Panelus dichrous Gillet, 1925: 4 Paulian 1935: 119 1939: 70 Lepanus dichrous : Matthews 1974: 111 Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species Gunter, Nicole L. Weir, Thomas A. Zootaxa 2021 2021-02-03 4923 1 1 66 6P88C (Gillet, 1925) Gillet 1925 [151,539,1806,1833] Insecta Scarabaeidae Lepanus Animalia Coleoptera 49 50 Arthropoda species dichrous  ( Figs. 13D, H, J, L; 14B; 15B; 16B; 17A, C)      Panelus dichrous Gillet, 1925: 4;  Paulian 1935: 119,  1939: 70.     Lepanus dichrous: Matthews 1974: 111.   Type material digitally examined. Holotype: “Queensl. Mjöberg”(printed) / “Ma– landa” (printed) / “Typ.” (printed) / “ dichrous Gillet type.” (handwritten in black, with “ type” in red ink) / “ Typus” (printed) / “261 68” (“261” printed, “68” handwritten) / “3555 E92 +” (printed) / “ NHRS_ JLKB 000027315” (printed) [ 17°21’00”S, 145°35’00”E].  Other material examined. 831 specimensfrom over 50 localities were also examined.    Queensland: North Bell Peaksummit, 17°05’19”S, 145°52’44”E,  15–16.ix.1981(T55406–T55421 16 specimensin QM);  North Bell Peaksummit, 17°05’19”S, 145°52’44”E,  16.ix.1981(T55422 1 specimenin QM);  North Bell Peaksummit, 17°05’19”S, 145°52’44”E,  19–22.xi.1990(T55423–T55435, T55835 14 specimensin QM);  Robson Creek, Atherton[ 17°07’00”S, 145°37’00”E],  17.i.1996, R.L. Kitching(25-058288 1 specimenin ANIC);  Robson Creek, Atherton[ 17°07’00”S, 145°37’00”E],  18.i.1996, R.J. Kitching& H. Mitchell(25-058287 1 specimenin ANIC);  Robson Creek[ 17°08’00”S, 145°38’00”E] (25-065483 1 specimenalso as COL3003, 25-065484 1 specimenalso as COL3004 specimens in ANIC);  Danbulla Forestry Reserve,  13 kmNE of Yungaburra[ 17°10’00”S, 145°38’00”E],  13.ii–6.iii.1987, R.I. Storey& De Faveri(25-032545 1 specimenin QDPC);  Danbulla Forestry Reserve,  13 kmNE of Yungaburra[ 17°10’00”S, 145°38’00”E],  13.i–13.ii.1987, R.I. Storey& De Faveri, (25-058283 2 specimensin ANIC);  Danbulla Forestry Reserve,  13 kmNE of Yungaburra[ 17°10’00”S, 145°38’00”E],  26.iii– 23.iv.1987, R.I. Storey& De Faveri, (25-058285 1 specimenin ANIC);  Danbulla Forestry Reserve,  13 kmNE of Yungaburra[ 17°10’00”S, 145°38’00”E],  28.vii–3.ix.1987, R.I. Storey& De Faveri, (25-058284 1 specimenin ANIC);  Danbulla Forestry Reserve,  13 kmNE of Yungaburra[ 17°10’00”S, 145°38’00”E],  6–26.iii.1987, R.I. Storey& De Faveri(25-058286 2 specimensin ANIC);  Danbulla Scientific Reserve, 17°11’09”S, 145°39’36”E,  10.xii.1995–  7.ii.1996(T55306 specimen in QM);  Malanda Falls, 17°21’22”S, 145°35’13”E,  750 m,  23–24.i.2012, G.B. Monteith(25-053935 17 specimens, 25-053936 24 specimensin ANIC);  Bartle Frere, Gourkaroad, 17°22’38”S, 145°44’34”E,  28–30.xi.1998(T70869–T70877 9 specimensin QM);  Bartle Frere, west base, 17°22’47”S, 145°45’51”E,  19–20.xi.2001(T111969–T111984 16 specimensin QM);  Bartle Frere, west base, 17°22’47”S, 145°45’51”E,  25.xi.1994–  10.i.1995(T55845–T55857 13 specimensin QM);  Bartle Frere, west base, 17°22’47”S, 145°45’51”E,  10.i–31.iii.1995(T55858–T55864 7 specimensin QM);  Bartle Frere, west base, 17°22’47”S, 145°45’51”E,  7.iii–15.v.1995(T55865–T55889 25 specimensin QM);  Bartle Frere, west base, 17°22’47”S, 145°45’51”E,  28–30.xi.1998(T70860–T70868 9 specimensin QM);  Bartle Frerewest base [ 17°23’00”S, 145°46’00”E],  750 m,  28.xi.1998, G.B. Monteith(25-060847 7 specimensalso as COL587– 589 specimensin ANIC);  Bartle Frerewest base [ 17°23’00”S, 145°46’00”E],  750 m,  28–29.xi.1998, G.B. Monteith(25-060850 2 specimensalso as COL616– 17 in ANIC);  Malanda, 3.5 kmS, 17°23’27”S, 145°35’46”E,  6–9.ii.1999(T75465– T75470, T75485 7 specimensin QM);  Topaz, Pei Road( Frithhouse in QM),  17°24’23”S, 145°41’05”E,  6.xii.1993–  25.ii.1994(T55220–T55226 7 specimensin QM);  The Crater National Park, 17°25’48”S, 145°29’24”E,  980 m,  