Microcaeculus weyrauchi Franz, 1964: 89–90 Taylor, Gunawardene and Kinnear, 2013: 449 A. weyrauchi, Coineau 1974a: 279 Revision and phylogeny of the genus Andocaeculus (Acari: Caeculidae) I: the A. weyrauchi species group Porta, Andrés O. Pizarro-Araya, Jaime Ramírez, Martín J. Zootaxa 2021 2021-03-17 4945 1 1 78 (Franz, 1964) Franz 1964 [151,630,294,321] Arachnida Caeculidae Andocaeculus GBIF Animalia Acari 6 7 Arthropoda species weyrauchi  ( Figs 2–17)      Microcaeculus weyrauchi Franz, 1964: 89–90;  Taylor, Gunawardene and Kinnear, 2013: 449.     A. weyrauchi, Coineau 1974a: 279.    Typematerial.  Typeseries from “  Puna El Infernilloin  3000 m Seehöhe am Fussevon flechtenbewachsenen Felsblöckenaus Moos, Rasenund Erdeunter diesem am  13. 9. 1963gesiebt”, Herbert Franzcollector, not examined (it should be deposited in MNHN, not found).  Examined material.  4 females(MACN-Ar 41058, 41059, 41135, 41136), Argentina, Provincia de Tucumán, Abrael Infiernillo, S26.74272°, W065.77283°(+/-  50 m), alt.  3012 masl( GPS), under stones, hand collected and on litter of xeric vegetation, extracted with Berlesefunnels, A. Porta leg., cleared with lactic acid, preserved in alcohol;  1 adult(MACN-Ar 41056), same data, not cleared, preserved in alcohol;  2 adults(MACN-Ar 41058), same data. preserved in alcohol;  10 adultsin a vial (MACN-Ar 4134), same data, preserved in alcohol;  1 larva(MACN-Ar 41065), same data, mounted on slide; 4 protonymphs (MACN-Ar 41066 to 41069), same data, mounted on slides; 2 deutonymphs (MACN-Ar 41070 and 41071), same data, mounted on slides; 6 tritonymphs (MACN-Ar 41072 to 41077), same data, mounted on slides;  14 females(MACN-Ar 41060 to 41065, 41126 to 41033), same data, mounted on slides;  1 male(MACN-Ar 41078), same data, mounted on slide; 3 individual stubs with exemplars mounted for SEM (MACN-Ar 41057), same data; 5 individuals, (MACN-Ar 41079 to 41083), vouchers of BOLDSYSTEM data SPDAR1609-16to 1609-20, same data, preserved in alcohol;  2 adultsin a vial (FML-Ar 00286), same data, preserved in alcohol.   Remark. The examined material has been assigned to  A. weyrauchion the basis of being the only species collected in the typelocality and because it exhibits a morphology that corresponds exactly with that of  A. weyrauchiin Franz’s description.   Diagnosis.  A. weyrauchidiffers from other species of the group by the following combination of characters: subtrapezoidal form of the idiosoma ( Figs. 3, 4); anterior margin of aspidosoma straight ( Fig. 5A); palp tarsus with 4 eupathidia; opisthosoma with a relatively less regressive chaetotaxy ( Figs. 3and 4): most setae normal-sized, elongated, b2and c2present in some exemplars, neotrichy sometimes present on d2seta sector ( Fig. 3and 4); with four pairs of setae on Pplates, setae e2and e2’with e2”separated by ridges ( Fig. 4); setae of series v’on basi and telofemur I of adults, elongated, distally blunt, relatively long for the group, 54–66 and 80–96 +long in females, RBaf: 0.52–0.6, RBaf +: 0.75–0.9, RTef: 0.5–0.57, RTef +: 0.75–0.84; genua I and II with 3 and tibiae I and II with 2 regressive setae of series d( Figs. 9Dand 10 D); setae of series v’on genua and tibiae I and II pointed; setae l1’always eupathidial on leg I and II ( Figs. 9A–J); solenidion present on tibia IV ( Figs. 11B, 13 B–C), RTi3: 2.6–4.3.   Redescription  Female (n=14). Color( Fig. 2). Gnathosoma and dorsal sclerotized plates on idiosoma brown, membranous integument pale cream, legs and hypostome dark brown.  