A Revision of the Yoyetta abdominalis (Distant) Species Group of Cicadas (Hemiptera: Cicadidae: Cicadettinae), Introducing Eight New Species Emery, David L. Emery, Nathan J. Popple, Lindsay W. Records of the Australian Museum 2019 2019-10-02 71 7 277 347 8K2CP Emery & Emery & Popple, 2019 Emery & Emery & Popple 2019 [963,1221,1423,1454] Insecta Cicadidae Yoyetta Animalia Hemiptera 38 313 Arthropoda species subalpina sp. nov.   Figs 2E, 3E, 26– 31, 52E, 53G     Holotype ♂, 18 kmE Cooma, NSW, 36°20'47"S 148°13'57"E,  10.xii.2010, L. Popple, 514-0013 ( AMK.559471).   Paratypes NEW SOUTH WALES: 1♂ 1♀, Piccadilly Circus, Brindabella Ra., ACT, 35°21'S 148°48'E,  26.xii.1978, D. C. F. Rentz, ANIC Databaseno. 20 010822;  1♀, Snowy River, Mt Kosciusko, 4000FT,  12.xii.1931, L. F. Graham, ANIC Database No.20 010821;  1♂, Old AdaminabyNSW,  24.i.1967, T. G. Campbell, ANIC Databaseno. 20 010820;  2♀♀, NSW Snowy Plains,  19.ii.1969, M. G. Stanger, on Eucalypt, [ 1♀] ANIC Databaseno. 20 010823;  1♀, CoomaNSW, 10.i.[19]06, Sellar, W. W. Froggattcollection, ANIC Database No.20 010815;  1♂, Tinderry Mountains, Round Flat, netted,  1142 m,  25.xii.2014, 35°42'52.08"S 149°17'40.48"E, D. J. & R. L. Ferguson, ANIC Database No.20 010718;  1♂, Tinderry Mountains, Tinderry Rdverge, netted,  1160 m, 35°43'49.17"S 149°18'07.92"E,  21.xii.2014, D.J. Ferguson, ANIC Database No.20 010816 ( ANIC);  2♂♂ 4♀♀, Mt Kosciusko, NSW, 36°27'S 148°16'E,  18.i.1931( HEM  2554–58);  1♀, JindabyneNSW, 36°25'S 148°37'E,  17.i.1953, ( HEM2559) ( MMV);  4♂♂, 3♀♀, 1 kmS. of Brown Mountain, via Nimmitabel, 36°36'16"S 149°22'31"E,  29.xii.2009, L. W. Popple, 514-0003 to 514-0009;  1♂, Tom Groggin Road,  18 kmESE of Cooma, 36°20'48"S 148°13'57"E,  10.xii.2010, L. W. Popple, 514-0014 ( LWP);  1♀, 7 miles[SW of?] Countegany on Kybean Rd, 16.xii.[19]62, L. R. G[reenup]?, 105;  1♂ 8♀♀, Mt David,  30 kmSW of Oberon,  26.xii.1985, S. &. B. Underwood;  1♂ 2♀♀, 12 mi.from Cooma on Nimitabelle[sic.] Rd,  27.xii.1973, Euc.pauciflora, L. R. G[reenup];  3♂♂ 2♀♀, Yarrangobilly Cavesroad, KosciuszkoNP, 35°33.258'S 148°30.842'E,  17.i.2011, 1305 m [elev.], K. Hill, D. Marshall, AU.NS.YAX, C. Simonlab voucher, Yoyetta“double-zit”;  1♂, same data; legs in ETOH, body pinned, 11.AU.NS.YAX.01 ( MSM);  1♂ 6♀♀, Piccadilly Circus, Brindabella Ranges, NSW, 35°21'34"S 148°47'59"E,  10–14.i.2002, D.Emery;  2♀♀, Kanangra Boyd National Park, NSW, 33°59'18"S 150°02'25"E,  8.i.2010, D. Emery& L. Popple(DE).  VICTORIA: 1♂ Mt Buffalo Nat. Pk,  16.i.1966, T.Weir( UQIC);  1♀, 36°29'S 147°54'E, 8 kmSE by E of Nariel Vic.,  16.i.1984, K. H.L. Key;  1♀,  9 kmE of Mt Pendergast, E of Benambra, Vic.,  1325m, 36°53'S 148°05'E,  29.xii.1983, K. H. L. Key, ANIC Database No.20 010819 ( ANIC);  3♂♂ 3♀♀, midway between Harrietville& St Bernard Hospice,  19 Dec 1933, A. Musgrave( AMK.307236–41);  1♂, Sommerville Ave, Lorne, Vic,  15.xii.2002, D. J. Hilton;  2♀♀,  1 kmSE Forrest, Otway Ranges, Vic, 38°54'39"S143°74'40"E, 5.ii.15, S. Emery& T. Corbin;  1♂, Kinglake, Vic, 37°28'11"S 145°20'12"E,  17.xii.2016, S. Emery& T. Corbin;  2♂♂, Country Fire Authority Station, Toolangi, 37°32'32"S 145°28'29"E,  11.i.2017, S. Emery& T. Corbin(to light);  2♂♂, same location,  23.i.2017, N., C. & D. Emery(DE);  2♂♂ 9♀♀, Mt Hotham Alpine Reserve,  1,830 m[elev.],  2.i.1976, M. S. & B. J. Moulds( 1♂ 1♀in cop);  4♂♂, same data,  1.i.1976,  1♂in cop; Mt Hotham Village, 36°59'S 147°11'E,  1–13.i.1990, G. R. Brown& M. A. Terras( MSM);  1♂, Kinglake, Vic, 37°32'S 145°20'E,  4.xii.1927, ( HEM2611);  5♀♀, Mt Buffalo, Vic, 36°43'S 146°46'E,  29.xii.1951( HEM  2612–16);  1♂, Country Fire Authority Station, Toolangi, 37°32'32"S 145°28'29"E,  27.xii.2018, S. Emery& T. Corbin(to light; T 22389); 5♂♂ 3♀♀, Allambee,  Victoria,  14.i.2019, J. Creen( HEM5765–7[ ♀], HEM5768–72 [ ♂]) ( MMV).  