18–19.ix.2010, G.B. Monteith(25-053860 1 specimenalso as COL1613, 25-059235 2 specimens, 25-060796 1 specimen, 25-060886 15 specimensalso as COL1160 and COL1161, 25-053932 11 specimensin ANIC);  The Crater National Park, 17°25’48”S, 145°29’24”E,  980 m,  22–23.i.2012, G.B. Monteith(25-053933 20 specimensin ANIC);  Thiaki, main rainforest, 17°25’55”S, 145°30’58”E,  990 m,  21–23.i.2012, G.B. Monteith(25-053849 1 specimenin ANIC);  Thiakicottage, 17°26’02”S, 145°30’40”E,  1000 m,  19–23.i.2012, G.B. Monteith(25-053843 2 specimensin ANIC);  Topaz National Park, SE of Malanda[ 17°26’00”S, 145°43’00”E],  720 m,  28.vii.1982, S. Peck& J. Peck(25-058290 1 specimenin ANIC);  Hugh  Nelson Range, 17°27’00”S, 145°29’00”E],  1150 m,  6.iii–4.iv.1995, P. Zborowski(25-058292 1 specimenin ANIC);  Longlands Gap, 3 ways road junction, 17°27’56”S, 145°28’30”E,  3–6.ii.1999(T75487 specimens in QM);  BS1 Longlands Gap[ 17°28’00”S, 145°29’00”E],  1150 m,  1.xii.1994–  3.i.1995, P. Zborowski(25-058291 1 specimenin ANIC);  BS1 Longlands Gap[ 17°28’00”S, 145°29’00”E],  1150 m,  27.ii–26.iii.1996, P. Zborowski(25-058293 1 specimenin ANIC);  Kenny Road, Atherton Tablelands, 17°28’13”S, 145°32’04”E,  25.xi.1994–  10.i.1995(T55296 specimens in QM);  Kenny Road, Atherton Tablelands, 17°28’13”S, 145°32’04”E,  10.i–31.iii.1995(T55297–T55301 5 specimensin QM);  Kenny Road, Atherton Tablelands, 17°28’13”S, 145°32’04”E,  7.iii–15.v.1995(T55302–T55303 2 specimensin QM);  Tumoulin Road, Evelyn, 17°29’52”S, 145°28’32”E,  5–11.ii.1998(T57499 1 specimen, T60022–T60026 5 specimensin QM);  Millaa Millaa Falls, 17°29’53”S, 145°36’32”E,  5.v.1983(T55230 1 specimenin QM);  Millaa Millaa Falls, 17°29’53”S, 145°36’32”E,  8–10.xii.2000(T105998 1 specimenin QM);  Millaa Millaa Falls, 17°29’53”S, 145°36’32”E,  6– 9.ii.1999(T75457–T75464 8 specimensin QM); above Millaa Millaa Falls, 17°29’53”S, 145°36’47”E,  830 m,  23–25.i.2012, G.B. Monteith(25-053937 32 specimens, 25-053938 41 specimensin ANIC); above Millaa Millaa Falls, 17°29’56”S, 145°36’50”E,  830 m,  23–25.i.2012, G.B. Monteith(25-053939 52 specimens, 25-053940 23 specimensin ANIC);  Millaa Millaa Falls(above) [ 17°30’00”S, 145°36’00”E],  800 m,  18–20.iv.1999, C. Reid& I. Reid(25-058295 4 specimensin ANIC);  Millaa Millaa Falls[ 17°30’00”S, 145°36’00”E],  780 m,  11.v.1974, R.I. Storey(25-058513 1 specimenin QM);  Millaa Millaa Falls[ 17°30’00”S, 145°36’00”E],  780 m,  23–25.i.2012, G.B. Monteith(25-053934 25 specimensin ANIC, 25-055612 6 specimensin ANIC);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  21–22.iv.1973, R.I. Storey(25-032538 2 specimensin QDPC, 25-058514 5 specimensin QM);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  830 m,  15–16.ix.2010, G.B. Monteith(25-059233 2 specimens, 25-060794 35 specimens, 25-060857 5 specimensin ANICalso as COL1104– 1108);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  760 m,  10.iv–4.v.1990, R.I. Storey& K.H. Halfpapp(25-032548 2 specimensin QDPC);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  760 m,  13.iii–10.iv.1990, R.I. Storey& K.H. Halfpapp(25-032541 1 specimenin QDPC);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  760 m,  15.viii–12.ix.1990, R.I. Storey& K.H. Halfpapp(25-032544 3 specimensin QDPC);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  760 m,  4.i–7.ii.1990, R.I. Storey& K.H. Halfpapp(25-032540 2 specimensin QDPC);  Millaa Millaa Fallsvia Millaa Millaa[ 17°30’00”S, 145°37’00”E],  760 m,  7.ii–13.iii.1990, R.I. Storey& K.H. Halfpapp(25-032539 3 specimensin QDPC);  Kjellberg Roadturnoff, 17°31’57”S, 145°36’27”E,  25.xi.1994–  10.i.1995(T55317 1 specimenin QM);  Kjellberg Roadturnoff, 17°31’57”S, 145°36’27”E,  7.iii–15.v.1995(T55318–T55328 11 specimensin QM);  Kjellberg Roadturnoff, 17°31’57”S, 145°36’27”E,  