Gnathosoma( Figs. 3, 5). Covered by aspidosoma in dorsal view. Chelicera ( Fig. 5A) typical for family, 54–58 wide at base, 132–170 long, movable digit hook-like with 3 minute teeth; cheliceral seta chslender, 6 long. Palp ( Figs. 14B, 15) 4-segmented, trochanter glabrous, femorogenu with 2 clavate setae in dorsal series, d126–40 long, d, 42–54 long; tibia with two calcars l’and d, with don projecting tubercle and 3 clavate setae  d1, ldand l”, setal lengths, l’: 24–30, l”: 30–36, ld: 30–38, d: 26–33, d1: 26–34. Tarsus subconical, 42–54 long, 16–22 wide at base, solenidion ω present on antiaxial face, with 6–7 setae,  v, l1’, l” in addition to 4 eupathidia, (u), l’and d.Subcapitulum ( Figs. 6, 7) posteriorly rounded, anteriorly subconical, 148–230 wide at level of palp insertion, 146–170 long, setae m22 long, anterior to n, 22 long, two pairs of adoral setae or 1–2, 12–14 long.  Idiosoma.In dorsal view subtrapezoidal ( Figs. 3, 4), 860–1110 long, 593–790 wide at level of coxa IV.  Aspidosoma( Fig. 5). Subtrapezoidal, 284–324 long, posterior margin 221–276 wide, anterior margin 70–90 wide, projecting horizontally over gnathosoma. Anterior margin straight, with a central depression. Seta Po( Fig. 5D) 100–120 long, expanded distally, with anterior third setulate, dorsally visible from anterior margin of aspidosoma. Trichobothrial seta bospatulate, 90–104 long, distal half upturned, broadening at tip ( Fig. 5D). Aspidosomal setation regressive, Pa( Fig. 5B) regressive, subcylindrical, 10–12 long, located on a transversal ridge that is united in some exemplars to longitudinal ridge; Pmregressive ( Fig. 5C), cylindrical, 10–11 long, on lateral border, setae Ppabsent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 18–24 diameter, posterior pair 24–32 ( Fig. 5A), median eye 40 long, 60 wide.  Hysterosoma. Dorsal view.With 5 dorsal plates D, (L), Mand P( Figs. 3, 4). Dorsal plate Dsubtrapezoidal, 279–398 long, anterior margin 184–232 long, posterior margin 280–320 long, with one longitudinal and two transversal and ridges, setae a1, b1, c1,ellipsoidal, lengths a1: 24–39, b1: 28–36, c1: 36–50; distance between setal insertions a1-a1: 106–130, b1-b1: 64–80, c1-c1: 80–98, a1-b1: 116–142, b1-c1: 108–152. Paired plates L344–440 long, 74–100 wide; a2regressive, 12–28 long; b2present with p=0.36 (n=14), 32–40 long, c2when present (p=0.43, n=14) ellipsoidal, 24–40 long; lyrifissure ialocated on external side of plate, halfway between a2and b2, 24–34 long; lyrifissure imlocated on median line of plate, halfway between b2and c2, 12–18 long. Median plates Mfused, with d1, d246–60 and 30–52 long, respectively; sometimes with neotrichy (p=0.29, n=14) on d2sector; distance between setal insertions d1-d1: 122–178, d1-d2: 126–154, d2-d2: 308–434. Posterior plate Pdivided by ridges, setae e1,  e2, e2’and  e2”,50–58, 34–44, 32–44 and 30–40 long, respectively. Setae e2’and e2”inserted on different sector of plate, which is separated by a ridge from e2; distance between setal insertions e1-e1: 110–138, e1-e2: 84–94. Seta hson posterior border of idiosoma, 40–46 long.  Ventral view, podosoma and opisthosoma( Fig. 6, 7). Membranous integument striate. Coxal setation formula 4:3:4:2–3, coxal setae clavate; their lengths, 1a: 20–28, 1b: 26–34, 1c: 32–38, 1d: 42–46; 2a: 23–26, 2b: 24–30, 2c: 26–30, 3a: 20–28, 3b: 24–28, 3c: 24, 3d: 22–24, 4a: 25–30, 4b: 24–26, 4c: 20–28, 4d: 20–30. Genital opening 140–174 long, 60–66 wide, with 6 pairs of genital setae, short, 16–22 long. Aggenital plate not observable. Anal opening 142–144 long, adanal plates 38–40 wide, each with two adanal clavate setae, 16–20 long. Pseudanal plate 80–82 long, 54–60 wide, with three pairs of setae, ps1clavate 30–49 long, normally developed, ps2and ps3regressive, ps216–22 long; ps316–20 long, membranous integument with 8 pairs of setae.   FIGURE 3.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, dorsal habitus, SEM. Scale bar, 200 µm.   FIGURE 4.  Andocaeculus weyrauchi(Franz), female, MACN-Ar 41058, dorsal idiosoma. Scale bar, 200 µm.   FIGURE 5.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, SEM, A, aspidosoma; B, Paseta; C, Pmseta; D, anterior view. Scale bars, A, 200 µm; B–C 5 µm; D 100 µm.   FIGURE 6.