A Revision of the Yoyetta abdominalis (Distant) Species Group of Cicadas (Hemiptera: Cicadidae: Cicadettinae), Introducing Eight New Species
Emery, David L.
Emery, Nathan J.
Popple, Lindsay W.
Records of the Australian Museum
2019
2019-10-02
71
7
277
347
8K2CN
Emery & Emery & Popple, 2019
Emery & Emery & Popple
2019
[914,1179,1625,1657]
Insecta
Cicadidae
Yoyetta
Animalia
Hemiptera
33
308
Arthropoda
species
spectabilis
sp. nov.
Figs 2D, 3D, 22–25, 52D, 53K
Holotype ♂, Wises Track, Royal National Park, NSW, 34°06'51"S 151°03'30"E, 9.i.2005, S. N. & D. Emery( AMK.536052). Paratypes NEW SOUTH WALES: 1♂ 1♀, AustraliaNSW, Morton NP, Malaiseon edge between open field & woodland Leptospermumunderstorey, off Touga Rd, 1–8.xii.2009, C. Manchester, D.Yeates, B. Lessard,ANIC Bulk Sample No.3381, 35°02'35"S 150°08'31"EGPS; 2♂♂, AustraliaNSW, Clyde Mountain(near top of highway pass), 28.xii.1994; A. Sundholm& R. De Keyzer; 1♂, Blue Mtns1899, W. W. Froggattcollection ( ANIC); 1♀, Park Rd, Waitara, NSW, 33°42'31"S 151°06'27"E, 17.xii.1996, M. S. Moulds; 1♀, Knapsack St, Glenbrook, NSW, 33°45'46"S 150°38'17"E, 10.xi.1997, D. Emery; 1♂ 1♀, Barren Ground Nature Reserve, NSW, 34°42'43"S 150°42'21"E, 15.i.1999, S., N. & D. Emery; 3♂♂ 2♀♀same location, 15.i.2000, S. & D. Emery; 6♂♂same location, 1.i.2004, S. & D. Emery; 2♂♂, same location, 29.xii, 2005, N. & D. Emery; 2♂♂, Clarence, NSW, 6.xii.2000, R. Chin; 2♂♂ 1♀, Blackheath, NSW, 33°36'44"S 150°19'28"E, 23.xi.2002, D. Emery& L. Popple; 1♂, same location, 14.xii.2002, D. Emery; 2♂♂, same location, 22.xii.2002, N. & D. Emery; 2♂♂, 20 kmSW Nowra, NSW, 35°01'35"S 150°28'37"E, 1.i.15, S., N., C. & D. Emery; 1♂, Berrara, 17.x.2015, N. Emery; 2♂♂, Medway, NSW, 34°29'25"S 150°16'27"E, 6.i.17, S., N. & D. Emery& T. Corbin; 3♂♂, Dolphin Point, NSW, 36°25'41"S 150°26'40"E, 19–28.i.2018, J. Poyitt( DE); 2♂♂, same location, 19–28.i.2018, J. Poyitt; 1♂, same location, 2.i.2014, J. Poyitt( JP); 1♂, Bendalong, NSW, 26.xii.1968, G. Daniels; 1♀, Royal National Park, NSW, 7.ii.1986, C. N. Smithers; 1♂, “ Tuglo”, 60 kmN Singleton, NSW, 13.xi.1983, B. Day& K. Kloo; 2♂♂, Macquarie Pass, NSW, 28.xii.1992, T. M. S. Hanlon; 6♂♂, Barren Ground Picnic Area, NSW, 10.i.2006, D.Britton( AM); 2♂♂, Blackheath, 23.xi.2002, L. Popple, D.Emery, 513-0001, 513-0002; 1♀, Barren Grounds, 31.xi.2003, L. Popple, D. Emery; 3♂♂, Burradoovia Jambaroo, 31.xi.2003, L.Popple, D. Emery, 513-0004 to 513-0006, 1♂, Genitaliaprep. 