10.i–31.iii.1995(T55329–T55338 10 specimensin QM);  Kjellberg Roadturnoff, 17°31’57”S, 145°36’27”E,  6–9.ii.1999(T75443–T75446, T75451–T75456 10 specimensin QM);  Kjellberg Road, State Forest, boundary [ 17°32’00”S, 145°35’00”E],  920 m,  6–9.ii.1999, G.B. Monteith& D. Cook(25-060841 11 specimensin ANIC);  Mount Fisher, Kjellberg Road, 17°32’34”S, 145°33’31”E,  1.xii.1993–  25.ii.1994(T55339– T55340 2 specimensin QM);  Kjellberg Road, State Forestboundary, 17°32’09”S, 145°34’39”E,  6–9.ii.1999(T75436–T75442, T75447–T75450 11 specimensin QM);  Wooroonooran National Park, 17°34’06”S, 145°42’21”E,  500 m,  9–14.ix.2004, L. Masner(CMNEN-00030220–000030221 2 specimensin CMNC);  Henrietta Creek, Palmerston National Park[ 17°34’00”S, 145°45’00”E],  22–24.iv.1973, R.I. Storey(25-058512 4 specimensin QM, CMNEN-00030222 1 specimenin CMNC);  Beatrice River, 2 kmS, 17°34’35”S, 145°41’25”E,  1.xii.1993–  25.ii.1994(T55304–T55305 2 specimensin QM);  Palmerston, Scenic Road, 17°34’44”S, 145°41’53”E,  650 m,  15–16.ix.2010, G.B. Monteith(25-060795 15 specimensin ANIC);  PalmerstonForestry turnoff, 17°35’20”S, 145°42’14”E,  700 m,  15–16.ix.2010, G.B. Monteith(25-059234 1 specimen, 25-060885 15 specimensin ANICalso as COL1157, COL1159);  Maalan State Forest, RavenshoeRoad, 17°35’23”S, 145°35’40”E,  7.iii–15.v.1995(T55104–T55133 30 specimensin QM);  Maalan State Forest, RavenshoeRoad, 17°35’23”S, 145°35’40”E,  10.i–31.iii.1995(T55134– T55152 19 specimensin QM);  Maalan State Forest, RavenshoeRoad, 17°35’23”S, 145°35’40”E,  25.xi.1994–  10.i.1995(T55153–T55186 34 specimensin QM);  Maalan State Forest, RavenshoeRoad, 17°35’23”S, 145°35’40”E,  6–9.ii.1999(T75471–T75484 14 specimensin QM);  Maalan Road,  2 kmS Palmerston Highway, 17°35’56”S, 145°41’58”E,  7.iii–15.v.1995(T55079–T55089 11, T55103 specimens in QM);  Maalan Road,  2 kmS Palmerston Highway, 17°35’56”S, 145°41’58”E, 10.i.  1–7.iii.1995(T55090–T55092 3 specimensin QM);  Maalan Road,  2 kmS Palmerston Highway, 17°35’56”S, 145°41’58”E,  26.xi.1994–  10.i.1995(T55093–T55102 10 specimensin QM);  Henrietta Creek, Palmerston National Park, 17°35’58”S, 145°45’28”E,  30.xi.1992–  15.iv.1993(T111989 1 specimenin QM);  Palmerston National Park, west boundary, 17°33’36”S, 145°41’15”E,  1.xii.1993–  25.ii.1994(T55836– T55839 4 specimensin QM);  Mount Father Clancy,  800 m, 17°35’58”S, 145°38’04”E,  6–14.xii.1988(T55840– T55844 5 specimensin QM); road to Maalan Palmerston National Park[ 17°36’00”S, 145°40’00”E],  23–24.iv.1978, R.I. Storey(25-058511 17 specimensin QM, CMNEN–00030223 1 specimenin CMNC);  Rosina Creek, 14 kmSE Millaa Millaa[ 17°36’00”S, 145°43’00”E],  720 m,  24.vi–2.viii.1982, S. Peck& J. Peck(25-058289 4 specimensin ANIC);  Palmerston National Park[ 17°37’00”S, 145°47’00”E],  1.iv.1968, E. Matthews(25-057885 4 specimensin ANIC);  South Johnstone Forestry Camp, 5 kmW, 17°38’12”S, 145°40’15”E,  1.xii.1993–  25.ii.1994(T55214– T55217 4 specimensin QM);  Vine Creek, Majors Mountain, 17°40’58”S, 145°32’02”E,  4–6.ii.1999(T75494, T76145, T76152 3 specimensin QM);  Vine Creek, Majors Mountain, 17°40’58”S, 145°32’02”E,  15–18.iv.1999(T79112 1 specimenin QM);  Charmillin Creekcrossing, 17°42’03”S, 145°31’21”E,  2–5.xii.1998(T70886–T70888 3 specimensin QM);  Charmillin Creekcrossing, 17°42’03”S, 145°31’21”E,  5–11.ii.1998(T60017–T60021, T60425–T60433 14 specimensin QM);  Charmillin Creek, 17°42’54”S, 145°31’34”E,  880 m,  18–19.ix.2010, G.B. Monteith(25-060856 2 specimensin ANIC);  Ravenshoe State Forest, Tully FallsRoad [ 17°43’00”S, 145°33’00”E],  15.xi–12.xii.1987, A. Walford-Huggins(25-058294 2 specimensin ANIC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  1.x–5.xi.1987, R.I. Storey& Dickinson(25-058272 5 specimensin ANIC, 25-032546 1specimenin QDPC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  