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, ventral idiosoma, SEM. Scale bar, 200 µm.   FIGURE 7.  Andocaeculus weyrauchi(Franz), female, MACN-Ar 41058, ventral idiosoma. Scale bar 200 µm.   FIGURE 8.  Andocaeculus weyrauchi(Franz), MACN-Ar 41060, A, right leg I; B, right leg II. Scale bars, A–B, 200 µm   FIGURE 9.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, Leg I SEM, A, right trochanter; B, right basi- and telofemur; C, left genu, dorsal view; D, right tibia, antiaxial view; E, solenidion φand famulus Κ”on right tibia; F, left tibia, dorsal view; G, left tarsus, dorsal view; H, right tarsus, antiaxial view; I, distal portion of right tarsus; J, left tarsus, dorsal view, K, full leg, antiaxial view. Scale bars, A–C, 100 µm; D, 50 µm; E, 5 µm; F–G, 100 µm; H–J, 50 µm; K, 200 µm.   FIGURE 10.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, Leg II SEM, A, left trochanter, dorsal view; B, right basi- and telofemur, dorsal view; C, left genu, dorsal view; D, right tibia, dorsal view; E, detail of the solenidion φ; F, right tibia, antiaxial view; G, left tarsus; H, right tarsus, distal portion; I, right tarsus, dorsal view. Scale bars, A–C, 100 µm; D, 50 µm; E, 20 µm; F–I, 50 µm.   FIGURE 11.  Andocaeculus weyrauchi(Franz), female, A, MACN-Ar 41061, left leg III; B, MACN-Ar 41062, left leg IV. Scale bars, A–B, 200 µm.   FIGURE 12.  Andocaeculus weyrauchi(Franz), Leg III MEB, MACN-Ar 41057, A,left trochanter, dorsal view; B, left basi and telofemur, dorsal view; C, left genu, dorsal view; D, right tibia, antiaxial view; E, right tarsus dorsoparaxial view; F, left tibia, dorsal view. Scale bars, A, 100 µm; B, 50 µm; C-F, 100 µm.   FIGURE 13.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, leg IV, A, right trochanter, dorsal view; B, right tibia, antiaxial view, C, same, detail of the solenidion; D, left tibia, dorsal view; E, right basi and telofemur, dorsal view; F, right tarsus, dorsal view; G, left genu, dorsal view. Scale bars, A, 50 µm; B, 100 µm; C, 20 µm; D, 100 µm; E–G, 50 µm.   FIGURE 14.  Andocaeculus weyrauchi(Franz), female, A, MACN-Ar 41063, right tarsus I, dorsal view; B-C, MACN-Ar 41064; B, right palpus, paraxial view; C, left tarsus I, antiaxial view. Scale bars, A, 100 µm; B–C, 50 µm.   FIGURE 15.  Andocaeculus weyrauchi(Franz), MACN-Ar 41057, A, left palpus, antiaxial view; B, anterior view. Scale bars A–B, 50 µm.  Legs( Figs. 8–14). Measurements see Table 2.   TABLE 2.  Andocaeculus weyrauchi, female, leg dimensions (in microns).    Trochanter Basifemur Telofemur  length width length width length width   Leg I 142–170 120–150 128–160 90–122 62–80 100–120  Leg II 134–160 100–120 102–138 76–104 50–60 86–102  Leg III 118–140 90–124 70–86 72–90 56–66 70–92  Leg IV 136–164 80–110 70–100 72–100 40–60 82–106   Continued.  Genu Tibia Tarsus  length width length width length width   Leg I 150–190 86–108 144–200 60–82 120–150 34–46  Leg II 126–152 76–94 120–168 52–70 104–140 35–38  Leg III 114–152 60–92 142–230 40–60 132–170 28–40  Leg IV 120–150 74–120 158–218 46–64 134–170 30–50  Leg I.Trochanter ( Fig. 9A) with one seta in series l’and one regressive seta in d( Figs. 8A, 9A). Basifemur ( Figs. 8A, 9B) with setae on v’not pointed, distally blunt, but longer than other species of group, 54–64 and 80–96 +long, RBaf: 0.52–0.6, RBaf +: 0.75–0.9, 2 setae in series v”, both regressive ( Fig. 9K), one pedicelated seta in series l’and one regressive in series l”and d. Telofemur ( Figs. 8A, 9B) with seta v’subequal to v’on basifemur, 54–66, 80–96 +long, RTef: 0.5–0.57, RTef +: 0.75–0.84, setae l’; l”and v”regressive. Genu ( Figs. 8A, 9C) with 3 setae in series d; 2 setae in series  l’,distal l’elongated, 54–60, 70–78 +long; 3 pedicelated setae in series l”; series v’with 3 pointed setae, with v1’medially displaced;  v’,62–74, 82–100 +long,  v2’,46–62, 64–80 +long; series v”with 2 setae ( Fig. 9K). Tibia ( Figs. 8A, 9D–E, K) with 2 regressive