Columbella ( Anachis ) Moravica Hoernes & Auinger 1880: 101 Columbella ( Macrurella ) moravica Hö. Au. Boettger 1906: 20 Mitrella ( Macrurella ) moravica ( Hoernes und Auinger 1880 ) Columbella ( Macrurella ) moravica Hoernes und Auinger 1880 The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group Harzhauser, Mathias Landau, Bernard M. Zootaxa 2021 2021-08-23 5025 1 1 75 8ND67 (Hoernes & Auinger, 1880) Harzhauser & Landau 2021 Hoernes & Auinger 1880 [151,741,330,357] Gastropoda Columbellidae Orthurella CoL Animalia Neogastropoda 38 39 Mollusca species moravica comb. nov.   Figs 4K, 13D  1–D  2, E  1–E  2, F  1–F  2, I [  Anachis] Moravicanov. form.—Hoernes 1880: 127 [ nomen nudum].    *  Columbella( Anachis) Moravicanov. form.—  Hoernes & Auinger 1880: 101, pl. 12, figs 1–2.    Columbella( Macrurella) moravicaHö. Au.var.—  Boettger 1906: 20.   Mitrella( Macrurella) moravica( Hoernes und Auinger 1880)— Kojumdgieva & Strachimirov 1960: 185, pl. 45, fig. 11.  Columbella( Macrurella) moravica Hoernes und Auinger 1880— Strausz 1966: 295, pl. 42, figs 4–5.   Type material.  Lectotype(designated herein): NHMW1861/0011/0519a, SL: 17.1 mm, MD: 6.9 mm, Jaroměřice nad Rokytnou(the Czech Republic), illustrated in Hoernes & Auinger (1880, pl. 12, figs 1a–c), Figs 13E  1–E  2.  Paralectotypes: NHMW1861/0011/0519b, SL: 14.9 mm, MD: 6.8 mm, Jaroměřice nad Rokytnou(the Czech Republic), illustrated in Hoernes & Auinger (1880, pl. 12, figs 2a–c), Figs 13D  1–D  2.  NHMW2020/0159/0001, SL: 19.7 mm, MD: 7.4 mm, Jaroměřice nad Rokytnou(the Czech Republic), Figs 4K, 13F  1–F  2.  NHMW2020/0159/0002, SL: 16.1 mm, MD: 7.1 mm, Jaroměřice nad Rokytnou(the Czech Republic); Fig. 13I.  NHMW1864/0001/0326, 12 spec., Jaroměřice nad Rokytnou(the Czech Republic);  NHMW2020/0160/0001, 1 spec., Boskovice(the Czech Republic).   Description.Shell medium sized, moderately broad fusiform, with strongly gradate spire and broad sutural shelf. Protoconch large, conical, multispiral of 3.5 convex whorls. Teleoconch of eight whorls. Early spire narrowly conical with moderately convex whorls and narrowly incised suture. Spire whorls becoming subcylindrical and rapidly increasing in width within 6 to 8 teleoconch whorls, resulting in broad sutural shelf bordered by crested shoulder. Last whorl subcylindrical, weakly keeled at periphery, strongly constricted, with high base. Sculpture of delicate spiral threads on entire teleoconch; numerous broad spiral cords with narrow interspaces on abapical portion of base and siphonal canal. Aperture narrow, elongate. Outer lip weakly thickened with about nine small weak denticles within. Anal canal narrowly incised, V-shaped. Siphonal canal very long, narrow, deflected. Columellatwisted, forming weak columellar fold. Columellar callus narrow, thin, adherent, without denticles. Parietal callus slightly thickened.  Shell measurements and ratios.SL: 14.9–19.7 mm, MD: 6.9–7.4 mm, AA: 35–41°, SL/MD: 2.5–2.7, AL/AW: 4.8–5.0, LWH/AH: 1.2–1.3.   Discussion.This species is characterised by its strikingly allometric growth. The early spire morphology, however, is identical with  Orthurella convexula, which develops a similar protoconch. In addition, both species develop an extraordinarily long siphonal canal. Therefore, we place  Columbella( Anachis) moravicain  Orthurella. The broad subhorizontal sutural shelf immediately distinguishes  Orthurella moravica( Hoernes & Auinger, 1880)from  O. convexula.  Palaeoenvironment.Unknown.   Distribution in Central Paratethys.Badenian (middle Miocene): Carpathian Foredeep: Jaroměřice nad Rokytnou, Boskovice (the Czech Republic); Pannonian Basin: Szob( Hungary) ( Strausz 1966); Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906); Dacian Basin: Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).  Genus  ClinurellaSacco, 1890    Typespecies.  Columbella( Clinurella) scalarisSacco 1890(in Sacco 1890b); subsequent designation by Pace (1902: 43). Early Miocene, Italy.   Discussion.  Clinurellacomprise medium sized, moderately broad fusiform shells with high to moderately high spire and convex, strongly constricted last whorl with long, narrow, slightly deflected siphonal canal. A flaring outer lip with prominent denticles and prominent spiral cords on the base and on the exterior part of the outer lip are characteristic.  Sacco (1890b: 44)established  Clinurellaas a subgenus of  Columbellaand listed  Columbella (Clinurella) scalaris(Sacco, 1890),  Columbella( Clinurella) albucianensisSacco, 1890,  Columbella borsoni Bellardi, 1848and  Columbella( Clinurella) vialensisSacco, 1890in his new subgenus.  Columbella borsoni,  C. minimaand  C. vialensishave relatively short siphonal canals and only moderately flaring apertures, if at all, and are placed herein in  Mitrella(see also Landau et al.2013: 189). Therefore, we place only  Clinurella scalaris,  C. albucianensisand  C. sophiae( Boettger, 1902)in  Clinurella.  Cossmann (1901: 242)synonymised  Clinurellawith  AtiliaH. Adams & A. Adams, 1853(=  AnachisH. Adams & A. Adams, 1853).  Anachis scalarina(G. B. Sowerby I, 1832), typespecies of  Anachis, and  Anachis suffusa(G.B. Sowerby I, 1844), typespecies of  Atilia, differ clearly from  Clinurellain their short siphonal canal and simple outer lips and the synonymy proposed by Cossmann (1901)is rejected herein.   Clinurellais reminiscent of  OrthurellaSacco, 1890but is distinguished by its broader shell, thickened and flaring outer lip and prominent denticulation on the columellar edge and the outer lip. In  Orthurellacolumellar denticles are absent, any sculpture resulting from the spiral cords over the base showing through the columellar callus.   Clinurellais known so far only from the Burdigalian and Langhian of the Proto-Mediterranean Sea and the Langhian (Badenian) of the Central Paratethys Sea. NHMW Czech Republic Jaromerice nad Rokytnou 38 39 NHMW1861 1 lectotype NHMW Czech Republic Jaromerice nad Rokytnou 38 39 NHMW1861 1 paralectotype NHMW Czech Republic Jaromerice nad Rokytnou 38 39 NHMW2020 1 paralectotype NHMW Czech Republic Jaromerice nad Rokytnou 38 39 NHMW2020 1 paralectotype NHMW Czech Republic Jaromerice nad Rokytnou 38 39 NHMW1864 12 paralectotype NHMW Czech Republic Boskovice 38 39 NHMW2020 1 paralectotype