A Revision of the Yoyetta abdominalis (Distant) Species Group of Cicadas (Hemiptera: Cicadidae: Cicadettinae), Introducing Eight New Species Emery, David L. Emery, Nathan J. Popple, Lindsay W. Records of the Australian Museum 2019 2019-10-02 71 7 277 347 8K2CN Emery & Emery & Popple, 2019 Emery & Emery & Popple 2019 [914,1179,1625,1657] Insecta Cicadidae Yoyetta Animalia Hemiptera 33 308 Arthropoda species spectabilis sp. nov.   Figs 2D, 3D, 22–25, 52D, 53K     Holotype ♂, Wises Track, Royal National Park, NSW, 34°06'51"S 151°03'30"E,  9.i.2005, S. N. & D. Emery( AMK.536052).   Paratypes NEW SOUTH WALES: 1♂ 1♀, AustraliaNSW, Morton NP, Malaiseon edge between open field & woodland  Leptospermumunderstorey, off Touga Rd,  1–8.xii.2009, C.  Manchester, D.Yeates, B. Lessard,ANIC Bulk Sample No.3381, 35°02'35"S 150°08'31"EGPS; 2♂♂, AustraliaNSW, Clyde Mountain(near top of highway pass),  28.xii.1994;  A. Sundholm& R. De Keyzer; 1♂, Blue Mtns1899, W. W. Froggattcollection ( ANIC); 1♀, Park Rd, Waitara, NSW, 33°42'31"S 151°06'27"E,  17.xii.1996, M. S. Moulds; 1♀, Knapsack St, Glenbrook, NSW, 33°45'46"S 150°38'17"E,  10.xi.1997, D. Emery; 1♂ 1♀, Barren Ground Nature Reserve, NSW, 34°42'43"S 150°42'21"E,  15.i.1999, S., N. & D. Emery; 3♂♂ 2♀♀same location,  15.i.2000, S. & D. Emery; 6♂♂same location,  1.i.2004,  S. & D. Emery; 2♂♂, same location, 29.xii, 2005, N. & D. Emery; 2♂♂, Clarence, NSW,  6.xii.2000,  R. Chin; 2♂♂ 1♀, Blackheath, NSW, 33°36'44"S 150°19'28"E,  23.xi.2002, D. Emery& L. Popple; 1♂, same location,  14.xii.2002, D. Emery; 2♂♂, same location,  22.xii.2002,  N. & D. Emery; 2♂♂,  20 kmSW Nowra, NSW, 35°01'35"S 150°28'37"E, 1.i.15, S., N., C. & D. Emery; 1♂, Berrara,  17.x.2015, N. Emery; 2♂♂,  Medway, NSW, 34°29'25"S 150°16'27"E, 6.i.17, S., N. & D. Emery& T. Corbin; 3♂♂, Dolphin Point, NSW, 36°25'41"S 150°26'40"E,  19–28.i.2018, J. Poyitt( DE); 2♂♂, same location,  19–28.i.2018, J. Poyitt; 1♂, same location,  2.i.2014,  J. Poyitt( JP); 1♂, Bendalong, NSW,  26.xii.1968, G. Daniels; 1♀, Royal National Park, NSW,  7.ii.1986,  C. N. Smithers; 1♂, “ Tuglo”,  60 kmN Singleton, NSW,  13.xi.1983,  B. Day& K. Kloo; 2♂♂, Macquarie Pass, NSW,  28.xii.1992, T. M. S. Hanlon; 6♂♂, Barren Ground Picnic Area, NSW,  10.i.2006,  D.Britton( AM); 2♂♂, Blackheath,  23.xi.2002,  L. Popple, D.Emery, 513-0001, 513-0002; 1♀, Barren Grounds,  31.xi.2003,  L. Popple, D. Emery; 3♂♂, Burradoovia Jambaroo,  31.xi.2003,  L.Popple, D. Emery, 513-0004 to 513-0006, 1♂, Genitaliaprep. 513-01; 2♂♂, Kanangra Walls, 33°59'19"S 150°05'32"E, S12588, Heathland,  9.i.2010,  Popple& Emery, 513-0008, 513-0009; 1♂, Kanangra-Boyd National Park, 33°53'10"S 150°45'23"E, S12588,  9.i.2010,  Popple& Emery, 513-0007; 1♂, Blackheath,  i.2010,  D. Emery, 513 ( LWP); 1♂, Ulladulla,  23.xii.1994,  M. S. & B. J. Moulds; 5♂♂, Hat Hill, Blue Mountain Nat. Pk,  5.xii.1971, G. Daniels; 1♂ 1♀, same data,  30.xi.1975;  3♂♂, Kanangra Walls,  23.xii.1973,  G. Daniels; 1♂, Wentworth Falls, Blue Mountains,  xii.1990,  Keith Chambers; 3♂♂ 1♀, Hat Hill, Blackheath,  24.xi.1973, G. Daniels; 1♂, Cudmirrah Faunal Reserve,  24.xii.1974,  G. Daniels; 1♂, Mt Leighton Bailey, Royal Nat. Pk,  2.ii.1978, G. Daniels; 1♂,  8 kmE Robertson,  12.xii.1976,  G. Daniels; 1♂, George Boyd Lookout, nr Ulladulla,  12.xii.1990,  A. Sundholm; 1♂, Bobbin Head, Ku-Ring-Gai Chase National Park,  16.ii.1970,  J. Barratt; 1♂, Budderoo Fire Trail, approx.  