setae in series d,4 setae on both lateral series; series v’with 4 pointed setae, 62–94, 74–106 +long, v2’medially displaced; series v”with 4 pointed setae ( Fig. 9K), v2”medially displaced, v”52–70 long. Solenidion φand famulus κ” inserted dorsoantiaxially on distal portion of segment ( Fig. 9D–E). Tarsus ( Figs. 9G–J, 14A and C) with 3–4 setae in series  l’,4 setae in series l”, with l1”eupathidial, 3–4 setae in series v’and 4 setae in series v”; trichobothria tb24–34 long. Solenidion ω inserted dorsally ( Figs. 9G, 14A) at level of l1”, simple in structure ( Figs. 14A and C). Famulus inserted at antiaxial facies ( Fig. 14C) at level of l1”. Four eupathidia: l1”, er’and (st). Two claws ol’and ol”lengths 8–10 and 30–42, respectively. Setal count (solenidia): 2, 6, 5, 13, 18(1), 20–22(1).  Leg II( Figs. 8Band 10). Trochanter ( Fig. 10A) with one seta in each lateral series and one regressive seta in series d. Basifemur ( Fig. 10B) d, l”and v1”regressive, v’and v”pedunculated. Telofemur ( Figs. 8B, 10B) with d, l’, l”and v”setae regressive and v’pedunculated. Genu ( Figs. 8A, 10C) with 3–4 regressive setae in dseries, 3 pedunculated setae in l’and 4 setae in l”, v1’displaced externally ( Fig. 8B), v”long and pointed ( Fig. 8B), 72–84 long. Tibia ( Fig.10D) with two regressive seta in d,4 pedunculated setae in l’and 3–4 on l”, 3 pointed setae in v’and v”, solenidion inserted as on tibia I ( Figs. 10E, F). Tarsus ( Figs. 8Band 10G–I) with 4 setae in series l’and l”, l1”always eupathidial, 4 setae in series v’and 3 in v”, solenidion ω inserted dorsally at level of l’,famulus antiaxial, (st)pair absent, trichobothria bt24–34 long. Two eupathidia: l1”and er’. Claws ol’and ol”lengths 7–10 and 28–36, respectively. Setal count: 3, 5, 5, 13–14, 15(1), 18(1).  Leg III( Figs. 11Aand 12). Setae on series v’and v”spatulate, eupathidia absent. Trochanter ( Fig. 12A) with anterior border rounded, with pedunculated setae: v’and l”and 1 regressive in d. Basifemur ( Fig. 12B) with l”and v’elongated and dand v”regressive. Telofemur ( Fig. 12B) with l’, l”, v”and dregressive and v’elongated and pedunculated. Genu ( Figs.11A, 12C) with 2–3, 1–2 regressive setae in series dand l’,respectively; 3 pedunculated setae in series v’and 3 regressive in series  v”.Tibia ( Figs. 12D, F) with 3–4 regressive setae in series d, 3–4 setae in series l”and 4–5 inboth ventral series, solenidion φ inserted on antiaxial face of segment. Tarsus ( Fig. 12E) with 3–4, 2, 4–5, 3–4 setae in series l’, l”, v’and v”, respectively, er”present and er’absent; trichobothria bt108–120 long, claws ol’and ol”of length 12–16 and 26–36, respectively. Setal count: 3, 4, 5, 12–14, 15–18(1), 14–17.  Leg IV( Figs. 11Band 13) Setae as on leg III. Trochanter ( Fig. 13A) with 3 setae: l”pedunculated and v’and dregressive. Basifemur ( Fig. 13E) with v’regressive and l’and v”pedunculated. Telofemur ( Fig. 11B, 13E) with l’, v’and dregressive, with setae of series v’and l”elongated and pedunculated. Genu ( Fig. 13G) with 2 and 1 regressive setae in series dand l’,respectively; 2 elongated and pedunculated setae each in series v’and l”. Tibia ( Figs. 13B–D) with 3 regressive setae in series d, 4 setae in vseries and 3 inseries l”, solenidion φ present ( Fig. 13 B–C). Tarsus ( Fig. 13F) with 2–4, 1–2, 3, 3 setae in series l’, l”, v’and v”, respectively, er”present and er’absent, claws ol’and ol”lengths 12–16 and 42–52, respectively. Trichobothria 108–150 long. Setal count: 3, 3, 5, 9, 14(1), 11–14.  Male(n=1). Externally similar to female except in size.  Gnathosoma. Chelicera 132–136 long. Palp 4-segmented, with chaetotaxy as in female. Subcapitulum 192 wide at level of palp insertion, 146 long, setae manterior to n, two pairs of adoral setae.  Idiosoma.860 long, 592 wide at level of coxa IV.  