20.iv–15.v.1988, R.I. Storey& Dickinson(25-058274 specimens in ANIC, 25-032572 1 specimenin QDPC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  20.iv–25.v.1988, R.I. Storey& Dickinson(25-058269 2 specimensin ANIC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  3.iii–20.iv.1988, R.I. Storey& Dickinson(25-058280 2 specimensin ANIC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  5.i–7.ii.1988, R.I. Storey& Dickinson(25-058276 2 specimensin ANIC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  7.xii.1987–  7.i.1988, R.I. Storey& Dickinson(25-058279 1 specimenin ANIC);  Tully Falls State Forest,  18 kmSSW Ravenshoe[ 17°46’00”S, 145°31’00”E],  730 m,  7.xi–7.xii.1987, R.I. Storey& Dickinson(25-058270 1 specimenin ANIC);  Tully Falls State Forest,  11 kmSSW Ravenshoe[ 17°47’00”S, 145°34’00”E],  900 m,  5.i.1988–  7.ii.1988, R.I. Storey& Dickinson(25-058275 1 specimenin ANIC);  Tully Falls State Forest,  11 kmSSW Ravenshoe[ 17°47’00”S, 145°34’00”E],  900 m,  5.xi–7.xii.1988, R.I. Storey& Dickinson(25-058282 2 specimensin ANIC);  Tully Falls State Forest,  9.5 kmSSW Ravenshoe[ 17°47’00”S, 145°34’00”E],  1000 m,  3.iii–20.iv.1988, R.I. Storey& Dickinson(25-058281 1 specimenin ANIC);  Tully Falls State Forest,  9.5 kmSSW Ravenshoe[ 17°47’00”S, 145°34’00”E],  1000 m,  7.xii.1987–  7.i.1988, R.I. Storey& Dickinson(25-058278 2 specimensin ANIC);  Tully Falls State Forest,  9.5 kmSSW Ravenshoe[ 17°47’00”S, 145°34’00”E],  1000 m,  7.i–9.ii.1988, R.I. Storey& Dickinson(25-058273 3 specimensin ANIC);  Tully Falls State Forest,  9.5 kmSSW Ravenshoe[ 17°47’00”S, 145°34’00”E],  1000 m,  9.ii–3.iii.1988, R.I. Storey& Dickinson(25-058277 1 specimenin ANIC);  Tully Falls State Forest, via Ravenshoe[ 17°47’00”S, 145°34’00”E],  13.v.1974, R.I. Storey(25-032536 1 specimenin QDPC, 25- 058510 1 specimenin QM);  Tully Falls State Forest, via Ravenshoe[ 17°47’00”S, 145°34’00”E],  20–21.iv.1973, R.I. Storey(25-032537 1 specimenin QDPC, 25-058509 11 specimensin QM);  Tully Falls State Forest[ 17°47’00”S, 145°34’00”E],  11–12.xi.1976, R.I. Storey(25-058271 4 specimensin ANIC);  Tully Falls,  30 miNE Tully[ 17°47’00”S, 145°34’00”E],  2500 feet,  5.vi.1972, G.B. & S.R. Monteith(25-058505 1 specimen, 25-058506 1 specimen, 25-058507 1 specimen, 25-058508 1 specimenin QM[ paratypesof  Lepanus nitidus]);  Tully Falls, Ravenshoe[ 17°47’00”S, 145°34’00”E],  20.xi.1977–  16.iv.1978, R.I. Storey(25-032547 1 specimenin QDPC).   Description.Colour ranging from uniformly light brown to shiny dark brown or with pronotum having orange tinges. Holotypeis shiny dark brown with mainly orange pronotum. Antennal club with apical antennomere (3 rd) yellowish white and basal 2 antennomeres (1st, 2nd) grey. Total length: 3.3–3.8 mm. Holotypemeasurements (taken from image): total length 3.8 mm; elytral width 2.6 mm.  Male.Head: Width to length ratio 29: 21. Broad V-shape between clypeal teeth. Surface without setae and finely punctate. Dorsal part of eyes moderate, separated by an interocular space approximately 10–11 times eye width (21: 2). Prothorax: Pronotum smooth, nitid, with very fine punctation towards the anterior and finely punctate posteriorly. Hypomeral surface finely reticulate. Hypomeral stria absent. Pronotum width to length ratio 47: 28. Elytra: Smooth and nitid. Intervals very finely punctate and with extremely fine setae. Apical area nitid with striae 1 and usually 2 superficial, appearing as a single line. Ratio of length of the elytra along suture to maximum elytral width 58: 64. Legs: as per  L. nitidusspecies group. Ridge on ventral side of protibia not joined to second tooth by a fine carina. Abdomen: Rest of pygidium surrounding central disc with fine punctures, each bearing a seta; finely reticulate only towards the basal angles, and nitid otherwise. Pterothorax: Lateral lobe of metaventrite finely punctate, setose and more or less finely reticulate. Medial lobe of metaventrite very finely punctate, smooth and nitid between metacoxae and with a very small tubercle, located closer to metacoxae than mesocoxae. Male genitalia: Aedeagus with left paramere excised apically.  