The Columbellidae (Gastropoda, Buccinoidea) in the Miocene Paratethys Sea — striking diversity of a negelected group Harzhauser, Mathias Landau, Bernard M. Zootaxa 2021 2021-08-23 5025 1 1 75 7P4KD Iredale, 1929 Iredale 1929 [237,596,978,1005] Gastropoda Columbellidae Graphicomassa Animalia Neogastropoda 8 9 Mollusca genus    Typespecies.  Columbella ligula Duclos, 1840; original designation by Iredale (1929: 289). Present-day, Australia.   Discussion.We place  Nassa columbelloides polonicaPusch, 1837and  Pyrene korytnicensis Bałuk, 1995in the Indo- West Pacific genus  Graphicomassa, based on their subcylindrical last whorls, presence of a subsutural concavity, giving the adapical portion of the last whorl a slightly pinched aspect, slightly gradate spires and absence of prominent axial sculpture. The absence of an internal row of columellar denticles allows a separation from  Columbella. A common feature of both Miocene species is a decollate early spire and the development of a low, spirally formed plug. The loss of the apex is known from several extant Columbellidae, such as  Mitrella ocellata( Gmelin, 1791)and even in some species of the deep water genus  Astyris, such as  Astyris atacamensis Araya, Catalán & Aliaga, 2016(pers. com. Marta deMaintenon, 7. 11. 2020). An abraded apex is frequently seen in  Graphicomassa ligula(e.g. deMaintenon 2016, figs 1B, 1C, 1E, 1F). The genus was represented in Europe during the early and middle Miocene by several species:  Graphicomassa ringens(Sacco, 1890),  G. scalarata(Sacco, 1891),  G. inflata(Sacco, 1890) (Proto-Mediterranean Sea)and  G. turonica( Mayer, 1869)(northeastern Atlantic) (see Sacco 1890b, 1891; Mayer 1869).