18 kmSE Robertson,  18.xi.1984,  G. Williams, ex. woodland; 1♂, Currarong,  24.i.1993,  T. M. S. Hanlon; 7♂♂, Hat Hill, Blackheath, Blue Mts,  11.xii.1971,  J. V. Peters; 1♂, Mt Wilson, Blue Mountains,  30.xi.1975,  G. Daniels; 1♂ 1♀, Jenolan Caves,  30.xii.1991,  R. Eastwood; 2♂♂, Clyde Mt,  28.xii.1994,  R. de Keyzer& A. Sundholm; 1♀, Tinderry Rg,  10.xii.1961, L. A. Greenup; 1♂, Jct Rawson Pdeand Mt Hay Rd, N. of Leura, 33°42.215'S 150°20.308'E,  980 m[elev.],  14.i.2011,   Figure 22.  Yoyetta spectabilis sp. nov., (A)male holotype, Kuringai Chase National Park (33°40'S 151°12'E), dorsal habitus; (B)male, ventral habitus; (C)female, Royal National Park (34°06'S 151°03'E), dorsal habitus; (D), female, Royal National Park, ventral habitus; (E), female, Barren Grounds (34°41'S 150°44'E), brown variant, dorsal habitus; (F), live male specimen, Blackheath (33°35'S 150°50'E), photo by NJE. Scale bar (for preserved specimens) = 10 mm.   Figure 23.  Yoyetta spectabilis sp. nov., illustration of male pygofer and internal genitalia; (a) viewed laterally from the left; (b)viewed ventrally; (c)aedeagus; and (d)apex of theca. Characters as depicted in Fig. 4. Specimen from Barren Grounds ( 34°41'S 150°44'E). K. Hill, D. Marshall,AU.NS.MHA, C. Simon lab voucher, Yoyetta“zizzer”, legs in ETOH, 11.AU.NS.MHA.03; ( MSM );  1♀, Tubrabucca, Barrington Tops, NSW, 31°52'S 151°25'E,  17.i.1948( HEM2218);  1♂, Barrington, NSW, 31°59'S 151°55'E,  17.xii.1921( HEM2219); ( MMV).  AUSTRALIAN CAPITAL TERRITORY: 2♂♂, Jervis Bay, ACT, 33°45'24"S 150°44'55"E, 12.xii.10, N. & D. Emery( DE).  VICTORIA: 1♂, Shady Ck V., 7 kmN. Yarragon,  7.i.1983, D. F. Crosby;  1♂ 1♀, N. Walkerville,  28.xii.1988, H. Abbott( MSM);  2♂♂, Tyers River, Moondarra Vic,  28.xii.1976, W. N. B. Quick( ANIC);  1♂, Yinnar, Vic, 38°19'S 146°19'E,  5.i.1952( HEM2237);  1♂, same location,  16.xii.1948( HEM2238) ( MMV ); 1♂, Narre Warren, 21.i.62;  2♀♀, UpperB’field,  13.i.1998( QM);  1♂, Berwick, Victoria, G. J. Hall,  13.i.1924,  Melampsalta abdominalisDist., det. G. J. Hallafter Nat. Mus. Vict., SAMA Databaseno. 20-014466 ( SAM).  Other material.  NEW SOUTH WALES: 4♂♂, Hat Hill Rd., Blackheath, NSW, 33°36'59"S 150°18'29"E,  19.xii.2007, C. & D. Emery;  1♀, same location,  10.i.2006, D. Emery;  1♂, Wises Track, Royal National Park, NSW, 34°06'51"S 151°03'30"E,  9–13.i.2005, S. N. & D.Emery;  4♂♂, same location,  22.i.2007, S., N. &, D. Emery;  5♂♂ 1♀, Budaroo National Park, NSW, 34°36'44"S 150°42'05"E,  30.xi.2003, S. & D. Emery& L. Popple;  11♂♂ 7♀♀, same location,  27.i.2007, S. & D. Emery;  8♂♂, same location,  1.i.2004, S. N. & D. Emery;  2♂♂ 1♀, Anvil Rock, NSW, 33°35'44"S 150°20'00"E,  6.xii.2009, D. Emery;  3♂♂ 1♀, same location,  9.xii.2009, D. Emery;  1♂, Kuringai Chase National Park, NSW, 33°40'34"S 151°12'52"E,  14.xii.2009, D. Emery;  2♂♂, St. Ives Showground, NSW, 33°42'22"S 151°11'21"E,  5–6.i.2010, D. Emery;  1♂, Kanangra Walls,NSW, 33°59'18"S 150°02'25"E,  8.i.2010, D. Emery& L. Popple;  6♂♂ 1♀, Kangaroo Track, Barren Ground Reserve, NSW, 34°41'40"S 150°44'19"E,  28.xii.2010, T., S., N., C. & D. Emery;  3♂♂, Lockley’s Pylon Track, Leura, NSW, 33°36'05"S 150°20'00"E,  20.xii.2012, N. & D. Emery;  5♂♂, same location, 7.xii.13, A. Greenville& N. Emery;  1♂ 1♀, same location, 2.xii.14, T., S., N., C. & D. Emery;  1♂, same location, 2.xii.14, A. Schreiber;  1♂, same location, 12.xii.14, N. Emery;  1♀, Bundanoon, NSW,  19.i.2012, G. Bellis( DE).  