513-01; 2♂♂, Kanangra Walls, 33°59'19"S 150°05'32"E, S12588, Heathland, 9.i.2010, Popple& Emery, 513-0008, 513-0009; 1♂, Kanangra-Boyd National Park, 33°53'10"S 150°45'23"E, S12588, 9.i.2010, Popple& Emery, 513-0007; 1♂, Blackheath, i.2010, D. Emery, 513 ( LWP); 1♂, Ulladulla, 23.xii.1994, M. S. & B. J. Moulds; 5♂♂, Hat Hill, Blue Mountain Nat. Pk, 5.xii.1971, G. Daniels; 1♂ 1♀, same data, 30.xi.1975; 3♂♂, Kanangra Walls, 23.xii.1973, G. Daniels; 1♂, Wentworth Falls, Blue Mountains, xii.1990, Keith Chambers; 3♂♂ 1♀, Hat Hill, Blackheath, 24.xi.1973, G. Daniels; 1♂, Cudmirrah Faunal Reserve, 24.xii.1974, G. Daniels; 1♂, Mt Leighton Bailey, Royal Nat. Pk, 2.ii.1978, G. Daniels; 1♂, 8 kmE Robertson, 12.xii.1976, G. Daniels; 1♂, George Boyd Lookout, nr Ulladulla, 12.xii.1990, A. Sundholm; 1♂, Bobbin Head, Ku-Ring-Gai Chase National Park, 16.ii.1970, J. Barratt; 1♂, Budderoo Fire Trail, approx. 18 kmSE Robertson, 18.xi.1984, G. Williams, ex. woodland; 1♂, Currarong, 24.i.1993, T. M. S. Hanlon; 7♂♂, Hat Hill, Blackheath, Blue Mts, 11.xii.1971, J. V. Peters; 1♂, Mt Wilson, Blue Mountains, 30.xi.1975, G. Daniels; 1♂ 1♀, Jenolan Caves, 30.xii.1991, R. Eastwood; 2♂♂, Clyde Mt, 28.xii.1994, R. de Keyzer& A. Sundholm; 1♀, Tinderry Rg, 10.xii.1961, L. A. Greenup; 1♂, Jct Rawson Pdeand Mt Hay Rd, N. of Leura, 33°42.215'S 150°20.308'E, 980 m[elev.], 14.i.2011, Figure 22. Yoyetta spectabilis sp. nov., (A)male holotype, Kuringai Chase National Park (33°40'S 151°12'E), dorsal habitus; (B)male, ventral habitus; (C)female, Royal National Park (34°06'S 151°03'E), dorsal habitus; (D), female, Royal National Park, ventral habitus; (E), female, Barren Grounds (34°41'S 150°44'E), brown variant, dorsal habitus; (F), live male specimen, Blackheath (33°35'S 150°50'E), photo by NJE. Scale bar (for preserved specimens) = 10 mm. Figure 23. Yoyetta spectabilis sp. nov., illustration of male pygofer and internal genitalia; (a) viewed laterally from the left; (b)viewed ventrally; (c)aedeagus; and (d)apex of theca. Characters as depicted in Fig. 4. Specimen from Barren Grounds ( 34°41'S 150°44'E). K. Hill, D. Marshall,AU.NS.MHA, C. Simon lab voucher, Yoyetta“zizzer”, legs in ETOH, 11.AU.NS.MHA.03; ( MSM ); 1♀, Tubrabucca, Barrington Tops, NSW, 31°52'S 151°25'E, 17.i.1948( HEM2218); 1♂, Barrington, NSW, 31°59'S 151°55'E, 17.xii.1921( HEM2219); ( MMV). AUSTRALIAN CAPITAL TERRITORY: 2♂♂, Jervis Bay, ACT, 33°45'24"S 150°44'55"E, 12.xii.10, N. & D. Emery( DE). VICTORIA: 1♂, Shady Ck V., 7 kmN. Yarragon, 7.i.1983, D. F. Crosby; 1♂ 1♀, N. Walkerville, 28.xii.1988, H. Abbott( MSM); 2♂♂, Tyers River, Moondarra Vic, 28.xii.1976, W. N. B. Quick( ANIC); 1♂, Yinnar, Vic, 38°19'S 146°19'E, 5.i.1952( HEM2237); 1♂, same location, 16.xii.1948( HEM2238) ( MMV ); 1♂, Narre Warren, 21.i.62; 2♀♀, UpperB’field, 13.i.1998( QM); 1♂, Berwick, Victoria, G. J. Hall, 13.i.1924, Melampsalta abdominalisDist., det. G. J. Hallafter Nat. Mus. Vict., SAMA Databaseno. 