AUSTRALIAN CAPITAL TERRITORY. 1♀, 35°21S 148°48E, Piccadilly Circus, Brindabella Ra. A.C.T.,  26.xii.1978, D. C. F. Rentz( ANIC);  1♂, Mt Franklin Rd, Brindabella Ranges, 35°32'S 148°46'E,  5.xii.2004, B.Kenway,514-0001;  1♂, Piccadilly Circus, Brindabella Range, 35°21'37"S 148°47'59"E,  28.xii.2009, L. W. Popple, 514-0002, LT2009371;  1♂ 1♀, 1.5 kmN. of Bull’s Head, 35°22'39"S 148°48'16"E,  15.i.2011, L. W. Popple, 514-0015, 514-0017, LT2009371;  1♀, Mt Franklin area, 35°29'23"S 148°45'48"E,  15.i.2011, L. W. Popple, 514-0018, LT2009371 ( LWP).   Figure 26. Map of eastern Australia showing the geographical distribution of  Yoyetta subalpina sp. nov.(open triangles),  Y. serrata sp. nov.(closed squares) and  Y. grandis sp. nov.(open circles).  Additional records from song recordings  NEW SOUTH WALES: Rocky Pic Road, Talleganda National Park, NSW, 35°37'12"S 149°29'54"E,  5.xii.2009, L. W. Popple.  VICTORIA: Ridge Rd, Mt Dandenong,  600 m, 37°49'28"S 145°21'34"E,  15.xii.2017, T. Corbin.   Distribution, habitat and seasonality. Cool temperate alpine areas from Kanangra-Boyd National Park in central New South Walessouth to the Upper Yarra region in central Victoria, with an isolated population in the Otway Ranges ( Fig. 26). Adults occur mainly on smooth-barked eucalypts on the main trunks and upper branches. They can be encountered from December to February.   Description  Male( Figs 2E, 3E, 27A– B, 27F, 51G, 52E). Headas wide as mesonotum, black, with an ochraceous diamond-shaped marking on midline, posterior to ocelli; ocelli pink to pale; dorsal postclypeus black with central ochraceous triangular marking, apex directed anteriorly; ventral postclypeus black with anterior brown marking on midline, and ochraceous, lateral and posterior margins; anteclypeus black, rostrum brown at base, becoming black at apex, reaching the middle of hind coxae; lora black with ochraceous anterior margin, gena black; eyes dull black; antennae black, supra-antennal plates black, ochraceous around base of pedicel.  Thoraxpredominantly black. Pronotum black, some specimens with brown in fissures; pronotal collar black, posterior margin ochraceous in some specimens, margins of lateral angles light brown, paranota black, anterior angles ochraceous in some specimens. Mesonotum black; cruciform elevation arms black, lateral depressions tinged brown. Metanotum black.  Legs.Coxae mainly black, fore coxae black with proximal ochraceous triangular stripe on posterior surface; mid and hind coxae black; coxal joint and membranes red, basisterna black; trochanters black with central segment orange-red; meracantha small, narrow, orange, black at base, pointed, minimally overlapping opercula; fore femora striped longitudinally orange-red and black, black along base of femoral spines femoral spines erect, black at base tending dark brown at tips; mid and hind femora black, orange-brown longitudinal   Figure 28.  