Female.As per  L. nitidusspecies group. Metaventral tubercle absent.   Distribution.Known from rainforest of the Australian Wet Tropics (WET) IBRA bioregion [Zones 7, 8, 9, 10, 11, and 12] from around Danbulla to Ravenshoe ( Fig. 17A), with all specimens from high elevations ( 650–1150 m). Within the BIOCLIM model ( Fig. 17C), there are some grids with suitable climate from which  L. dichroushas not yet been recorded, however the absence of records may reflect the limited sampling due to accessibility of highelevation forests in these grids.   Comments.Most specimens have been collected in dung or mushroom-baited pitfall traps or in flight intercept traps.  Lepanus dichrouswas one of the most abundant species collected in the food preference study of Ebert et al.(2019)with 203 specimenscollected from all lures except both fermenting fruit baits. Both macropod and fowl dung baits collected approximately 25% of the total specimens each ( n =49 and 52 respectively), while vertebrate carrion ( n =26) and invertebrate carrion ( n =22, 9, 21) were more attractive than cucumber ( n =12) and mushroom ( n =12) ( Ebert et al.2019). These results support the generalist feeding habits of the species. Specimens from Charmillin Creek, Millaa Millaa Falls, Palmerston National Park, and Mount Hypipamee National Park were sequenced for this study (see Fig. 1). They form a close relationship with each other, but the relationship within the  L. nitidusspecies group could not be resolved. In Bayesian analysis, it was recovered at the base of the clade containing  L. vangerweniand  L. carbinensiswhile in maximum likelihood analysis it was recovered at the base of the clade containing  L. nitidusand  L. kulki.Neither relationship was strongly supported (BA PP = 81, ML BS = 28). 3034608737 NHRS -17.35 21 145.58333 50 51 JLKB 000027315 3 holotype 3034609302 T55406, T55421 1981-09-15 1981-09-16 1981-09-15 QM -17.088612 North Bell Peak 21 145.87889 50 51 16 Queensland 3034609035 T55422 [315,1359,330,357] 1981-09-16 QM -17.088612 North Bell Peak 21 145.87889 50 51 1 Queensland 3034608313 T55423, T55435, T55835 1990-11-19 1990-11-22 1990-11-19 QM -17.088612 North Bell Peak 21 145.87889 50 51 14 Queensland 3034608560 [151,1432,402,429] 1996-01-17 ANIC R. L. Kitching -17.116667 Atherton 21 145.61667 Robson Creek 50 51 1 Queensland 3034608736 1996-01-18 ANIC R. J. Kitching & H. Mitchell -17.116667 Atherton 21 145.61667 Robson Creek 50 51 1 Queensland 3034608997 ANIC -17.133333 Robson Creek 21 145.63333 50 51 2 Queensland 3034608571 1987-02-13 1987-03-06 1987-02-13 QDPC R. I. Storey & De Faveri -17.166666 Danbulla Forestry Reserve 21 145.63333 50 51 1 Queensland 3034609284 1987-01-13 1987-02-13 1987-01-13 ANIC R. I. Storey & De Faveri -17.166666 Danbulla Forestry Reserve 21 145.63333 50 51 2 Queensland 3034609226 1987-03-26 1987-04-23 1987-03-26 ANIC R. I. Storey & De Faveri -17.166666 Danbulla Forestry Reserve 21 145.63333 50 51 1 Queensland 3034608360 1987-07-28 1987-09-03 1987-07-28 ANIC R. I. Storey & De Faveri -17.166666 Danbulla Forestry Reserve 21 145.63333 50 51 1 Queensland 3034608547 1987-03-06 1987-03-26 1987-03-06 ANIC R. I. Storey & De Faveri -17.166666 Danbulla Forestry Reserve 21 145.63333 50 51 2 Queensland 3034608720 T55306 1995-12-10 1996-02-07 1995-12-10 QM -17.185833 Danbulla Scientific Reserve 21 145.65999 50 51 1 Queensland 3034609059 2012-01-23 2012-01-24 2012-01-23 ANIC