20-014466 ( SAM). Other material. NEW SOUTH WALES: 4♂♂, Hat Hill Rd., Blackheath, NSW, 33°36'59"S 150°18'29"E, 19.xii.2007, C. & D. Emery; 1♀, same location, 10.i.2006, D. Emery; 1♂, Wises Track, Royal National Park, NSW, 34°06'51"S 151°03'30"E, 9–13.i.2005, S. N. & D.Emery; 4♂♂, same location, 22.i.2007, S., N. &, D. Emery; 5♂♂ 1♀, Budaroo National Park, NSW, 34°36'44"S 150°42'05"E, 30.xi.2003, S. & D. Emery& L. Popple; 11♂♂ 7♀♀, same location, 27.i.2007, S. & D. Emery; 8♂♂, same location, 1.i.2004, S. N. & D. Emery; 2♂♂ 1♀, Anvil Rock, NSW, 33°35'44"S 150°20'00"E, 6.xii.2009, D. Emery; 3♂♂ 1♀, same location, 9.xii.2009, D. Emery; 1♂, Kuringai Chase National Park, NSW, 33°40'34"S 151°12'52"E, 14.xii.2009, D. Emery; 2♂♂, St. Ives Showground, NSW, 33°42'22"S 151°11'21"E, 5–6.i.2010, D. Emery; 1♂, Kanangra Walls,NSW, 33°59'18"S 150°02'25"E, 8.i.2010, D. Emery& L. Popple; 6♂♂ 1♀, Kangaroo Track, Barren Ground Reserve, NSW, 34°41'40"S 150°44'19"E, 28.xii.2010, T., S., N., C. & D. Emery; 3♂♂, Lockley’s Pylon Track, Leura, NSW, 33°36'05"S 150°20'00"E, 20.xii.2012, N. & D. Emery; 5♂♂, same location, 7.xii.13, A. Greenville& N. Emery; 1♂ 1♀, same location, 2.xii.14, T., S., N., C. & D. Emery; 1♂, same location, 2.xii.14, A. Schreiber; 1♂, same location, 12.xii.14, N. Emery; 1♀, Bundanoon, NSW, 19.i.2012, G. Bellis( DE). VICTORIA: 1♂, [labels in handwriting of J. H. Ashton] M. abdominalis Dist. Vic., K30137 [typed], genitalia preparation [handwritten] Y029, AM K.307108 (donated in 1911).
Distribution, habitat and seasonality. Highlandlocations from Barrington Tops south to the Southern Highlands of New South Wales. Specimens have been found in coastal locations from northern Sydney, south to eastern Victoria, close the Melbourne ( Fig. 18). Adults occur mainly on small heath shrubs and sometimes on eucalypts. Males constantly Figure 24. Male calling song structure of Yoyetta spectabilis sp. nov, illustrated in waveform plots, including (A)a series of echemes, including an introductory echeme and three typical echemes, illustrating the “round whirling” and “abrupt whirling” modulations (explained in the text); (B)expanded section of the round whirling from the introductory echeme (in A) showing rounded amplitude modulations; (C)further expanded detail of a round whirling amplitude modulation; (D)expanded detail of a typical echeme (in A) showing the abrupt whirling modulations; (E)expanded detail of an abrupt whirling amplitude modulation (from D); (F)expanded (from D) to show an abrupt modulation followed by the terminal modulation (explained in text). The final subfigure (G)is a spectrogram displaying song frequency. The specimen was recorded in the field at Wises Track, Audley (34°08'S 151°03'E) by LWP using RS1 (see Methods and terminology). move between singing stations that are usually < 3 mhigh. Females are located close to ground level. Since their song is somewhat similar to that of Y. regalis sp. nov., the location of Y. spectabilis sp. nov.on the heathland shrubs contrasts with the distinct preference of the former species for smooth-barked eucalypts when the two species cohabit open woodland. This species can be encountered from late November to January.