Yoyetta subalpina sp. nov., illustration of male pygofer and internal genitalia; (a) viewed laterally from the left; (b)viewed ventrally; (c)aedeagus; and (d)apex of theca. Characters as depicted in Fig. 4. Specimen from Piccadilly Circus ( 35°22'S 148°48'E). stripe on medial aspect, distal femoral joint orange-red; fore tibiae black, brown at distal tip; mid tibiae black, with medial ochraceous stripe; hind tibiae black proximally ochraceous over distal half; fore tarsi black with brown distal margins; mid and hind tarsi blackish-brown becoming black towards claws; claws dark brown, black at tips.  Wingswith fore wing costal veins black, browner central rib, becoming ochraceous distally; pterostigma mottled red; basal cell translucent with black anterior border; basal membranes bright orange; other venation dark brown to black; with eight apical cells; hind wing plagas white over entire anal cell 3 and medial third of ac2(v), thin along vein 2A, an irregular grey-black marking at proximal end of jugum, reddish stippling at base of ac2(v), veins 2A and 3A pale, remainder dark brown to black, with six apical cells.  Opercula( Fig. 3E) medium, spatulate, following body axis ventrolaterally, depressed centrally, variably black over basal half, orange-brown across remainder, clearly separated.  Timbals( Fig. 2E) with five distinct long ribs; long ribs 1–4 extending across surrounding membrane and fused dorsally along basal spur; long rib 5 independent of basal spur, comparatively shorter, extending ventrally across half of membrane; large ridged dome on posterior timbal plate extending across three-quarters of timbal; apodeme pit ovalshaped and conspicuous.  Abdomen. Tergite 1 black, with orange-red membranes around timbal cavity; tergite 2 black; tergites 3–7 black with orange posterior margins, extending laterally to epipleurites, increasing on tergites 6–7 on either side of midline; tergite 8 shiny black, with ventral margins orange. Epipleurites with medial black strip, orange over reminder. Sternite II mainly black, with central orange spot; sternite III black medially, becoming orange laterally along posterior margin; sternites IV–VI orange, posterior halves becoming progressively reddish, diffuse black midline markings over anterior twothirds of each sternite, decreasing posteriorly, sternite VIII dull orange-red, with black pubescence.   Figure 29. Typical male calling song structure of  Yoyetta subalpina sp. nov., illustrated in waveform plots, including (A)two phrases; (B)expanded section from A showing a single phrase with an echeme followed by a long macrosyllable; (C)expanded section from B showing more detail from the echeme and the macrosyllable. The final subfigure (D)is a spectrogram displaying song frequency. The specimen was recorded in the field at Piccadilly Circus (35°22'S 148°48'E) by LWP using RS2 (see Methods and terminology).   Figure 30. Comparative male calling song structure of  Yoyetta subalpina sp. nov., illustrated in waveform plots, including (A)a 20 s section containing echemes, interspersed with a “soft ticking” segment; (B)expanded “soft ticking” section (from A) showing 10 syllables.The final subfigure (C)is a spectrogram displaying song frequency. The specimen was recorded in the field at Talleganda (35°37'S 149°30'E) by LWP using RS2 (see Methods and terminology).  