G. B. Monteith 750 -17.356112 Malanda Falls 21 145.58694 50 51 41 Queensland 3034608652 T70869, T70877 1998-11-28 1998-11-30 1998-11-28 QM -17.377222 Gourka 21 145.74278 Bartle Frere 50 51 9 Queensland 3034609392 2001-11-19 2001-11-20 2001-11-19 QM -17.379723 Bartle Frere 21 145.76416 50 51 16 Queensland 3034609170 T55845, T55857 1994-11-25 1995-01-10 1994-11-25 QM -17.379723 Bartle Frere 21 145.76416 50 51 13 Queensland 3034609232 T55858, T55864 1995-01-10 1995-03-31 1995-01-10 QM -17.379723 Bartle Frere 21 145.76416 50 51 7 Queensland 3034608688 T55865, T55889 1995-03-07 1995-05-15 1995-03-07 QM -17.379723 Bartle Frere 21 145.76416 50 51 25 Queensland 3034609366 T70860, T70868 1998-11-28 1998-11-30 1998-11-28 QM -17.379723 Bartle Frere 21 145.76416 50 51 9 Queensland 3034609396 1998-11-28 ANIC G. B. Monteith 750 -17.383333 Bartle Frere 21 145.76666 50 51 596 Queensland 3034609160 1998-11-28 1998-11-29 1998-11-28 ANIC G. B. Monteith 750 -17.383333 Bartle Frere 21 145.76666 50 51 2 Queensland 3034608640 T75465, T75470, T75485 1999-02-06 1999-02-09 1999-02-06 QM -17.390833 Malanda 21 145.5961 50 51 7 Queensland 3034609388 [580,992,1195,1221] QM Frith Pei Road Topaz 50 51 1 Queensland 3034609324 T55220, T55226 1993-12-06 1994-02-25 1993-12-06 QM -17.40639 21 145.68472 50 51 7 Queensland 3034609250 2010-09-18 2010-09-19 2010-09-18 ANIC G. B. Monteith 980 -17.43 The Crater National Park 21 145.49 50 51 30 Queensland 3034608839 2012-01-22 2012-01-23 2012-01-22 ANIC G. B. Monteith 980 -17.43 The Crater National Park 21 145.49 50 51 20 Queensland 3034608874 2012-01-21 2012-01-23 2012-01-21 ANIC G. B. Monteith 990 -17.431944 Thiaki 21 145.51611 50 51 1 Queensland 3034609269 2012-01-19 2012-01-23 2012-01-19 ANIC G. B. Monteith 1000 -17.433887 Thiaki 21 145.51111 50 51 2 Queensland 3034608824 1982-07-28 ANIC S. Peck & J. Peck 720 -17.433332 Topaz National Park 21 145.71666 50 51 1 Queensland 3034609182 [558,620,1483,1509] Hugh 50 51 1 Queensland 3034608990 1995-03-06 1995-04-04 1995-03-06 ANIC P. Zborowski 1150 -17.45 Range 21 145.48334 50 51 1 Nelson 3034608769 T75487 1999-02-03 1999-02-06 1999-02-03 QM -17.465557 Longlands Gap 21 145.47499 50 51 1 Nelson 3034608429 1994-12-01 1995-01-03 1994-12-01 ANIC P. Zborowski 1150 -17.466667 Longlands Gap 21 145.48334 50 51 BS1 1 Nelson 3034608410 1996-02-27 1996-03-26 1996-02-27 ANIC P. Zborowski 1150 -17.466667 Longlands Gap 21 145.48334 50 51 BS1 1 Nelson 3034608855 T55296 1994-11-25 1995-01-10 1994-11-25 QM -17.470278 Atherton Tablelands 21 145.53445 Kenny Road 50 51 1 Nelson 3034608786 T55297, T55301 1995-01-10 1995-03-31 1995-01-10 QM -17.470278 Atherton Tablelands 21 145.53445 Kenny Road 50 51 5 Nelson 3034608892 T55302, T55303 1995-03-07 1995-05-15 1995-03-07 QM -17.470278 Atherton Tablelands 21 145.53445 Kenny Road 50 51 2 Nelson 3034608506 T57499, T60022, T60026 1998-02-05 1998-02-11 1998-02-05 QM -17.497778 Evelyn 21 145.47556 Tumoulin Road 50 51 6 Nelson 3034609436 T55230 1983-05-05 QM -17.498056 Millaa Millaa Falls 21 145.6089 50 51 1 Nelson 3034609003 2000-12-08 2000-12-10 2000-12-08 QM -17.498056 Millaa Millaa Falls 21 145.6089 50 51 1 Nelson 3034609249 T75457, T75464 1999-02-06 1999-02-09 1999-02-06 QM -17.498056 Millaa Millaa Falls 21 145.6089 50 51 8 Nelson 3034609203 2012-01-23 2012-01-25 2012-01-23 ANIC G. B. Monteith 830 -17.498056 Millaa Millaa Falls 21 145.61307 50 51 73 Nelson 3034608995 2012-01-23 2012-01-25 2012-01-23 ANIC G. B. Monteith 830 -17.49889 Millaa Millaa Falls 21 145.61389 50 51 75 Nelson 3034608787 1999-04-18 1999-04-20 1999-04-18 ANIC C. Reid & I. Reid 800 51 52 -17.5 Millaa Millaa Falls 21 145.6 50 51 4 Nelson 3034609115 1974-05-11 QM R. I. Storey 780 -17.5 Millaa Millaa Falls 21 145.6 51 52 1 Nelson 3034608639 2012-01-23 2012-01-25 2012-01-23 ANIC