Description Male( Figs 2D, 3D, 22A,B,F, 52D). Headalmost as wide as mesonotum, black in mountain specimens, with a small, triangular brown fascia posterior to ocelli, reducing anteriorly, and a small to large ochraceous spot on posterior margin between ocelli and eye; covered with black pubescence. In coastal specimens, including the holotype, pale brown to ochraceous posteriorly, extending halfway around eye, black surrounding ocelli, black spot medial to eye; ocelli pink; postclypeus dorsally dark brown to black with yellowish triangular median fascia, apex directed anteriorly, ventral surface black with black transverse grooves, midline with ochraceous line over anterior half in some specimens, lateral and posterior borders ochraceous, tending reddish; anteclypeus black, rostrum black at base, dark brown centrally, black at apex, reaching anterior edge of hind coxae; lora ochraceous anteriorly, variably black on medial and posterior sections; eyes variably dull black to dark brown; antennae black, supraantennal plates black, coastal specimens with central brown spot on anterior margin extending laterally above pedicel. Thoraxpredominantly dull black on mountain specimens, brown on coastal cicadas. Pronotum black with distinct ochraceous marking over central third of midline, variably bordered by black, fissures variably black, tending brown on raised areas; pronotal collar brown centrally, variably tending black laterally, lateral angles black, paranota black on posterior and lateral margins, becoming brown elsewhere. Mesonotum black on mountain cicadas, coastal specimens brown with sigilla, scutal depressions and surrounds black; cruciform elevation arms black, lateral depressions brown. Metanotum black. Legs.Coxae mainly black and ochraceous, fore coxae with anterior black markings ranging from proximal spot to a longitudinal stripe, mid and hind coxae black, variably ochraceous laterally, membranes red; meracantha small, narrow, reddish, black at base, pointed, overlapping onequarter of opercula; trochanters red with black medial stripe; femora red laterally, black medially; femoral spines erect, reddish to black at base tending dark brown at tips; fore tibia black, tending brown distally, browner on coastal specimens; mid tibiae ochraceous with proximal black stripe, hind tibiae pale brown with variable, small black marking on medial aspect of proximal segment; tarsi on coastal specimens reddish tending black at distal joints, fore tarsi brown on mountain specimens becoming black towards claws, mid tarsi light red with black at claws, hind tarsi red with brown at claws; claws dark brown, black at tips. Wingswith fore wing costal veins orange-red, proximal margin black posteriorly; pterostigma mottled red; basal cell hyaline with black anterior border; basal membranes bright orange-red, other veins dark brown to black, with eight apical cells; hind wing plagas white over entire anal cell 3 and thin along vein 2A, some reddish stippling at base, with six apical cells. Opercula( Fig. 3D) small, spatulate, following body axis ventrolaterally, depressed centrally, variably black at base, orange-red across remainder, clearly separated. Timbals( Fig. 2D) with five distinct long ribs; long ribs 1–3 extending across surrounding membrane and fused dorsally along basal spur; long rib 4 discontinuous at proximal third; long rib 5 independent of basal spur, comparatively shorter, extending ventrally across half of membrane; large ridged dome on posterior timbal plate extending across two-thirds of timbal; apodeme pit oval-shaped and conspicuous. Abdomen. Tergite 1 black with red lateral membranes; tergites 2–3 black with diffuse dark brown shading on some coastal specimens, intersegmental membranes orange; tergites 4–7 black with thin orange