VICTORIA: 1♂, [labels in handwriting of J. H. Ashton] M. abdominalis Dist. Vic., K30137 [typed], genitalia preparation [handwritten] Y029, AM K.307108 (donated in 1911).   Distribution, habitat and seasonality. Highlandlocations from Barrington Tops south to the Southern Highlands of New South Wales. Specimens have been found in coastal locations from northern Sydney, south to eastern Victoria, close the Melbourne ( Fig. 18). Adults occur mainly on small heath shrubs and sometimes on eucalypts. Males constantly   Figure 24. Male calling song structure of  Yoyetta spectabilis sp. nov, illustrated in waveform plots, including (A)a series of echemes, including an introductory echeme and three typical echemes, illustrating the “round whirling” and “abrupt whirling” modulations (explained in the text); (B)expanded section of the round whirling from the introductory echeme (in A) showing rounded amplitude modulations; (C)further expanded detail of a round whirling amplitude modulation; (D)expanded detail of a typical echeme (in A) showing the abrupt whirling modulations; (E)expanded detail of an abrupt whirling amplitude modulation (from D); (F)expanded (from D) to show an abrupt modulation followed by the terminal modulation (explained in text). The final subfigure (G)is a spectrogram displaying song frequency. The specimen was recorded in the field at Wises Track, Audley (34°08'S 151°03'E) by LWP using RS1 (see Methods and terminology). move between singing stations that are usually < 3 mhigh. Females are located close to ground level. Since their song is somewhat similar to that of  Y. regalis sp. nov., the location of  Y. spectabilis sp. nov.on the heathland shrubs contrasts with the distinct preference of the former species for smooth-barked eucalypts when the two species cohabit open woodland. This species can be encountered from late November to January.   Description  Male( Figs 2D, 3D, 22A,B,F, 52D). Headalmost as wide as mesonotum, black in mountain specimens, with a small, triangular brown fascia posterior to ocelli, reducing anteriorly, and a small to large ochraceous spot on posterior margin between ocelli and eye; covered with black pubescence. In coastal specimens, including the holotype, pale brown to ochraceous posteriorly, extending halfway around eye, black surrounding ocelli, black spot medial to eye; ocelli pink; postclypeus dorsally dark brown to black with yellowish triangular median fascia, apex directed anteriorly, ventral surface black with black transverse grooves, midline with ochraceous line over anterior half in some specimens, lateral and posterior borders ochraceous, tending reddish; anteclypeus black, rostrum black at base, dark brown centrally, black at apex, reaching anterior edge of hind coxae; lora ochraceous anteriorly, variably black on medial and posterior sections; eyes variably dull black to dark brown; antennae black, supraantennal plates black, coastal specimens with central brown spot on anterior margin extending laterally above pedicel.  