Aspidosoma.Subtrapezoidal, 269 long, posterior margin 221 wide, anterior margin 70 wide, projecting horizontally over gnathosoma. Seta Po100 long, trichobothrial seta bo90 long. Aspidosomal setation regressive, Paand Pmminute, Ppabsent; anterior pair of lateral eyes 20–22 diameter, posterior pair 26–32.  Hysterosoma. Dorsal view. Dorsal plate Dsubtrapezoidal, 299 long, anterior margin 197 long, posterior margin 254 long, a1: 22–26, b1: 26, c1: 32; distance between setal insertions, a1-a1: 92, b1-b1: 50, c1-c1: 58, a1- b1:108–110, b1-c1: 114–118. Paired plates L336–344 long, 70 wide; a2regressive, 16 long; b2and c2absent, iaand im,30–32 and 16 long, respectively. Median plates Mfused, with d1, d2, 46–48 and 30 long, respectively, distance between setal insertions, d1-d1: 96, d2-d2: 292, d1-d2: 100–102. Posterior plates P, divided by ridges, setae e1, e2, e2’,44–50, 34–36 and 30 long, respectively. Setae e2’separated by ridge from e2, distance between setal insertions, e1-e1: 90, e1-e2: 68–76. Seta hson posterior border of idiosoma, 50 long.  Ventral view, podosoma and opisthosoma. Coxal setation formula 4:3:4:3. Genital opening 162 long, with 6 pairs of genital setae, 24–30 long. Aggenital plate subtriangular, poorly sclerotized. Anal opening 154 long, with two pairs of clavate adanal setae, 16–18 long. Pseudanal plate 168–180 long, 60 wide, with three pairs of setae. Genital sclerites as other species of group ( Fig. 39), with nine pairs of simple, unbranched setae, 14–16 long.  Legs.Measurement see Table 3. Ratios: RBaf: 0.54; RBaf  +: 0.78; RTef: 0.54; RTef +: 0.76.   TABLE 3.  A. weyrauchi, male, leg dimensions.    Trochanter Basifemur Telofemur  length width length width length width  Leg I 130 110 112 92 52 92  Leg II 124–130 98–100 100–102 74–82 46–48 78–82  Leg III 104–106 90–92 70 58–66 54–56 68  Leg IV 128 82 60 64 44 80   Continued.  Genu Tibia Tarsus  length width length width length width  Leg I 142 80 126–134 54 116–118 30–34  Leg II 104–108 70–72 110–120 54–58 98–110 36  Leg III 114–118 60 150–154 50–56 120 34  Leg IV 114 68 150 46 132 36  Larva (n=1)  Gnathosoma.Covered by aspidosoma in dorsal view. Chelicera 42–44 long; cheliceral seta chpresent. Palp 4-segmented, trochanter glabrous, femorogenu with 1 clavate seta in dorsal series, 9–10 long; tibia with a spiniform dseta, 11 long, located on a projecting tubercle, 2 clavate setae l’and  l”,and two spiniform setae ld; 8–11 long, tarsus subconical, solenidion ω present on antiaxial face, 5 setae present:  v, l’,and din addition to 2 eupathidia, (u).Subcapitulum 90 wide at level of palp insertion, 60 long, setae m12–13 long, anterior to n, 10 long, two pairs of adoral setae or 1-2.  Idiosoma355 long, 269 wide at level of coxa III.  Aspidosoma.Subtrapezoidal, 136 long, posterior margin 132 wide, anterior margin 40 wide, projecting horizontally over gnathosoma. Seta Po38 long, trichobothrial seta bo42 long. Aspidosomal setation regressive, Paand Pmminute, Ppabsent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 8 diameter, posterior pair 8–10.  Hysterosoma. Dorsal view( Fig. 16A). Dorsal plates poorly defined; only setae a1, b1, c1, a2, d1, e1and hspresent. Lengths of anterior hysterosomal setae, a1: 8–10, b1: 9–10, c1: 9; a2regressive, 10 long, distance between setal insertions, a1-a1: 58, b1-b1: 38, c1-c1: 52, a1-b1: 45, b1-c1: 43–44. Lengths of posterior hysterosomal setae, d1: 22, e1: 28, distance between setal insertions d1-d1: 48, e1-e1: 38.  Ventral view, podosoma and opisthosoma( Fig. 16B). Coxal setation formula 2:0:1, setae 16–18 long, Claparede’s organs present on anterior margin of coxae II. Anal opening 70 long, hs16 long, ventral lyrifissure ihobservable at level of anal plate.  Legs( Fig. 17). Measurements see table 7. Femora entire on all legs, Rf: 0.65–0.73  Leg I. Tr, 0; Fe,1d*, 1l’, 1l”, 1v’, 2v”; Ge, 2d*, 1l’, 2l”, 1v’, 1v”; Ti, 1d*, 2l’, 2l”, 2v’, 2v”, κ”, φ; Ta, 3l’, 2l”, 3v’, 3v”, tb, ε (dorsal), ω, st’ζ.  