G. B. Monteith 780 -17.5 Millaa Millaa Falls 21 145.6 51 52 31 Nelson 3034608347 1973-04-21 1973-04-22 1973-04-21 QDPC, QM R. I. Storey -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 7 Nelson 3034609074 2010-09-15 2010-09-16 2010-09-15 ANIC G. B. Monteith 830 -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 42 Nelson 3034608630 1990-04-10 1990-05-04 1990-04-10 QDPC R. I. Storey & K. H. Halfpapp 760 -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 2 Nelson 3034609449 1990-03-13 1990-04-10 1990-03-13 QDPC R. I. Storey & K. H. Halfpapp 760 -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 1 Nelson 3034609036 1990-08-15 1990-09-12 1990-08-15 QDPC R. I. Storey & K. H. Halfpapp 760 -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 3 Nelson 3034608829 1990-01-04 1990-02-07 1990-01-04 QDPC R. I. Storey & K. H. Halfpapp 760 -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 2 Nelson 3034608964 1990-02-07 1990-03-13 1990-02-07 QDPC R. I. Storey & K. H. Halfpapp 760 -17.5 Millaa Millaa 21 145.61667 Millaa Millaa Falls 51 52 3 Nelson 3034608628 T55317 1994-11-25 1995-01-10 1994-11-25 QM -17.5325 Kjellberg Road 21 145.60751 51 52 1 Nelson 3034609437 T55318, T55328 1995-03-07 1995-05-15 1995-03-07 QM -17.5325 Kjellberg Road 21 145.60751 51 52 11 Nelson 3034608703 T55329, T55338 1995-01-10 1995-03-31 1995-01-10 QM -17.5325 Kjellberg Road 21 145.60751 51 52 10 Nelson 3034608392 T75443, T75446, T75451, T75456 1999-02-06 1999-02-09 1999-02-06 QM -17.5325 Kjellberg Road 21 145.60751 51 52 10 Nelson 3034609178 1999-02-06 1999-02-09 1999-02-06 ANIC G. B. Monteith & D. Cook 920 -17.533333 State Forest 21 145.58333 Kjellberg Road 51 52 11 Nelson 3034609248 T55339, T55340 1993-12-01 1994-02-25 1993-12-01 QM -17.542778 Kjellberg Road 21 145.55861 Mount Fisher 51 52 2 Nelson 3034608745 T75436, T75442, T75447, T75450 1999-02-06 1999-02-09 1999-02-06 QM -17.535833 State Forest 21 145.5775 Kjellberg Road 51 52 11 Nelson 3034609139 2004-09-09 2004-09-14 2004-09-09 CMNC L. Masner 500 -17.568335 Wooroonooran National Park 21 145.70583 51 52 2 Nelson 3034609273 1973-04-22 1973-04-24 1973-04-22 QM, CMNC R. I. Storey -17.566668 Palmerston National Park 21 145.75 Henrietta Creek 51 52 5 Nelson 3034609263 T55304, T55305 1993-12-01 1994-02-25 1993-12-01 QM -17.57639 Beatrice River 21 145.69028 51 52 2 Nelson 3034608831 2010-09-15 2010-09-16 2010-09-15 ANIC G. B. Monteith 650 -17.57889 Scenic Road 21 145.69806 Palmerston 51 52 15 Nelson 3034608758 2010-09-15 2010-09-16 2010-09-15 ANIC G. B. Monteith 700 -17.58889 Palmerston 21 145.70389 51 52 16 Nelson 3034609026 T55104, T55133 1995-03-07 1995-05-15 1995-03-07 QM -17.589724 Ravenshoe 21 145.59444 Maalan State Forest 51 52 30 Nelson 3034609350 T55134, T55152 1995-01-10 1995-03-31 1995-01-10 QM -17.589724 Ravenshoe 21 145.59444 Maalan State Forest 51 52 19 Nelson 3034608341 T55153, T55186 1994-11-25 1995-01-10 1994-11-25 QM -17.589724 Ravenshoe 21 145.59444 Maalan State Forest 51 52 34 Nelson 3034608980 T75471, T75484 1999-02-06 1999-02-09 1999-02-06 QM -17.589724 Ravenshoe 21 145.59444 Maalan State Forest 51 52 14 Nelson 3034609414 T55079, T55089, T55103 1995-03-07 1995-05-15 1995-03-07 QM -17.59889 Maalan Road 21 145.69945 51 52 1 Nelson 3034608332 T55090, T55092 1995-03-01 1995-03-07 1995-03-01 QM -17.59889 Maalan Road 21 145.69945 51 52 3 Nelson 3034609058 T55093, T55102 1994-11-26 1995-01-10 1994-11-26 QM -17.59889 Maalan Road 21 145.69945 51 52 10 Nelson 3034609404 1992-11-30 1993-04-15 1992-11-30 QM -17.599445 Palmerston National Park 21 145.75778 Henrietta Creek 51 52 1 Nelson 3034609096 T55836, T55839 1993-12-01 1994-02-25 1993-12-01 QM -17.56 Palmerston National Park 21 145.6875 51 52 4 Nelson 3034609069 T55840, T55844 1988-12-06 1988-12-14 1988-12-06 QM 800 -17.599445 Mount