Thoraxpredominantly dull black on mountain specimens, brown on coastal cicadas. Pronotum black with distinct ochraceous marking over central third of midline, variably bordered by black, fissures variably black, tending brown on raised areas; pronotal collar brown centrally, variably tending black laterally, lateral angles black, paranota black on posterior and lateral margins, becoming brown elsewhere. Mesonotum black on mountain cicadas, coastal specimens brown with sigilla, scutal depressions and surrounds black; cruciform elevation arms black, lateral depressions brown. Metanotum black.  Legs.Coxae mainly black and ochraceous, fore coxae with anterior black markings ranging from proximal spot to a longitudinal stripe, mid and hind coxae black, variably ochraceous laterally, membranes red; meracantha small, narrow, reddish, black at base, pointed, overlapping onequarter of opercula; trochanters red with black medial stripe; femora red laterally, black medially; femoral spines erect, reddish to black at base tending dark brown at tips; fore tibia black, tending brown distally, browner on coastal specimens; mid tibiae ochraceous with proximal black stripe, hind tibiae pale brown with variable, small black marking on medial aspect of proximal segment; tarsi on coastal specimens reddish tending black at distal joints, fore tarsi brown on mountain specimens becoming black towards claws, mid tarsi light red with black at claws, hind tarsi red with brown at claws; claws dark brown, black at tips.  Wingswith fore wing costal veins orange-red, proximal margin black posteriorly; pterostigma mottled red; basal cell hyaline with black anterior border; basal membranes bright orange-red, other veins dark brown to black, with eight apical cells; hind wing plagas white over entire anal cell 3 and thin along vein 2A, some reddish stippling at base, with six apical cells.  Opercula( Fig. 3D) small, spatulate, following body axis ventrolaterally, depressed centrally, variably black at base, orange-red across remainder, clearly separated.  Timbals( Fig. 2D) with five distinct long ribs; long ribs 1–3 extending across surrounding membrane and fused dorsally along basal spur; long rib 4 discontinuous at proximal third; long rib 5 independent of basal spur, comparatively shorter, extending ventrally across half of membrane; large ridged dome on posterior timbal plate extending across two-thirds of timbal; apodeme pit oval-shaped and conspicuous.  Abdomen. Tergite 1 black with red lateral membranes; tergites 2–3 black with diffuse dark brown shading on some coastal specimens, intersegmental membranes orange; tergites 4–7 black with thin orange posterior margins, an increasing area of lateral orange coloration posteriorly forming a triangular shape with base on tergite 7, extending to lateral border and to epipleurite 7; tergite 8 shiny black, with orange-red posteriorly and extending laterally in some specimens. Epipleurites with variable black medial markings, fiery red laterally and posteriorly. Sternite II mainly black; sternite III black centrally, orange-red laterally; sternites IV–VI orange, posterior halves becoming red, black midline markings over anterior two-thirds of each sternite, decreasing posteriorly, sternite VIII fiery red, with yellowish pubescence ( Fig. 51D).  