Columbella curta Dujardin
Columbella ( Columbella ) curta Duj.
Boettger 1902: 16
Columbella ( Mitrella ) curta Duj.
Boettger 1906: 14
Columbella ( Mitrella )
turgidula Brocc.
Columbella
turgidula Brocc.
Friedberg 1938: 128
Columbella ( Alia ) curta var. convexa Friedberg 1911
Kojumdgieva & Strachimirov 1960: 182
Mitrella convexa ( Friedberg, 1911 )
Pyrene ( Mitrella ) korytnicensis
Bałuk 1995: 236
The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group
Harzhauser, Mathias
Landau, Bernard M.
Zootaxa
2021
2021-08-23
5025
1
1
75
8NCF8
(Baluk, 1995)
Harzhauser & Landau
2021
Baluk
1995
[151,682,1120,1147]
Gastropoda
Columbellidae
Graphicomassa
Animalia
Neogastropoda
11
12
Mollusca
species
korytnicensis
comb. nov.
Figs 3G 1–G 2, H 1–H 2, I 1–I 2,J 1–J 2, K, L Columbella curtaBell.—Hörnes 1852: 118 (pars), pl. 11, figs 4–6 [ non Dujardin, 1837].
Columbella curtaDujardin— Hoernes & Auinger 1880(pars): 93, pl. 7, figs 15–19 [ non Dujardin, 1837]. Columbella( Columbella) curtaDuj.— Boettger 1902: 16[ non Dujardin, 1837]. Columbella( Mitrella) curtaDuj.— Boettger 1906: 14[ non Dujardin, 1837]. Columbella( Mitrella)an turgidulaBrocc.—Friedberg 1928: 586, pl. 38, fig. 2. Columbellaan turgidulaBrocc.— Friedberg 1938: 128. Columbella( Alia) curtavar. convexa Friedberg 1911—Kojumdgieva in Kojumdgieva & Strachimirov 1960: 182, pl. 45, fig. 3 [non Mitrella convexa( Friedberg, 1911)]. * Pyrene( Mitrella) korytnicensis sp. n.— Bałuk 1995: 236, pl. 31, figs 11–12, pl. 32, fig. 7.
Type material. Holotype: Institute of Geologyof the University of Warsaw( ZPALUWBkK-G619), SL: 16.0 mm, MD: 8.5 mm, Korytnica, Poland, illustrated in Bałuk (1995, pl. 31, fig. 12). Illustrated material. NHMW1858/0043/0016, SL: 21.4 mm, MD: 11.4 mm, Lăpugiu de Sus( Romania), Figs 3J 1–J 2. NHMW1854/0035/0106, SL: 23.4 mm, MD: 11.2 mm, Lăpugiu de Sus( Romania), Fig. 3K. NHMW1854/ 0035/0167a, SL: 20.1 mm, MD: 10.4 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 15a–b), Figs 3I 1–I 2. NHMW1854/0035/016c7, SL: 18.9 mm, MD: 10.6 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 17a–b), Figs 3G 1–G 2. NHMW1854/0035/0167d, SL: 15.9 mm, MD: 7.9 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 18a–c), Figs 3H 1–H 2; NHMW2020/0153/0001, Lăpugiu de Sus( Romania), Fig 3L. Additional material. NHMW1846/0037/0155a, SL: 15.9 mm, MD: 7.0 mm, Bad Vöslau( Austria), illustrated in Hörnes (1852, pl. 11, fig. 5). NHMW1846/0037/0155b, SL: 13.6 mm, MD: 6.6 mm, Bad Vöslau( Austria), illustrated in Hörnes (1852, pl. 11, fig. 6). NHMW1854/0035/0167b, SL: 20.4 mm, MD: 10.5 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 16a–b). NHMW1854/0035/0167e, SL: 11.4 mm, MD: 5.5 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 19a–c). NHMW1869/0001/0024, 12 spec., Bad Vöslau( Austria); NHMW1868/0001/0066, 5 spec., Baden-Sooss( Austria); NHMW1872/0030/0118, 1 spec., Baden( Austria); NHMW1870/0033/0041, 74 spec., Lăpugiu de Sus( Romania); NHMW2020/0153/0001, 6 spec., Lăpugiu de Sus( Romania); NHMW 2020/0172/0005 (ex B. Landaucoll.), 2 spec., Lăpugiu de Sus( Romania). Revised description.Shell large, robust, broad ovoid, with decollate apex and incised suture. Protoconch unknown. Spire moderately low, conical to cyrtoconoid, weakly gradate. Spire whorls flat-sided to weakly convex with narrow shoulder and periphery at abapical suture. Last whorl high, subcylindrical to weakly convex, rarely slightly pinched below shoulder. Base