Leg II. Tr, 0; Fe,1d*, 1l’, 1l”, 1v’, 2v”; Ge, 2d*, 1l’, 1l”, 1v’, 1v”; Ti, 1d*, 2l’, 2l”, 2v’, 2v”, φ; Ta, 3l’, 2l”, 3v’, 3v”, tb, ε (dorsal/antiaxial), ω.  Leg III. Tr, 0; Fe,1d*, 1l’, 1l”, 1v’, 2v”; Ge, 1d*, 1l’, 1l”, 1v’, 1v”; Ti, 1d*, 1l’, 2l”, 2v’, 2v”, φ; Ta, 2l’, 1l”, 3v’, 2v”, tb.   FIGURE 16.  Andocaeculus weyrauchi(Franz), larva, MACN-Ar 41065 A, idiosoma dorsal; B, idiosoma ventral. Scale bar 100 µm.  Protonymph (n=4)  Gnathosoma.Covered by aspidosoma in dorsal view. Chelicera 64 long; cheliceral seta chslender present. Palp 4-segmented, trochanter glabrous, femorogenu with 1 clavate seta in dorsal series, 20–22 long, tibia with prominent dcalcar on projecting tubercle, 14 long, 2 clavate setae l’and  l”,and one calcar ld, 12–14 long; tarsus subconical, 18–20 long, 12–14 wide at base, solenidion ω present on antiaxial face, 5 setae: v, l’, din addition to 2 eupathidia (u). Subcapitulum 130 wide at level of palp insertion, 86 long, setae manterior to n, two pairs of adoral setae or 1–2.  Idiosoma.434–528 long, 260–368 wide at level of coxa IV.  Aspidosoma.160–176 long, posterior margin 140 wide, anterior margin 42–50 wide, projecting horizontally over gnathosoma. Anterior margin circular. Seta Po54–60 long, trichobothrial seta bo56–80 long. Aspidosomal setation regressive, Paand Pmminute, Ppabsent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 10–12 diameter, posterior pair 12–18.   FIGURE 17.  Andocaeculus weyrauchi(Franz), larva, MACN-Ar 41065, A, right leg I; B, right leg II; C, right leg III. Scale bars, A–C, 50 µm.  Hysterosoma. Dorsal view. Unpaired dorsal plate Dsubtrapezoidal, 150–166 long, anterior margin 110–140 long, posterior margin 150–160 long, setae a1, b1, c1, present, lengths, a1: 10–14, b1: 14–15, c1: 20; distance between setal insertions, a1-a1: 56–70, b1-b1: 47–50, c1-c1: 47–52, a1-b1: 56–66, b1-c1: 59–70. Paired plates L180–200 long, 38–50 wide; a2regressive, 8–10 long; b2and c2absent; iaand im18–20 and 6–10 long, respectively. Median plate Mfused, d130–32 long, d2absent; distance between setal insertions d1-d1: 54–72. Posterior plates Pwith setae  e128–32 long, e2absent, distance between setal insertions e1-e1: 50–64. Seta hson posterior border of idiosoma, 20 long.  Ventral view, podosoma and opisthosoma. Coxal setation formula 2:1:2:1, coxal setae clavate. Genital opening 22–28 long, with 1 pair of setae, 8–10 long. Anal opening 80–86 long, with two pairs of clavate setae, 20 long. Pseudanal plate 70–80 long, 30 wide, with three pairs of setae, ps1, ps2and  ps3,14–18, 10–12, 8–10 long, respectively.  Legs.Measurement see table 6. Femora entire in leg IV, RTef: 0.68–0.73, RTef +: 0.84–0.95.  Leg I. Tr, 1d (p=0.5), 1l’; PrFe,1d (p=0.5) v”; TeFe,1d*, 1l’, 1l”, 1v’, 1v”; Ge, 2d*, 2l’, 1l”, 1v’, 1v”; Ti, 1d*, 2l’, 2l”, 2v’, 2v”, κ”, φ; Ta, 3l’, 2l”, 3v’, 3v”, tb, ε (antiaxial), ω, (st)ζ.  Leg II. Tr, 1d (p=0.5, n=4), 1l’, 1l”; PrFe, v’, TeFe,1d*, 1l’, 1l”, 1v’, 1v”; Ge, 2d*, 1l’, 1l”, 1v’, 1v”; Ti, 1d*, 2l’, 2l”, 2v’, 2v”, φ; Ta, 3l’, 2l”, 3v’, 3v”, tb, ε (antiaxial), ω.  Leg III. Tr, d*(p=0.5, n=4), v’; PrFe,v’; TeFe,1d*, 1l’*, 1l”*, 1v’, 1v”; Ge, 1d*, 1-2l’*, 1l”, 1v’, 1v”; Ti, 1d*, 1l’, 2l”, 2v’, 2v”, φ; Ta, 2l’, 1l”, 3v’, 2v”, tb.  Leg IV. Tr, 0-1v’; Fe,1d*, 1v’, 0-1v”; Ge, 1d*, 1l’*,1l”*, 1v’, 1v”; Ti, 1d*, 1l’, 2l”, 2v’, 2v”, φ; Ta, 1l’, 1l”, 2v’, 2v”, tb.   TABLE 4.  A. weyrauchi, tritonymph, leg dimensions.    Genu Tibia Tarsus  length width length width length width  Leg I 126–130 62–76 136–142 54–62 106–114 32–40  Leg II 106–110 66 110–114 52–54 104–106 26–36  Leg III 98–110 56–68 150–160 42–50 118–130 28–36  Leg IV 92–102 62–68 140 40–56 120–124 34–40   Continued.  