posterior margins, an increasing area of lateral orange coloration posteriorly forming a triangular shape with base on tergite 7, extending to lateral border and to epipleurite 7; tergite 8 shiny black, with orange-red posteriorly and extending laterally in some specimens. Epipleurites with variable black medial markings, fiery red laterally and posteriorly. Sternite II mainly black; sternite III black centrally, orange-red laterally; sternites IV–VI orange, posterior halves becoming red, black midline markings over anterior two-thirds of each sternite, decreasing posteriorly, sternite VIII fiery red, with yellowish pubescence ( Fig. 51D). Genitalia( Fig. 23). Pygofer black, dorsal beak black, anal styles bright red; upper lobe black; basal lobe black. Uncus reddish-brown, orange-brown laterally; in lateral view beak-like; lobes in ventral view narrow, with rounded lateral termination; claspers clearly divided, short, with apices tapering laterally. Aedeagus with pseudoparameres extending around three-quarters the length of theca; theca recurved ventrally around 180° towards apex, with transparent flange along dorsal margin of distal half of recurvature, broadly smooth, equal to width of theca; apex short, knob-like, with 6–8 prominent cornuti. Female( Fig. 22C–E). Headand thoraxsimilar to male, and with patterns of black and brown markings within the range described for coastal and mountain male specimens. Abdomensimilar to coastal and mountain male specimens for the range of markings on tergites and sternites; sternites III–VII becoming more orange-red posteriorly, with midline black marking also reducing progressively from sternite III to sternite VII; abdominal segment 9 black dorsally with orange longitudinal stripes either side of midline, curving anteriorly and becoming orange to red laterally and ventrally; dorsal beak black. Ovipositor brown, black at tip, not extending beyond apex of abdominal segment 9. Anal styles bright red; ovipositor sheath dark brown to black. Measurements(in mm; range with mean in parentheses: 12 males, 12 females). Body length: male 19.6–22.7 (21.0); female 21.4–26.5 (22.6). Fore wing length: male 23.4–26.7 (24.8); female 26.3–28.4 (26.9). Head width: male 6.0–6.8 (6.5); female 6.9–7.3 (7.1). Pronotum width: male 6.0–6.7 (6.3); female 6.3–7.3 (6.8). Abdomen width: male 5.8–6.5 (6.4); female 6.0–7.3 (6.7). Ovipositor length 6.1– 6.8 (6.5).
Etymology. “ Spectabilis” (Latin) depicts the impressive coloration and markings that range from orange-red to enamel black in this species. Distinguishing features. Yoyetta spectabilis sp. nov.is readily distinguished from Y. denisoni, Y. timothyisp. Figure 25. Comparative structures of a single phrase from the calling songs of Yoyetta spectabilis sp. nov.from a range of locations, including: (i)Wises Track, Audley (34°08'S 151°03'E); (ii)Budderoo (34°39'S 150°45'E); (iii)Barren Grounds (34°41'S 150°44'E); (iv)Kanangra Boyd National Park (33°53'S 150°03'E); (v)Blackheath (33°35'S 150°50'E). Specimens were recorded in the field by LWP using RS1 (i, ii, iii, v) or RS2 (iv). nov., Y. kershawi stat. rev., comb. nov., Y. regalis sp. nov., Y. grandis sp. nov., Y. subalpina sp. nov., Y. grandis sp. nov.and Y. verrens sp. nov.by the following combination of characters: (1) smaller size with body length < 23 mm, (2) presence of symmetrical, orange triangular patterns on tergites 4–7, and (3) ovipositor sheaths of females not extending beyond the apex of abdominal segment 9. It differs from Y. hunterorum, which has the opercula and sternite 8 dull brown, the female abdominal segment 9 without red markings and the female ovipositor sheath extending> 1.5 mmbeyond the apex of the abdomen. The orange triangular patterning on the tergites of Y. spectabilis sp. nov.is distinct from