Columbella ( Anachis ) Moravica
Hoernes & Auinger 1880: 101
Columbella ( Macrurella ) moravica Hö. Au.
Boettger 1906: 20
Mitrella ( Macrurella ) moravica ( Hoernes und Auinger 1880 )
Columbella ( Macrurella ) moravica Hoernes und Auinger 1880
The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group
Harzhauser, Mathias
Landau, Bernard M.
Zootaxa
2021
2021-08-23
5025
1
1
75
8ND67
(Hoernes & Auinger, 1880)
Harzhauser & Landau
2021
Hoernes & Auinger
1880
[151,741,330,357]
Gastropoda
Columbellidae
Orthurella
CoL
Animalia
Neogastropoda
38
39
Mollusca
species
moravica
comb. nov.
Figs 4K, 13D 1–D 2, E 1–E 2, F 1–F 2, I [ Anachis] Moravicanov. form.—Hoernes 1880: 127 [ nomen nudum].
* Columbella( Anachis) Moravicanov. form.— Hoernes & Auinger 1880: 101, pl. 12, figs 1–2. Columbella( Macrurella) moravicaHö. Au.var.— Boettger 1906: 20. Mitrella( Macrurella) moravica( Hoernes und Auinger 1880)— Kojumdgieva & Strachimirov 1960: 185, pl. 45, fig. 11. Columbella( Macrurella) moravica Hoernes und Auinger 1880— Strausz 1966: 295, pl. 42, figs 4–5.
Type material. Lectotype(designated herein): NHMW1861/0011/0519a, SL: 17.1 mm, MD: 6.9 mm, Jaroměřice nad Rokytnou(the Czech Republic), illustrated in Hoernes & Auinger (1880, pl. 12, figs 1a–c), Figs 13E 1–E 2. Paralectotypes: NHMW1861/0011/0519b, SL: 14.9 mm, MD: 6.8 mm, Jaroměřice nad Rokytnou(the Czech Republic), illustrated in Hoernes & Auinger (1880, pl. 12, figs 2a–c), Figs 13D 1–D 2. NHMW2020/0159/0001, SL: 19.7 mm, MD: 7.4 mm, Jaroměřice nad Rokytnou(the Czech Republic), Figs 4K, 13F 1–F 2. NHMW2020/0159/0002, SL: 16.1 mm, MD: 7.1 mm, Jaroměřice nad Rokytnou(the Czech Republic); Fig. 13I. NHMW1864/0001/0326, 12 spec., Jaroměřice nad Rokytnou(the Czech Republic); NHMW2020/0160/0001, 1 spec., Boskovice(the Czech Republic).
Description.Shell medium sized, moderately broad fusiform, with strongly gradate spire and broad sutural shelf. Protoconch large, conical, multispiral of 3.5 convex whorls. Teleoconch of eight whorls. Early spire narrowly conical with moderately convex whorls and narrowly incised suture. Spire whorls becoming subcylindrical and rapidly increasing in width within 6 to 8 teleoconch whorls, resulting in broad sutural shelf bordered by crested shoulder. Last whorl subcylindrical, weakly keeled at periphery, strongly constricted, with high base. Sculpture of delicate spiral threads on entire teleoconch; numerous broad spiral cords with narrow interspaces on abapical portion of base and siphonal canal. Aperture narrow, elongate. Outer lip weakly thickened with about nine small weak denticles within. Anal canal narrowly incised, V-shaped. Siphonal canal very long, narrow, deflected. Columellatwisted, forming weak columellar fold. Columellar callus narrow, thin, adherent, without denticles. Parietal callus slightly thickened. Shell measurements and ratios.SL: 14.9–19.7 mm, MD: 6.9–7.4 mm, AA: 35–41°, SL/MD: 2.5–2.7, AL/AW: 4.8–5.0, LWH/AH: 1.2–1.3.
Discussion.This species is characterised by its strikingly allometric growth. The early spire morphology, however, is identical with Orthurella convexula, which develops a similar protoconch. In addition, both species develop an extraordinarily long siphonal canal. Therefore, we place Columbella( Anachis) moravicain Orthurella. The broad subhorizontal sutural shelf immediately distinguishes Orthurella moravica( Hoernes & Auinger, 1880)from O. convexula. Palaeoenvironment.Unknown.
Distribution in Central Paratethys.Badenian (middle Miocene): Carpathian Foredeep: Jaroměřice nad Rokytnou, Boskovice (the Czech Republic); Pannonian Basin: Szob( Hungary) ( Strausz 1966); Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906); Dacian Basin: Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960). Genus ClinurellaSacco, 1890
Typespecies. Columbella( Clinurella) scalarisSacco 1890(in Sacco 1890b); subsequent designation by Pace (1902: 43). Early Miocene, Italy.
Discussion. Clinurellacomprise medium sized, moderately broad fusiform shells with high to moderately high spire and convex, strongly constricted last whorl with long, narrow, slightly deflected siphonal canal. A flaring outer lip with prominent denticles and prominent spiral cords on the base and on the exterior part of the outer lip are characteristic. Sacco (1890b: 44)established Clinurellaas a subgenus of Columbellaand listed Columbella (Clinurella) scalaris(Sacco, 1890), Columbella( Clinurella) albucianensisSacco, 1890, Columbella borsoni Bellardi, 1848and Columbella( Clinurella) vialensisSacco, 1890in his new subgenus. Columbella borsoni, C. minimaand C. vialensishave relatively short siphonal canals and only moderately flaring apertures, if at all, and are placed herein in Mitrella(see also Landau et al.2013: 189). Therefore, we place only Clinurella scalaris, C. albucianensisand C. sophiae( Boettger, 1902)in Clinurella. Cossmann (1901: 242)synonymised Clinurellawith AtiliaH. Adams & A. Adams, 1853(= AnachisH. Adams & A. Adams, 1853). Anachis scalarina(G. B. Sowerby I, 1832), typespecies of Anachis, and Anachis suffusa(G.B. Sowerby I, 1844), typespecies of Atilia, differ clearly from Clinurellain their short siphonal canal and simple outer lips and the synonymy proposed by Cossmann (1901)is rejected herein. Clinurellais reminiscent of OrthurellaSacco, 1890but is distinguished by its broader shell, thickened and flaring outer lip and prominent denticulation on the columellar edge and the outer lip. In Orthurellacolumellar denticles are absent, any sculpture resulting from the spiral cords over the base showing through the columellar callus. Clinurellais known so far only from the Burdigalian and Langhian of the Proto-Mediterranean Sea and the Langhian (Badenian) of the Central Paratethys Sea.
NHMW
Czech Republic
Jaromerice nad Rokytnou
38
39
NHMW1861
1
lectotype
NHMW
Czech Republic
Jaromerice nad Rokytnou
38
39
NHMW1861
1
paralectotype
NHMW
Czech Republic
Jaromerice nad Rokytnou
38
39
NHMW2020
1
paralectotype
NHMW
Czech Republic
Jaromerice nad Rokytnou
38
39
NHMW2020
1
paralectotype
NHMW
Czech Republic
Jaromerice nad Rokytnou
38
39
NHMW1864
12
paralectotype
NHMW
Czech Republic
Boskovice
38
39
NHMW2020
1
paralectotype