Genu Tibia Tarsus  length width length width length width  Leg I 126–130 62–76 136–142 54–62 106–114 32–40  Leg II 106–110 66 110–114 52–54 104–106 26–36  Leg III 98–110 56–68 150–160 42–50 118–130 28–36  Leg IV 92–102 62–68 140 40–56 120–124 34–40  Deutonymph (n=2)  Gnathosoma.Covered by aspidosoma in dorsal view. Chelicera 44 wide at base, 90 long; cheliceral seta chpresent. Palp 4-segmented, trochanter glabrous, femorogenu with 2 clavate setae in dorsal series, dand d1, 24–30 and 14–20 long, respectively; tibia with 2 calcars dand ld, 17–20 and 18–20 long, respectively, and 2 clavate setae l’and  l”,12–16 long; tarsus subconical, ω solenidion present on antiaxial face, 5 setae v, l’and din addition to 2 eupathidia (u). Subcapitulum 122–164 wide at level of palp insertion, 106–118 long, setae m24–26 long, anterior to n, 20 long, two pairs of adoral setae or 1–2.  Idiosoma.569–711 long, 410–458 wide at the level of coxa IV.  Aspidosoma.192–220 long, posterior margin 164–170 wide, anterior margin 44–50 wide, projecting horizontally over gnathosoma. Seta Po62–72 long, trichobothrial seta bo72–80 long. Aspidosomal setation regressive, Paand Pmminute, Ppabsent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 10–18 diameter, posterior pair 16–21.  Hysterosoma. Dorsal view.Unpaired dorsal plate Dsubtrapezoidal, 180–221 long, anterior margin 150–184 long, posterior margin 180 long, lengths, a1: 18–20, b1: 22–24, c1: 25–26; distance between setal insertions, a1-a1: 81–88, b1-b1: 42–56, c1-c1: 50–66, a1-b1: 72–84, b1-c1: 69–94. Paired plates L298 long, 80 wide; a2regressive 11–12 long; b2and c2absent; iaand im21–30 and 12 long, respectively. Median plate Mfused, d1: 30–32 long, d2absent in the second exemplar; distance between setal insertions d1-d1: 72–84, d1-d2: 72–86, d2-d2: 237. Posterior plates Pwith setae e1and  e230–44 and 16–30 long, respectively; distance between setal insertions e1-e1: 56–70, e1-e2: 44–56. Seta hson posterior border of idiosoma, 26–30 long.  Ventral view, podosoma and opisthosoma.Coxal setation formula 4:1–2:5:2. Genital opening 34 long, with 1 pair of genital setae, 14–16 long. Anal opening 94–110 long, adanal plate 30 wide, with two pairs of adanal clavate setae, 10–14 long.  Legs.Measurement see table 5. Ratios: RBaf: 0.43–0.55; RBaf  +: 0.6–0.7, RTef: 0.48–0.61, RTef +: 0.59–0.77  Leg I. Tr, 1d, 1l’; PrFe, 1d, l’, l”, v’ v”; TeFe,1d*, 1l’, 1l”, 1v’, 1v”; Ge, 2d*, 1l’, 2l”, 2v’, 2v”; Ti, 1d*, 2l’, 1-2l”, 3v’, 3v”, κ”, φ; Ta, 2-3l’, 2l”, 3v’, 3v”, tb, er’ζ, er”, ε, ω, (st)ζ,  Leg II. Tr, 1d*, 1l’, 1l”; PrFe, 0-1d*, l”, 1v’,1v”; TeFe,1d*, 1l’*, 1l”*, 1v’, 1v”; Ge, 2d*, 1l’, 2l”, 2v’, 1v”; Ti, 1d*, 2l’, 2l”, 2v’, 2v”, φ; Ta, 2l’, 2l”, 3v’, 3v”, er’ζ, er” tb, ε, ω.  Leg III. Tr,0-1d*, l”, v’; PrFe,0-1l”, 1v’, 1v”; TeFe,1d*, 1l’*, 1l”*, 1v’, 1v”; Ge, 2d*, 2l”, 2v’, 2v”; Ti, 2d*, 2l”, 3v’, 3v”, φ; Ta, 2l’, 1l”, 3v’, 2-3v”, er”, tb.  Leg IV. Tr, 0-1d*, 0-1v’; PrFe, 1v’; TeFe, 1d*, 0-1l’, 1l”, 1v’, 1v”; Ge, 1-2d*, 1l’*, 1-2l”, 2v’, 1v”; Ti, 1d*, 1l’*, 2l”, 2v’, 2v”, φ; Ta, 1l’, 1l”, 3v’, 2v”, er”, tb.   TABLE 5.  A. weyrauchi, deutonymph, leg dimensions.    Trochanter Basifemur Telofemur  length width Length width length width  Leg I 70–76 76–78 78 68–70 36 66–68  Leg II 66–68 68 62–72 52 30–32 58  Leg III 66–70 68–70 50 42 34–35 54–56  Leg IV 70 58–60 40–62 48–50 30–32 54–56   Continued.  Genu Tibia Tarsus  length width length width length width  Leg I 84–86 54–60 80 46–48 70 26–30  Leg II 70 50–52 70–74 42–44 66–68 28  Leg III 70–72 46 84–88 36–38 76–80 24–26  Leg IV 68–70 46–48 80–98 40 80 26–28   TABLE 6.  A. weyrauchi, protonymph, leg dimensions.    Trochanter Basifemur Telofemur  length width Length width length width  Leg I Leg II Leg III 60–70 60–66 50–64 58–74 48–58 46–60 52–66 42–58 40–64 42–56 34–44 30–48 30–36 26–30 26–32 44–62 40–56 42–52  Femur  Leg IV 46–66 40–56 42–56 32–48   Continued.  