the annular banding on Y. electrica sp. nov.which is also smaller (BL < 18 mm). Specimens of Y. spectabilis sp. nov.can be difficult to distinguish from Y. serrata sp. nov.and some specimens of Y. abdominalis, which have a similar range of shapes, sizes and markings. While the calling songs are quite distinctive, Y. spectabilis sp. nov.can also be distinguished from Y. abdominalisand Y. serrata sp. nov.by the following combination of characters: (1) lack of longitudinal central fascia on the pronotum in male specimens; (2), the colouration of the male opercula, which is predominantly red (cf. dull reddish-brown in Y. serrata sp. nov.and black to pale brown in Y. abdominalis); (3), the forewing costa and basal membrane is bright orange-red; (4) epipleurites boldly red and black. Calling song( Figs 24, 25, 53K). The male calling song of Y. spectabilis sp. nov.consists of a series of “whirling” amplitude modulations and produced at rest. Close examination reveals that these are organized into long echemes. The calling song commences with an introductory echeme with rapidly-emitted and regular amplitude modulations (“round whirling”; Fig. 24A–C). Towards the end of the echeme, the rate of modulation decreases, with each modulation commencing abruptly and fading (“abrupt whirling”; Fig. 24A,D,E). Each echeme ends with a sharp terminal modulation equivalent to a macrosyllable ( Fig. 24F). The first echeme is the longest, lasting about 5–8 s. The second echeme sometimes has the same form, though with a shorter round whirling section. The typical echemes following these introductory echeme(s) are simplified, losing the round whirling altogether and only exhibiting the abrupt whirling and sharp terminal modulation. A comparison between the typical echemes recorded at multiple sites in provided in Fig. 24. These typical echemes are around 2 s duration. The highest frequency plateau for the 513 calling song ranges between 8.9–13.0 kHz, with a dominant frequency between 9.5–12.1 kHz ( Fig. 24). Within the genus Yoyetta, the calling song of Y. spectabilis sp. nov.most closely resembles that of Y. regalis sp. nov.The song of Y. spectabilis sp. nov.differs most noticeably in the slower modulation rate and the presence of both round whirling and abrupt whirling typesof amplitude modulation (cf. one typeof rapidly emitted amplitude modulation in Y. regalis sp. nov.). Yoyettaspectabilis sp. nov.also does not produce intervening short chirps between bouts of echemes, which is a feature of the song of Y. regalis sp. nov.
2005-01-09
AM
S. N. & D. Emery
-34.114166
Wises Track
19
151.05833
Royal National Park
33
308
1
1
holotype
2009-12-01
2009-12-08
2009-12-01
Morton NP
Australia
Touga Rd
Malaise
33
308
2
1
1
New South Wales
paratype
1994-12-28
D. Yeates & B. Lessard & Bulk Sample No.
-35.043056
Clyde Mountain
19
150.14194
Australia
33
308
2
2
Manchester
paratype
1996-12-17
ANIC
A. Sundholm & De Keyzer & W. W. Froggatt
Blue Mtns
-33.70861
Waitara
19
151.10751
Park Rd
33
308
2
1
1
Manchester
paratype
1997-11-10
Moulds
-33.76278
Glenbrook
19
150.63805
Knapsack St
33
308
1
1
Manchester
paratype
1999-01-15
Emery
34
309
-34.711945
Barren Ground Nature Reserve
19
150.70583
33
308
2
1
1
Manchester
paratype
[166,648,1579,1599]
2000-01-15
Emery
-34.711945
Barren Ground Nature Reserve
19
150.70583
34
309
5
2
3
Manchester
paratype
2004-01-01
Emery
-34.711945
Barren Ground Nature Reserve
19
150.70583
34
309
6
6
Manchester
paratype
2000-12-06
S. & D. Emery & N. & D. Emery
-34.711945
Clarence
19
150.70583
34
309
4
4
Manchester
paratype
2002-11-23
R. Chin
-33.61222
Blackheath
19
150.32445
34
309
3
1
2
Manchester
paratype
2002-12-14
Emery & Popple
-33.61222
Blackheath
19
150.32445
34
309
1
1
Manchester
paratype
2002-12-22
Emery
-33.61222
Blackheath
19
150.32445
34
309
2
2