Father Clancy 21 145.63445 51 52 5 Nelson 3034609229 1978-04-23 1978-04-24 1978-04-23 QM, CMNC R. I. Storey -17.6 Maalan Palmerston National Park 21 145.66667 51 52 18 Nelson 3034608779 1982-06-24 1982-08-02 1982-06-24 ANIC S. Peck & J. Peck 720 -17.6 Millaa Millaa 21 145.71666 51 52 4 Nelson 3034608607 1968-04-01 ANIC E. Matthews -17.616667 Palmerston National Park 21 145.78334 51 52 4 Nelson 3034608842 T55214, T55217 1993-12-01 1994-02-25 1993-12-01 QM -17.636667 South Johnstone Forestry Camp 21 145.67084 51 52 4 Nelson 3034609316 T75494, T76145, T76152 1999-02-04 1999-02-06 1999-02-04 QM -17.682777 Majors Mountain 21 145.53389 Vine Creek 51 52 3 Nelson 3034608546 T79112 1999-04-15 1999-04-18 1999-04-15 QM -17.682777 Majors Mountain 21 145.53389 Vine Creek 51 52 1 Nelson 3034609195 T70886, T70888 1998-12-02 1998-12-05 1998-12-02 QM -17.700834 Charmillin Creek 21 145.52249 51 52 3 Nelson 3034608590 T60017, T60021, T60425, T60433 1998-02-05 1998-02-11 1998-02-05 QM -17.700834 Charmillin Creek 21 145.52249 51 52 14 Nelson 3034609333 2010-09-18 2010-09-19 2010-09-18 ANIC G. B. Monteith 880 -17.715 Charmillin Creek 21 145.52611 51 52 2 Nelson 3034608943 1987-11-15 1987-12-12 1987-11-15 ANIC A. Walford-Huggins -17.716667 Tully Falls 21 145.55 Ravenshoe State Forest 51 52 2 Nelson 3034608572 1987-10-01 1987-11-05 1987-10-01 ANIC, QDPC R. I. Storey & Dickinson 730 -17.766666 Tully Falls State Forest 21 145.51666 51 52 6 Nelson 3034609022 1988-04-20 1988-05-15 1988-04-20 ANIC, QDPC R. I. Storey & Dickinson 730 52 53 -17.766666 Tully Falls State Forest 21 145.51666 51 52 1 Nelson 3034608953 1988-04-20 1988-05-25 1988-04-20 ANIC R. I. Storey & Dickinson 730 -17.766666 Tully Falls State Forest 21 145.51666 52 53 2 Nelson 3034609043 1988-03-03 1988-04-20 1988-03-03 ANIC R. I. Storey & Dickinson 730 -17.766666 Tully Falls State Forest 21 145.51666 52 53 2 Nelson 3034609033 1988-01-05 1988-02-07 1988-01-05 ANIC R. I. Storey & Dickinson 730 -17.766666 Tully Falls State Forest 21 145.51666 52 53 2 Nelson 3034609186 1987-12-07 1988-01-07 1987-12-07 ANIC R. I. Storey & Dickinson 730 -17.766666 Tully Falls State Forest 21 145.51666 52 53 1 Nelson 3034608579 1987-11-07 1987-12-07 1987-11-07 ANIC R. I. Storey & Dickinson 730 -17.766666 Tully Falls State Forest 21 145.51666 52 53 1 Nelson 3034609264 1988-01-05 1988-02-07 1988-01-05 ANIC R. I. Storey & Dickinson 900 -17.783333 Tully Falls State Forest 21 145.56667 52 53 1 Nelson 3034608830 1988-11-05 1988-12-07 1988-11-05 ANIC R. I. Storey & Dickinson 900 -17.783333 Tully Falls State Forest 21 145.56667 52 53 2 Nelson 3034608650 1988-03-03 1988-04-20 1988-03-03 ANIC R. I. Storey & Dickinson 1000 -17.783333 Tully Falls State Forest 21 145.56667 52 53 1 Nelson 3034608657 1987-12-07 1988-01-07 1987-12-07 ANIC R. I. Storey & Dickinson 1000 -17.783333 Tully Falls State Forest 21 145.56667 52 53 2 Nelson 3034608497 1988-01-07 1988-02-09 1988-01-07 ANIC R. I. Storey & Dickinson 1000 -17.783333 Tully Falls State Forest 21 145.56667 52 53 3 Nelson 3034608761 1988-02-09 1988-03-03 1988-02-09 ANIC R. I. Storey & Dickinson 1000 -17.783333 Tully Falls State Forest 21 145.56667 52 53 1 Nelson 3034608967 1974-05-13 QDPC, QM R. I. Storey -17.783333 Ravenshoe 21 145.56667 Tully Falls State Forest 52 53 2 Nelson 3034608942 1973-04-20 1973-04-21 1973-04-20 QDPC, QM R. I. Storey -17.783333 Ravenshoe 21 145.56667 Tully Falls State Forest 52 53 12 Nelson 3034609009 1976-11-11 1976-11-12 1976-11-11 ANIC R. I. Storey -17.783333 Tully Falls State Forest 21 145.56667 52 53 4 Nelson 3034609371 1972-06-05 QM G. B. & S. R. Monteith 762 -17.783333 Tully Falls 21 145.56667 52 53 4 Nelson paratype 3034608680 1977-11-20 1978-04-16 1977-11-20 QDPC R. I. Storey -17.783333 Ravenshoe 21 145.56667 Tully Falls 52 53 1 Nelson paratype