Genu Tibia Tarsus  length width length width length width  Leg I Leg II Leg III 72–88 58–70 58–64 42–60 36–52 32–48 66–86 60–70 64–82 30–48 30–44 26–40 62–74 62–70 60–74 20–30 18–28 20–28  Leg IV 46–62 32–46 62–72 28–40 64–70 18–28  Tritonymph (n=6)  Gnathosoma.Covered by aspidosoma in dorsal view. Chelicera 106–110 long. Palp 4-segmented, trochanter glabrous, femorogenu with 2 clavate setae in dorsal series, dand d1, 36–40 and 26–30 long, respectively; tibia with 2 calcars dand ld, 22–26 and 26 long, respectively, and 2 clavate setae l’and  l”,16–26 long; tarsus subconical, 34–40 long, 16–22 width at base, ω present on antiaxial face, 5 setae v, l’and din addition to 2 eupathidia (u). Subcapitulum 192–204 wide at level of palp insertion, 92–130 long, setae m26–28 long, anterior to n, 22–24 long, two pairs of adoral setae or 1-2, 9–11 long.  Idiosoma.700–943 long, 553–711 wide at level of coxa IV.  Aspidosoma.Subtrapezoidal, 253–276 long, posterior margin 215–241 wide, anterior margin 78–82 wide, projecting horizontally over gnathosoma. Seta Po82–90 long, trichobothrial seta bo, 76–90 long. Aspidosomal setation regressive, Paand Pmminute, Ppabsent. Lateral eye plates separated from other plates; anterior pair of lateral eyes 24 diameter, posterior pair 26–30.  Hysterosoma. Dorsal view.Unpaired dorsal plate Dsubtrapezoidal, 221–291 long, anterior margin 176–205 long, posterior margin 246 long, a1: 22–24, b1: 26–30, c1: 26–32; distance between setal insertions, a1-a1: 96–100, b1-b1: 70, c1-c1: 66–76, a1-b1: 108–112, b1-c1: 106–114. Paired plates L344–352 long, 74–80 wide; a2regressive, 12–13 long; b2present with p=1/12, 22 long, c2present with p=1/2, 18–22 long; iaand im26–32 and 12–18 long, respectively. Median plates Mfused, with d1, d238–42 and 30 long, respectively, distance between setal insertions, d1-d1: 114, d2-d2: 316–324, d1-d2: 100–110. Posterior plates Pdivided by ridges, setae e1, e2, e2’, 42, 36–38 and 26–32 long, respectively. Setae e2’separated by ridge from e2, distance between setal insertions, e1-e1: 96, e1-e2: 64–80. Seta hson posterior border of idiosoma, 38 long.  Ventral view, podosoma and opisthosoma. Coxal setation formula 3–4:2–3:3–4:2–3. Genital opening 70–94 long, with 3 pairs of genital setae, 12–20 long. Anal opening 146–164 long, adanal plate 30–36 wide, with two pairs of adanal clavate setae, 12–15 long. Pseudanal plate 200 long, 135 wide, with three pairs of setae.  Legs.Measurements see table 4. Ratios: RBaf: 0.57–0.68; RBaf  +: 0.75–0.85; RTef: 0.55–0.61; RTef +: 0.71– 0.77.  Leg I. Tr, 1d*, 1l’; PrFe, 1d*, l’, l”, v’, 2v”; TeFe,1d*, 1l’, 1l”, 1v’, 1v”; Ge, 2-3d*, 1-2l’, 2l”, 3v’, 2v”; Ti, 2d*, 3l’, 3l”, 3v’, 3v”, κ”, φ; Ta, 3l’, 3l” (1”ζ), 2–3v’, 3–4v”, tb, ε, ω, (st)ζ, er’ζ, er”.  Leg II. Tr, 1d*, 1l’, 1l”; PrFe, 1d*, l’*, 1v’,2v”; TeFe,1d*, 1l’*, 1l”*, 1v’, 1v”; Ge, 3d*, 3-4l’, 2l”, 1-2v’, 1v”; Ti, 1-2d*, 3l’, 3l”, 3v’, 2-3v”, φ; Ta, 3l’, 3l” (l’ ζ,), 3v’, 3v”, tb, ε, ω,, er’ζ, er”.  Leg III. Tr, 1d*, 1l”, 1v’; PrFe,1d*, 1l”, 1v’, 1–2v”; TeFe,1d*, 1l’*, 1l”*, 1v’, 1v”; Ge, 2d*,0–2l’*, 2l”, 2v’, 2v”; Ti, 3d*, 3l”, 4v’, 4v”, φ; Ta, 3l’, 2l”, 3v’, 3v”, er”, tb.  Leg IV. Tr, 1d*, l”, 1v’; PrFe, 0-1d, 0-1l”, 1v’, 1v”; TeFe, 1d*, 1l’*, 1l”, 1v’, 1v”; Ge, 2d*, 0-1l’*, 2l”, 2v’, 2v”; Ti, 2–3d*, 2-3l”, 3v’, 3v”, φ; Ta, 2l’, 1l”, 3v’, 3v”, er”, tb. 3071274319 1963-09-13 MNHN Herbert Franz Erde 3000 Seehohe am Fusse Rasen 6 7 1 holotype 3071274330 GPS Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 4 4 Tucuman 3071274323 [151,973,716,743] GPS Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 1 1 Tucuman 3071274303 GPS Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 2 2 Tucuman 3071274301 [399,1203,752,779] GPS Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 10 10 Tucuman 3071274329 GPS Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 1 1 Tucuman 3071274312 GPS Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 14 14 Tucuman 3071274322 SPDAR Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 SPDAR1609-16, 1609-20 1 1 Tucuman 3071274317 SPDAR Berlese & A. Porta Argentina 50 -26.74272 Infiernillo 1 -65.77283 Abra 6 7 2 2 Tucuman