Manchester
paratype
2015-10-17
N. & D. Emery & S., N., C. & D. Emery
-35.02639
Berrara
19
150.47694
34
309
3
3
Manchester
paratype
2018-01-19
2018-01-28
2018-01-19
S., N. & D. Emery & T. Corbin
-36.42806
Dolphin Point
19
150.44444
34
309
3
3
Medway
paratype
[288,709,1795,1815]
2018-01-19
2018-01-28
2018-01-19
DE
Poyitt
-36.42806
Dolphin Point
19
150.44444
34
309
2
2
Medway
paratype
2014-01-02
2018-01-28
2018-01-19
DE
Poyitt
-36.42806
Dolphin Point
19
150.44444
34
309
1
1
Medway
paratype
[398,790,1819,1839]
1968-12-26
JP
J. Poyitt
Bendalong
34
309
1
1
Medway
paratype
[166,618,1843,1862]
1986-02-07
Daniels
Royal National Park
34
309
1
1
Medway
paratype
1983-11-13
C. N. Smithers
Tuglo
34
309
1
1
Medway
paratype
1992-12-28
B. Day & K. Kloo
Macquarie Pass
34
309
2
2
Medway
paratype
2006-01-10
Hanlon
Barren Ground Picnic Area
34
309
6
6
Medway
paratype
[407,790,1915,1935]
2002-11-23
AM
D. Britton
Blackheath
34
309
2
2
Medway
paratype
[166,790,1939,1959]
2003-11-31
L. Popple & D. Emery
Barren Grounds
34
309
1
1
Medway
paratype
[166,679,1963,1983]
2003-11-31
L. Popple & D. Emery
Jambaroo
Burradoo
34
309
3
3
Medway
paratype
2010-01-09
L. Popple & D. Emery
Genitalia
-33.988613
Heathland
19
150.09222
Kanangra Walls
34
309
3
3
Medway
paratype
2010-01-09
Popple & Emery
-33.886112
Kanangra-Boyd National Park
19
150.7564
34
309
1
1
Medway
paratype
2010-01
Popple & Emery
Blackheath
34
309
1
1
Medway
paratype
[905,1335,1627,1647]
1994-12-23
LWP
D. Emery
Ulladulla
34
309
1
1
Medway
paratype
1971-12-05
M. S. & B. J. Moulds
Mountain Nat. Pk
Hat Hill
34
309
5
5
Medway
paratype
1975-11-30
Daniels
Mountain Nat. Pk
Hat Hill
34
309
2
1
1
Medway
paratype
[1069,1363,1675,1695]
1973-12-23
Kanangra Walls
34
309
3
3
Medway
paratype
1990-12
G. Daniels
Blue Mountains
Wentworth Falls
34
309
1
1
Medway
paratype
1973-11-24
Keith Chambers
Blackheath
Hat Hill
34
309
4
1
3
Medway
paratype
1974-12-24
Daniels
Cudmirrah Faunal Reserve
34
309
1
1
Medway
paratype
1978-02-02
G. Daniels
Royal Nat. Pk
Mt Leighton Bailey
34
309
1
1
Medway
paratype
[958,1361,1771,1791]
1976-12-12
Daniels
8 km E Robertson
34
309
1
1
Medway
paratype
1990-12-12
G. Daniels
Ulladulla
George Boyd Lookout
34
309
1
1
Medway
paratype
1970-02-16
A. Sundholm
Chase National Park
Bobbin Head
34
309
1
1
Medway
paratype
1984-11-18
J. Barratt
Budderoo Fire Trail
34
309
1
1
Medway
paratype
1993-01-24
G. Williams
Currarong
34
309
1
1
Medway
paratype
1971-12-11
T. M. S. Hanlon
Hat Hill
Blue Mts
Blackheath
34
309
7
7
Medway
paratype
1975-11-30
J. V. Peters
Blue Mountains
Mt Wilson
34
309
1
1
Medway
paratype
[1078,1464,1915,1934]
1991-12-30
G. Daniels
Jenolan Caves
34
309
2
1
1
Medway
paratype
[839,1200,1939,1959]
1994-12-28
R. Eastwood
Clyde Mt
34
309
2
2
Medway
paratype
1961-12-10
R. de Keyzer & A. Sundholm
Tinderry Rg
34
309
1
1
Medway
paratype
2011-01-14
Greenup
Jct Rawson Pde
980
-33.703583
Leura
1
150.33847
Mt Hay Rd
34
309
1
1
Medway
paratype
1948-01-17
HEM
-31.866667
Barrington Tops
1214
151.41667
Tubrabucca
35
310
HEM2218
1
1
1921-12-17
MMV
-31.983334
Barrington
1213
151.91667
35
310
HEM2219
1
1
DE
N. & D. Emery
-33.756668
Jervis Bay
19
150.74861
35
310
2
2
Australian Capital Territory
1983-01-07
Shady Ck V. & D. F. Crosby
Yarragon
35
310
1
1
Victoria
1988-12-28
MSM
N. Walkerville & Abbott
35
310
2
1
1
Victoria
1976-12-28
ANIC
W. N. B. Quick
Moondarra Vic
Tyers River
35
310
2
2
Victoria
[257,745,1677,1697]
1952-01-05
HEM
-38.316666
Vic
1176
146.31667
Yinnar
35
310
HEM2237
1
1
Victoria
[121,573,1701,1721]
1948-12-16
MMV
-38.316666
Vic
1176
146.31667
Yinnar
35
310
HEM2238
1
1
Victoria
[190,504,1726,1745]
1998-01-13
QM
Upper
35
310
2
2
Victoria
1924-01-13
SAM
Berwick & G. J. Hall & Nat. Mus. Vict. & Database
G. J. Hall
Victoria
35
310
1
1
Victoria
2007-12-19
C. & D. Emery
-33.616386
Blackheath
19
150.30806
Hat Hill Rd.
