZMUC W089, same data as holotype; 1 ♂, ZMH W089, same data as holotype; 1 ♂, 1 ♀, CAS W089, same data as holotype; 1 ♂, ZSM W089, same data as holotype.    Othermaterial examined:4 ♂, 4 ♀, W089, same data as holotype, voucher specimens deposited at the University Antananarivo; 5 ♂, 4 ♀, 2 imm., FMMC W049B, Andohahela, Isaka-Ivondro Sud,  46 m, rainforest, 24°49.690’ S, 46°51.971’ E, leg. Weseneret al.,  08.VI.2007; 5 ♂, 4 ♀, 2 imm., W049B, same data as previous, voucher specimens deposited at the University Antananarivo; 2 ♂, 2 ♀, FMMC W099, Ambatotsirongorongo Mountain, Grand Lavasoa, rainforest, 25°5’10.23” S, 46°44’55.93” E, rainforest, leg. Weseneret al.,  14.VI.2007; 2 ♂, 1 ♀, 2 imm., W099, same data as previous, voucher specimens deposited at the University Antananarivo.    Differentialdiagnosis:  H. brachiosauroidesdiffers in size and colour from the sympatric living  H. albicollaris sp. n.The posterior gonopods ( Figs 55M, O) of  H. brachiosauroidesdiffer from other, same-sized  Hylekobolusspecies: the lateral margin carries a thick, swollen, well-rounded projection ( Fig. 55M), a feature only shared with  H. montanus sp. n.  H. brachiosauroidesdiffers from  H. montanus sp. n.in the presence of a protruding apical process ( x) on the posterior gonopods, a weakly protruding lateral retrorse process of the anterior gonopod ( Fig. 55N), and a different colour pattern.    Description. Measurements: males with 50 or 51 rings, 53–62 mmlong, 4.8–5.2 mmwide. Females with 50 or 51 rings, 58–72 mmlong, 5.1–6.0 mm wide.  Coloration:anterior part of head and collum, posterior part of telson and posterior margin of body rings lightly red or white ( Figs 53A, 55A, C). Legs and antennae white ( Fig. 53A). Anterior part of body rings, telson, posterior part of head and collum black ( Figs 53A, 55A). Eyeswith 40–44 ocelli ( Fig. 55D).  Anterior gonopodsternite basally wide, apically elevated into a triangular lobe with a triangular, wide, well-rounded tip ( Fig. 55L). Mesal process of coxite wide and only slightly protruding ( Fig. 55L). Apical process of telopodite large and well-rounded. Retrorse margin wide, not projecting above telopodite margin ( Fig. 55N). Margin of apical process basally wide and projecting ( Fig. 55N).  Posterior gonopodstelopodite bent 90° ( Fig. 55O). Lateral margin with a thick, swollen, well-rounded projection ( w, Fig. 55M). Apical process ( x) apically slender, basally with a widely protruding margin ( Fig. 55M). Membranous lobe ( y) rectangular ( Fig. 55M). Projection of sperm canal ( z) long and slender, slightly curved towards apical process ( Fig. 55M).   Intraspecificvariation:coloration of the posterior margin of body ring varies in adults between white and reddish, while it is always white in imm..    Distributionand ecology: this species was found in the isolated Ambatotsirongorongo Mountain, the Andohahela Isaka-Ivondro corridor as well as on the western slopes of the Vohimena Mountain ( Fig. 54).  H. brachiosauroidescould not be collected north of the Isaka-Ivondro corridor, where  H. goodmani sp. n.and  H. montanus sp. n.occur. In the eastern Vohimena slopes,  H. rufus sp. n.replaces  H. brachiosauroides. All individuals were collected inside dead logs.    Etymology:  brachiosauroides, adjective, refers to the dinosaur genus  Brachiosaurus, whose neck and head closely resemble the general shape of the posterior gonopods of this millipede. 2608818240 2007-05-23 FMMC Wesener Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 W089 27 21 holotype 2608818212 [472,1103,1363,1390] 2007-05-23 FMMC Wesener Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 W089 4 4 paratype 2608818309 [140,652,1399,1425] 2007-05-23 ZMUC Wesener Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 W089 1 1 paratype 2608818296 [661,1103,1399,1425] 2007-05-23 ZMH Wesener Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 W089 1 paratype 2608818227 [140,629,1434,1460] 2007-05-23 CAS Wesener Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 W089 1 1 paratype 2608818332 [639,1080,1434,1460] 2007-05-23 ZSM Wesener Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 W089 1 paratype 2608818349 2007-05-23 FMMC University Ivorona 114 -24.833767 Vohimena Mountains 1 46.94905 West 101 102 4 4 Antananarivo 2608818355 2007-06-08 FMMC Wesener 46 -24.828167 Isaka-Ivondro Sud 1 46.866184 Andohahela 101 102 W049 6 4 Antananarivo 2608818354 2007-06-08 FMMC University 46 -24.828167 Isaka-Ivondro Sud 1 46.866184 Andohahela 101 102 6 4 Antananarivo 2608818258 2007-06-14 FMMC Wesener 102 103 -25.086176 Grand Lavasoa 1 46.74887 Ambatotsirongorongo Mountain 101 102 W099 2 2 Antananarivo 2608818361 2007-06-14 FMMC University -25.086176 Grand Lavasoa 1 46.74887 Ambatotsirongorongo Mountain 102 103 3 1 Antananarivo