Genitalia( Fig. 23). Pygofer black, dorsal beak black, anal styles bright red; upper lobe black; basal lobe black. Uncus reddish-brown, orange-brown laterally; in lateral view beak-like; lobes in ventral view narrow, with rounded lateral termination; claspers clearly divided, short, with apices tapering laterally. Aedeagus with pseudoparameres extending around three-quarters the length of theca; theca recurved ventrally around 180° towards apex, with transparent flange along dorsal margin of distal half of recurvature, broadly smooth, equal to width of theca; apex short, knob-like, with 6–8 prominent cornuti.  Female( Fig. 22C–E). Headand thoraxsimilar to male, and with patterns of black and brown markings within the range described for coastal and mountain male specimens.  Abdomensimilar to coastal and mountain male specimens for the range of markings on tergites and sternites; sternites III–VII becoming more orange-red posteriorly, with midline black marking also reducing progressively from sternite III to sternite VII; abdominal segment 9 black dorsally with orange longitudinal stripes either side of midline, curving anteriorly and becoming orange to red laterally and ventrally; dorsal beak black. Ovipositor brown, black at tip, not extending beyond apex of abdominal segment 9. Anal styles bright red; ovipositor sheath dark brown to black.  Measurements(in mm; range with mean in parentheses: 12 males, 12 females). Body length: male 19.6–22.7 (21.0); female 21.4–26.5 (22.6). Fore wing length: male 23.4–26.7 (24.8); female 26.3–28.4 (26.9). Head width: male 6.0–6.8 (6.5); female 6.9–7.3 (7.1). Pronotum width: male 6.0–6.7 (6.3); female 6.3–7.3 (6.8). Abdomen width: male 5.8–6.5 (6.4); female 6.0–7.3 (6.7). Ovipositor length 6.1– 6.8 (6.5).   Etymology. “ Spectabilis” (Latin) depicts the impressive coloration and markings that range from orange-red to enamel black in this species.  Distinguishing features.  Yoyetta spectabilis sp. nov.is readily distinguished from  Y. denisoni, Y. timothyisp.   Figure 25. Comparative structures of a single phrase from the calling songs of  Yoyetta spectabilis sp. nov.from a range of locations, including: (i)Wises Track, Audley (34°08'S 151°03'E); (ii)Budderoo (34°39'S 150°45'E); (iii)Barren Grounds (34°41'S 150°44'E); (iv)Kanangra Boyd National Park (33°53'S 150°03'E); (v)Blackheath (33°35'S 150°50'E). Specimens were recorded in the field by LWP using RS1 (i, ii, iii, v) or RS2 (iv). nov.,  Y. kershawi stat. rev., comb. nov.,  Y. regalis sp. nov.,  Y. grandis sp. nov.,  Y. subalpina sp. nov.,  Y. grandis sp. nov.and  Y. verrens sp. nov.by the following combination of characters: (1) smaller size with body length < 23 mm, (2) presence of symmetrical, orange triangular patterns on tergites 4–7, and (3) ovipositor sheaths of females not extending beyond the apex of abdominal segment 9. It differs from  Y. hunterorum, which has the opercula and sternite 8 dull brown, the female abdominal segment 9 without red markings and the female ovipositor sheath extending> 1.5 