35
310
4
4
New South Wales
2006-01-10
D. Emery
-33.616386
Blackheath
19
150.30806
Hat Hill Rd.
35
310
1
1
NEW SOUTH WALES
2005-01-09
2005-01-13
2005-01-09
S. N. & D. Emery
-34.114166
Royal National Park
19
151.05833
Wises Track
35
310
1
1
NEW SOUTH WALES
2007-01-22
2005-01-13
2005-01-09
D. Emery
-34.114166
Royal National Park
19
151.05833
Wises Track
35
310
4
4
NEW SOUTH WALES
2003-11-30
S. & D. Emery & L. Popple
-34.61222
Budaroo National Park
19
150.70139
35
310
6
1
5
NEW SOUTH WALES
[121,569,1966,1985]
2007-01-27
S. & D. Emery
-34.61222
Budaroo National Park
19
150.70139
35
310
18
7
11
NEW SOUTH WALES
2004-01-01
S. N. & D. Emery
-34.61222
Budaroo National Park
19
150.70139
35
310
8
8
NEW SOUTH WALES
[385,1090,1485,2009]
2009-12-06
D. Emery
-33.595554
Anvil Rock
19
150.33333
35
310
3
1
2
NEW SOUTH WALES
2009-12-09
D. Emery
-33.595554
Anvil Rock
19
150.33333
35
310
4
1
3
NEW SOUTH WALES
2009-12-14
D. Emery
-33.676113
Kuringai Chase National Park
19
151.21445
35
310
1
1
NEW SOUTH WALES
2010-01-05
2010-01-06
2010-01-05
D. Emery
-33.706112
St. Ives Showground
19
151.18916
35
310
2
2
NEW SOUTH WALES
2010-01-08
D. Emery & L. Popple
-33.988335
Kanangra Walls
19
150.04028
35
310
1
1
NEW SOUTH WALES
2010-12-28
T., S., N., C. & D. Emery
-34.694447
Barren Ground Reserve
19
150.73862
Kangaroo Track
35
310
7
1
6
NEW SOUTH WALES
2012-12-20
N. & D. Emery
-33.601387
Leura
19
150.33333
Lockley's Pylon Track
35
310
3
3
NEW SOUTH WALES
2012-12-20
A. Greenville & N. Emery
-33.601387
Leura
19
150.33333
Lockley's Pylon Track
35
310
5
5
NEW SOUTH WALES
2012-12-20
T., S., N., C. & D. Emery
-33.601387
Leura
19
150.33333
Lockley's Pylon Track
35
310
2
1
1
NEW SOUTH WALES
[909,1254,1702,1721]
2012-12-20
A. Schreiber
-33.601387
Leura
19
150.33333
Lockley's Pylon Track
35
310
1
1
NEW SOUTH WALES
2012-12-20
N. Emery
-33.601387
Leura
19
150.33333
Lockley's Pylon Track
35
310
1
1
NEW SOUTH WALES
[982,1418,1726,1745]
2012-01-19
DE
G. Bellis
Bundanoon
35
310
1
1
NEW SOUTH WALES
J. H. Ashton
VICTORIA
35
310
1
1
Victoria