Columbella curta Dujardin Columbella ( Columbella ) curta Duj. Boettger 1902: 16 Columbella ( Mitrella ) curta Duj. Boettger 1906: 14 Columbella ( Mitrella ) turgidula Brocc. Columbella turgidula Brocc. Friedberg 1938: 128 Columbella ( Alia ) curta var. convexa Friedberg 1911 Kojumdgieva & Strachimirov 1960: 182 Mitrella convexa ( Friedberg, 1911 ) Pyrene ( Mitrella ) korytnicensis Bałuk 1995: 236 The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group Harzhauser, Mathias Landau, Bernard M. Zootaxa 2021 2021-08-23 5025 1 1 75 8NCF8 (Baluk, 1995) Harzhauser & Landau 2021 Baluk 1995 [151,682,1120,1147] Gastropoda Columbellidae Graphicomassa Animalia Neogastropoda 11 12 Mollusca species korytnicensis comb. nov.   Figs 3G  1–G  2, H  1–H  2, I  1–I  2,J  1–J  2, K, L   Columbella curtaBell.—Hörnes 1852: 118 (pars), pl. 11, figs 4–6 [ non Dujardin, 1837].    Columbella curtaDujardin— Hoernes & Auinger 1880(pars): 93, pl. 7, figs 15–19 [ non Dujardin, 1837].    Columbella( Columbella) curtaDuj.—  Boettger 1902: 16[ non Dujardin, 1837].    Columbella( Mitrella) curtaDuj.—  Boettger 1906: 14[ non Dujardin, 1837].   Columbella( Mitrella)an  turgidulaBrocc.—Friedberg 1928: 586, pl. 38, fig. 2.    Columbellaan  turgidulaBrocc.—  Friedberg 1938: 128.    Columbella( Alia) curtavar. convexa Friedberg 1911—Kojumdgieva in  Kojumdgieva & Strachimirov 1960: 182, pl. 45, fig. 3 [non  Mitrella convexa( Friedberg, 1911)].   *  Pyrene( Mitrella) korytnicensis sp. n.—  Bałuk 1995: 236, pl. 31, figs 11–12, pl. 32, fig. 7.   Type material.  Holotype: Institute of Geologyof the University of Warsaw( ZPALUWBkK-G619), SL: 16.0 mm, MD: 8.5 mm, Korytnica, Poland, illustrated in Bałuk (1995, pl. 31, fig. 12).  Illustrated material.  NHMW1858/0043/0016, SL: 21.4 mm, MD: 11.4 mm, Lăpugiu de Sus( Romania), Figs 3J  1–J  2.  NHMW1854/0035/0106, SL: 23.4 mm, MD: 11.2 mm, Lăpugiu de Sus( Romania), Fig. 3K.  NHMW1854/ 0035/0167a, SL: 20.1 mm, MD: 10.4 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 15a–b), Figs 3I  1–I  2.  NHMW1854/0035/016c7, SL: 18.9 mm, MD: 10.6 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 17a–b), Figs 3G  1–G  2.  NHMW1854/0035/0167d, SL: 15.9 mm, MD: 7.9 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 18a–c), Figs 3H  1–H  2;  NHMW2020/0153/0001, Lăpugiu de Sus( Romania), Fig 3L.  Additional material.  NHMW1846/0037/0155a, SL: 15.9 mm, MD: 7.0 mm, Bad Vöslau( Austria), illustrated in Hörnes (1852, pl. 11, fig. 5).  NHMW1846/0037/0155b, SL: 13.6 mm, MD: 6.6 mm, Bad Vöslau( Austria), illustrated in Hörnes (1852, pl. 11, fig. 6).  NHMW1854/0035/0167b, SL: 20.4 mm, MD: 10.5 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 16a–b).  NHMW1854/0035/0167e, SL: 11.4 mm, MD: 5.5 mm, Lăpugiu de Sus( Romania), illustrated in Hoernes & Auinger (1880, pl. 7, figs. 19a–c).  NHMW1869/0001/0024, 12 spec., Bad Vöslau( Austria);  NHMW1868/0001/0066, 5 spec., Baden-Sooss( Austria);  NHMW1872/0030/0118, 1 spec., Baden( Austria);  NHMW1870/0033/0041, 74 spec., Lăpugiu de Sus( Romania);  NHMW2020/0153/0001, 6 spec., Lăpugiu de Sus( Romania);  NHMW 2020/0172/0005 (ex B. Landaucoll.), 2 spec., Lăpugiu de Sus( Romania).  