mmbeyond the apex of the abdomen. The orange triangular patterning on the tergites of  Y. spectabilis sp. nov.is distinct from the annular banding on  Y. electrica sp. nov.which is also smaller (BL < 18 mm). Specimens of  Y. spectabilis sp. nov.can be difficult to distinguish from  Y. serrata sp. nov.and some specimens of  Y. abdominalis, which have a similar range of shapes, sizes and markings. While the calling songs are quite distinctive,  Y. spectabilis sp. nov.can also be distinguished from  Y. abdominalisand  Y. serrata sp. nov.by the following combination of characters: (1) lack of longitudinal central fascia on the pronotum in male specimens; (2), the colouration of the male opercula, which is predominantly red (cf. dull reddish-brown in  Y. serrata sp. nov.and black to pale brown in  Y. abdominalis); (3), the forewing costa and basal membrane is bright orange-red; (4) epipleurites boldly red and black.  Calling song( Figs 24, 25, 53K). The male calling song of  Y. spectabilis sp. nov.consists of a series of “whirling” amplitude modulations and produced at rest. Close examination reveals that these are organized into long echemes. The calling song commences with an introductory echeme with rapidly-emitted and regular amplitude modulations (“round whirling”; Fig. 24A–C). Towards the end of the echeme, the rate of modulation decreases, with each modulation commencing abruptly and fading (“abrupt whirling”; Fig. 24A,D,E). Each echeme ends with a sharp terminal modulation equivalent to a macrosyllable ( Fig. 24F). The first echeme is the longest, lasting about 5–8 s. The second echeme sometimes has the same form, though with a shorter round whirling section. The typical echemes following these introductory echeme(s) are simplified, losing the round whirling altogether and only exhibiting the abrupt whirling and sharp terminal modulation. A comparison between the typical echemes recorded at multiple sites in provided in Fig. 24. These typical echemes are around 2 s duration. The highest frequency plateau for the 513 calling song ranges between 8.9–13.0 kHz, with a dominant frequency between 9.5–12.1 kHz ( Fig. 24). Within the genus  Yoyetta, the calling song of  Y. spectabilis sp. nov.most closely resembles that of  Y. regalis sp. nov.The song of  Y. spectabilis sp. nov.differs most noticeably in the slower modulation rate and the presence of both round whirling and abrupt whirling typesof amplitude modulation (cf. one typeof rapidly emitted amplitude modulation in  Y. regalis sp. nov.).  Yoyettaspectabilis sp. nov.also does not produce intervening short chirps between bouts of echemes, which is a feature of the song of  Y. regalis sp. nov. 2005-01-09 AM S. N. & D. Emery -34.114166 Wises Track 19 151.05833 Royal National Park 33 308 1 1 holotype 2009-12-01 2009-12-08 2009-12-01 Morton NP Australia Touga Rd Malaise 33 308 2 1 1 New South Wales paratype 1994-12-28 D. Yeates & B. Lessard & Bulk Sample No. -35.043056 Clyde Mountain 19 150.14194 Australia 33 308 2 2 Manchester paratype 1996-12-17 ANIC A. Sundholm & De Keyzer & W. W. Froggatt Blue Mtns -33.70861 Waitara 19 151.10751 Park Rd 33 308 2 1 1 Manchester paratype 1997-11-10 Moulds -33.76278 Glenbrook 19 150.63805 Knapsack St 33 308 1 1 Manchester