weakly to moderately constricted. Shell surface smooth except for few weak spiral cords on base.Aperture moderately narrow.Anal canal sharply incised, V-shaped, callused. Outer lip thickened with weakly developed internal labial callus pad in upper third and straight margin, denticulate with about 10–12 small denticles within. Siphonal canal short, wide, moderately deeply notched at tip. Columellar callus strongly thickened forming sharply delimited callus rim, slightly detached over base, abapically thickened with about seven indistinct denticles at outer columellar edge. Callus thins mid-aperture, thickening again in parietal area. Colour pattern of close-set, roughly spirally arranged, small subtrigonal whitish dots on brownish ground. Shell measurements and ratios.SL: 15.9–23.4 mm, MD: 7.0– 11.4 mm, AA: 53–61°, SL/MD: 1.9, AL/AW: 3.5–3.6, LWH/AH: 1.3.
Discussion.This species has been mixed with Graphicomassa polonicaby Hörnes (1852) and Hoernes & Auinger (1880). It differs from G. polonicain its stout outline, cyrtoconoid spire and shorter last whorl. In addition, the colour pattern of G. korytnicensisis delicate, comprising small dots, whereas G. polonicashows broad squarish flammulae. Nevertheless, subadult shells of Graphicomassa polonicacan be difficult to distinguish from G. korytnicensis. Both species seem to have been ecologically separated. Graphicomassa polonicais found in inner neritic silty-sandy deposits whereas G. korytnicensisis found in offshore clays. The stout, thickened shell and cyrtoconoid spire make this species superficially similar to Columbella mayeriHoernes, 1856, but it is readily separated by the absence of the row of inner columellar denticles, diagnostic to the genus (see above). Moreover, the spire is even lower and more strongly cyrtoconoid in C. mayeri. Palaeoenvironment.Middle neritic to outer neritic. The specimens from the ViennaBasin are found in clay of the Baden Fm., suggesting about 250 mwater depth ( Hohenegger et al.2008).
Distribution in Central Paratethys.Badenian (middle Miocene): Carpathian Foredeep:Korytnica ( Poland) ( Bałuk 1995); Zborów ( Ukraine) ( Friedberg 1938); Vienna Basin: Baden, Bad Vöslau, Baden-Sooss (own data); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Hoernes & Auinger 1880; Boettger 1906); Dacian Basin: Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).
ZPALUW
Poland
Korytnica
Institute of Geology
University of Warsaw
11
12
1
holotype
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1858
1
[232,1269,1648,1675]
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1854
1
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1854
1
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1854
1
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1854
1
[151,838,1828,1855]
NHMW
Romania
Lapugiu de Sus
11
12
NHMW2020
3
[449,1313,1864,1891]
NHMW
Austria
Bad Voslau
11
12
NHMW1846
1
[533,1432,1900,1927]
NHMW
Austria
Bad Voslau
11
12
NHMW1846
1
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1854
1
NHMW
Romania
Lapugiu de Sus
11
12
NHMW1854
1
[151,789,151,178]
NHMW
Austria
Bad Voslau
12
13
NHMW1869
12
[801,1432,150,178]
NHMW
Austria
Baden-Sooss
12
13
NHMW1868
5
[151,717,187,214]
NHMW
Austria
Baden
12
13
NHMW1872
1
[729,1432,187,214]
NHMW
Romania
Lapugiu de Sus
12
13
NHMW1870
74
[151,859,222,250]
NHMW
Romania
Lapugiu de Sus
12
13
NHMW2020
6
NHMW
B. Landau
Romania
Lapugiu de Sus
12
13
NHMW 2020
2