Revised description.Shell large, robust, broad ovoid, with decollate apex and incised suture. Protoconch unknown. Spire moderately low, conical to cyrtoconoid, weakly gradate. Spire whorls flat-sided to weakly convex with narrow shoulder and periphery at abapical suture. Last whorl high, subcylindrical to weakly convex, rarely slightly pinched below shoulder. Base weakly to moderately constricted. Shell surface smooth except for few weak spiral cords on base.Aperture moderately narrow.Anal canal sharply incised, V-shaped, callused. Outer lip thickened with weakly developed internal labial callus pad in upper third and straight margin, denticulate with about 10–12 small denticles within. Siphonal canal short, wide, moderately deeply notched at tip. Columellar callus strongly thickened forming sharply delimited callus rim, slightly detached over base, abapically thickened with about seven indistinct denticles at outer columellar edge. Callus thins mid-aperture, thickening again in parietal area. Colour pattern of close-set, roughly spirally arranged, small subtrigonal whitish dots on brownish ground.  Shell measurements and ratios.SL: 15.9–23.4 mm, MD: 7.0– 11.4 mm, AA: 53–61°, SL/MD: 1.9, AL/AW: 3.5–3.6, LWH/AH: 1.3.   Discussion.This species has been mixed with  Graphicomassa polonicaby Hörnes (1852) and Hoernes & Auinger (1880). It differs from  G. polonicain its stout outline, cyrtoconoid spire and shorter last whorl. In addition, the colour pattern of  G. korytnicensisis delicate, comprising small dots, whereas  G. polonicashows broad squarish flammulae. Nevertheless, subadult shells of  Graphicomassa polonicacan be difficult to distinguish from  G. korytnicensis. Both species seem to have been ecologically separated.  Graphicomassa polonicais found in inner neritic silty-sandy deposits whereas  G. korytnicensisis found in offshore clays. The stout, thickened shell and cyrtoconoid spire make this species superficially similar to  Columbella mayeriHoernes, 1856, but it is readily separated by the absence of the row of inner columellar denticles, diagnostic to the genus (see above). Moreover, the spire is even lower and more strongly cyrtoconoid in  C. mayeri.  Palaeoenvironment.Middle neritic to outer neritic. The specimens from the ViennaBasin are found in clay of the Baden Fm., suggesting about 250 mwater depth ( Hohenegger et al.2008).   Distribution in Central Paratethys.Badenian (middle Miocene): Carpathian Foredeep:Korytnica ( Poland) ( Bałuk 1995); Zborów ( Ukraine) ( Friedberg 1938); Vienna Basin: Baden, Bad Vöslau, Baden-Sooss (own data); Făget Basin: Lăpugiu de Sus, Coşteiu de Sus ( Romania) ( Hoernes & Auinger 1880; Boettger 1906); Dacian Basin: Târnene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960). ZPALUW Poland Korytnica Institute of Geology University of Warsaw 11 12 1 holotype NHMW Romania Lapugiu de Sus 11 12 NHMW1858 1 [232,1269,1648,1675] NHMW Romania Lapugiu de Sus 11 12 NHMW1854 1 NHMW Romania Lapugiu de Sus 11 12 NHMW1854 1 NHMW Romania Lapugiu de Sus 11 12 NHMW1854 1 NHMW Romania Lapugiu de Sus 11 12 NHMW1854 1 [151,838,1828,1855] NHMW Romania Lapugiu de Sus 11 12 NHMW2020 3 [449,1313,1864,1891] NHMW Austria Bad Voslau 11 12 NHMW1846 1 [533,1432,1900,1927] NHMW Austria Bad Voslau 11 12 NHMW1846 1 NHMW Romania Lapugiu de Sus 11 12 NHMW1854 1 NHMW Romania Lapugiu de Sus 11 12 NHMW1854 1 [151,789,151,178] NHMW Austria Bad Voslau 12 13 NHMW1869 12 [801,1432,150,178] NHMW Austria Baden-Sooss 12 13 NHMW1868 5 [151,717,187,214] NHMW Austria Baden 12 13 NHMW1872 1 [729,1432,187,214] NHMW Romania Lapugiu de Sus 12 13 NHMW1870 74 [151,859,222,250] NHMW Romania Lapugiu de Sus 12 13 NHMW2020 6 NHMW B. Landau Romania Lapugiu de Sus 12 13 NHMW 2020 2