paratype 1999-01-15 Emery 34 309 -34.711945 Barren Ground Nature Reserve 19 150.70583 33 308 2 1 1 Manchester paratype [166,648,1579,1599] 2000-01-15 Emery -34.711945 Barren Ground Nature Reserve 19 150.70583 34 309 5 2 3 Manchester paratype 2004-01-01 Emery -34.711945 Barren Ground Nature Reserve 19 150.70583 34 309 6 6 Manchester paratype 2000-12-06 S. & D. Emery & N. & D. Emery -34.711945 Clarence 19 150.70583 34 309 4 4 Manchester paratype 2002-11-23 R. Chin -33.61222 Blackheath 19 150.32445 34 309 3 1 2 Manchester paratype 2002-12-14 Emery & Popple -33.61222 Blackheath 19 150.32445 34 309 1 1 Manchester paratype 2002-12-22 Emery -33.61222 Blackheath 19 150.32445 34 309 2 2 Manchester paratype 2015-10-17 N. & D. Emery & S., N., C. & D. Emery -35.02639 Berrara 19 150.47694 34 309 3 3 Manchester paratype 2018-01-19 2018-01-28 2018-01-19 S., N. & D. Emery & T. Corbin -36.42806 Dolphin Point 19 150.44444 34 309 3 3 Medway paratype [288,709,1795,1815] 2018-01-19 2018-01-28 2018-01-19 DE Poyitt -36.42806 Dolphin Point 19 150.44444 34 309 2 2 Medway paratype 2014-01-02 2018-01-28 2018-01-19 DE Poyitt -36.42806 Dolphin Point 19 150.44444 34 309 1 1 Medway paratype [398,790,1819,1839] 1968-12-26 JP J. Poyitt Bendalong 34 309 1 1 Medway paratype [166,618,1843,1862] 1986-02-07 Daniels Royal National Park 34 309 1 1 Medway paratype 1983-11-13 C. N. Smithers Tuglo 34 309 1 1 Medway paratype 1992-12-28 B. Day & K. Kloo Macquarie Pass 34 309 2 2 Medway paratype 2006-01-10 Hanlon Barren Ground Picnic Area 34 309 6 6 Medway paratype [407,790,1915,1935] 2002-11-23 AM D. Britton Blackheath 34 309 2 2 Medway paratype [166,790,1939,1959] 2003-11-31 L. Popple & D. Emery Barren Grounds 34 309 1 1 Medway paratype [166,679,1963,1983] 2003-11-31 L. Popple & D. Emery Jambaroo Burradoo 34 309 3 3 Medway paratype 2010-01-09 L. Popple & D. Emery Genitalia -33.988613 Heathland 19 150.09222 Kanangra Walls 34 309 3 3 Medway paratype 2010-01-09 Popple & Emery -33.886112 Kanangra-Boyd National Park 19 150.7564 34 309 1 1 Medway paratype 2010-01 Popple & Emery Blackheath 34 309 1 1 Medway paratype [905,1335,1627,1647] 1994-12-23 LWP D. Emery Ulladulla 34 309 1 1 Medway paratype 1971-12-05 M. S. & B. J. Moulds Mountain Nat. Pk Hat Hill 34 309 5 5 Medway paratype 1975-11-30 Daniels Mountain Nat. Pk Hat Hill 34 309 2 1 1 Medway paratype [1069,1363,1675,1695] 1973-12-23 Kanangra Walls 34 309 3 3 Medway paratype 1990-12 G. Daniels Blue Mountains Wentworth Falls 34 309 1 1 Medway paratype 1973-11-24 Keith Chambers Blackheath Hat Hill 34 309 4 1 3 Medway paratype 1974-12-24 Daniels Cudmirrah Faunal Reserve 34 309 1 1 Medway paratype 1978-02-02 G. Daniels Royal Nat. Pk Mt Leighton Bailey 34 309 1 1 Medway paratype [958,1361,1771,1791] 1976-12-12 Daniels 8 km E Robertson 34 309 1 1 Medway paratype 1990-12-12 G. Daniels Ulladulla George Boyd Lookout 34 309 1 1 Medway paratype 1970-02-16 A. Sundholm Chase National Park Bobbin Head 34 309 1 1 Medway paratype 1984-11-18 J. Barratt Budderoo Fire Trail 34 309 1 1 Medway paratype 1993-01-24 G. Williams Currarong 34 309 1 1 Medway paratype 1971-12-11 T. M. S. Hanlon Hat Hill Blue Mts Blackheath 34 309 7 7 Medway paratype 1975-11-30 J. V. Peters Blue Mountains Mt Wilson 34 309 1 1 Medway paratype [1078,1464,1915,1934] 1991-12-30 G. Daniels Jenolan Caves 34 309 2 1 1 Medway paratype [839,1200,1939,1959] 1994-12-28 R. Eastwood Clyde Mt 34 309 2 2 Medway paratype 1961-12-10 R. de Keyzer & A. Sundholm Tinderry Rg 34 309 1 1 Medway paratype 2011-01-14 Greenup Jct Rawson Pde 980 -33.703583 Leura 1 150.33847 Mt Hay Rd 34 309 1 1 Medway paratype 1948-01-17 HEM -31.866667 Barrington Tops 1214 151.41667 Tubrabucca 35 310 HEM2218 1 1 1921-12-17 MMV -31.983334 Barrington 1213 151.91667 35 310 HEM2219 1 1 DE N. & D. Emery -33.756668 Jervis Bay 19 150.74861 35 310 2 2 Australian Capital Territory 1983-01-07 Shady Ck V. & D. F. Crosby Yarragon 35 310 1 1 Victoria 1988-12-28 MSM N. Walkerville & Abbott 35 310 2 1 1 Victoria 1976-12-28 ANIC W. N. B. Quick Moondarra Vic Tyers River 35 310 2 2 Victoria [257,745,1677,1697] 1952-01-05 HEM -38.316666 Vic 1176 146.31667 Yinnar 35 310 HEM2237 1 1 Victoria [121,573,1701,1721] 1948-12-16 MMV -38.316666 Vic 1176 146.31667 Yinnar 35 310 HEM2238 1 1 Victoria [190,504,1726,1745] 1998-01-13 QM Upper 35 310 2 2 Victoria 1924-01-13 SAM Berwick & G. J. Hall & Nat. Mus. Vict. & Database G. J. Hall Victoria 35 310 1 1 Victoria 2007-12-19 C. & D. Emery -33.616386 Blackheath 19 150.30806 Hat Hill Rd. 35 310 4 4 New South Wales 2006-01-10 D. Emery -33.616386 Blackheath 19 150.30806 Hat Hill Rd. 35 310 1 1 NEW SOUTH WALES 2005-01-09 2005-01-13 2005-01-09 S. N. & D. Emery -34.114166 Royal National Park 19 151.05833 Wises Track 35 310 1 1 NEW SOUTH WALES 2007-01-22 2005-01-13 2005-01-09 D. Emery -34.114166 Royal National Park 19 151.05833 Wises Track 35 310 4 4 NEW SOUTH WALES 2003-11-30 S. & D. Emery & L. Popple -34.61222 Budaroo National Park 19 150.70139 35 310 6 1 5 NEW SOUTH WALES [121,569,1966,1985] 2007-01-27 S. & D. Emery -34.61222 Budaroo National Park 19 150.70139 35 310 18 7 11 NEW SOUTH WALES 2004-01-01 S. N. & D. Emery -34.61222 Budaroo National Park 19 150.70139 35 310 8 8 NEW SOUTH WALES [385,1090,1485,2009] 2009-12-06 D. Emery -33.595554 Anvil Rock 19 150.33333 35 310 3 1 2 NEW SOUTH WALES 2009-12-09 D. Emery -33.595554 Anvil Rock 19 150.33333 35 310 4 1 3 NEW SOUTH WALES 2009-12-14 D. Emery -33.676113 Kuringai Chase National Park 19 151.21445 35 310 1 1 NEW SOUTH WALES 2010-01-05 2010-01-06 2010-01-05 D. Emery -33.706112 St. Ives Showground 19 151.18916 35 310 2 2 NEW SOUTH WALES 2010-01-08 D. Emery & L. Popple -33.988335 Kanangra Walls 19 150.04028 35 310 1 1 NEW SOUTH WALES 2010-12-28 T., S., N., C. & D. Emery -34.694447 Barren Ground Reserve 19 150.73862 Kangaroo Track 35 310 7 1 6 NEW SOUTH WALES 2012-12-20 N. & D. Emery -33.601387 Leura 19 150.33333 Lockley's Pylon Track 35 310 3 3 NEW SOUTH WALES 2012-12-20 A. Greenville & N. Emery -33.601387 Leura 19 150.33333 Lockley's Pylon Track 35 310 5 5 NEW SOUTH WALES 2012-12-20 T., S., N., C. & D. Emery -33.601387 Leura 19 150.33333 Lockley's Pylon Track 35 310 2 1 1 NEW SOUTH WALES [909,1254,1702,1721] 2012-12-20 A. Schreiber -33.601387 Leura 19 150.33333 Lockley's Pylon Track 35 310 1 1 NEW SOUTH WALES 2012-12-20 N. Emery -33.601387 Leura 19 150.33333 Lockley's Pylon Track 35 310 1 1 NEW SOUTH WALES [982,1418,1726,1745] 2012-01-19 DE G. Bellis Bundanoon 35 310 1 1 NEW SOUTH WALES J. H. Ashton VICTORIA 35 310 1 1 Victoria