Genitalia( Fig. 28). Pygofer black, upper lobe mainly black, apex brown, basal lobe black; dorsal beak black, dark brown posterior margin, anal styles orange-red. Uncus reddishbrown; in lateral view beak-like and elongated; lobes in ventral view narrow, with rounded lateral termination; claspers clearly divided, short, bulbous, with apices tapering laterally.Aedeagus with pseudoparameres almost extending as far as theca; theca recurved ventrally at around 180° towards apex, with prominent transparent flange along distal margin of recurvature, smooth and broadening to> 3× width of theca; remainder of theca is short, with <20% of theca extending apically beyond margin of recurvature; apex blunt, with 2–3 cornuti on ventral surface and several small cornuti on dorsal side at tip.  Female( Fig. 27C–E). Head, thoraxand abdomenof dark specimens similar to male. Paler, brown-coloured females apparently occur at a ratio of approximately 1:2.  Headin dark specimens, similar to male; in paler specimens mainly brown with black markings around ocelli extending to the supra-antennal plate, brown at anterior margin, black spot at medial border of eye; dorsal postclypeus ochraceous centrally, dark brown laterally; ventral postclypeus black with central ochraceous spot anteriorly; anteclypeus and rostrum similar to male; gena variably ochraceous and black, lateral margins ochraceous posteriorly, black anteriorly.  Thoraxin dark specimens, similar to male. In paler specimens; pronotum with black markings restricted to lateral and posterior margins of a broad ochraceous midline strip over central third, and variably along fissures and lateral margins; pronotal collar brown, with lateral angles black; mesonotum brown with black submedian and lateral sigilla, and a diffuse black central triangular marking with base around scutal depressions, apex directed anteriorly on midline to posterior third of submedian sigilla; cruciform elevation black on central, medial portion and lateral margins, yellow-ochraceous on posterior margins, lateral depressions ochraceous, anterior and posterior depressions dark brown to black; scutal depressions black; metanotum black.  Abdomen.Colour variation similar to thorax. On dark specimens, similar to male; abdominal segment 9 black dorsally with ochraceous triangular markings either side of midline, tapering posteriorly, ending midway to dorsal beak, becoming orange laterally, reddish on ventral margin. On brown specimens; tergite 1 black, tergite 2 black on midline, orange merging to black laterally; tergites 3–8 black with orange posterior margins, increasing posteriorly, extending laterally to epipleurites; epipleurites with medial black strip, orange over remainder; sternite II black; sternites III–VI pale brown with midline black marking; sternite VII pale brown with anterior black spot either side of midline; abdominal segment 9 orange-brown with three longitudinal black stripes faint on midline and prominent either side of midline. On all specimens, dorsal beak orange to black at base, black at tip; ovipositor dark brown, becoming black at tip, extending < 1 mmbeyond apex of abdominal segment 9. Anal styles orange-red; ovipositor sheath dark brown to black.  Measurements(in mm; range with mean in parentheses: 10 males, 8 females). Body length: male 25.0–25.1 (25.1); female 24.0–27.0 (26.0). Fore wing length: male 27.7– 28.5 (28.1); female 29.4–33.2 (31.9). Head width: male 7.0–7.6 (7.3); female 6.8–7.6 (7.4). Pronotum width: male 6.7–7.4 (7.1); female 6.5–8.4 (7.5). Abdomen width: male 7.4–8.0 (7.7); female 7.5–8.2 (7.8). Ovipositor length 7.6–8.4 (8.0).   Etymology. Since most specimens have been collected in cool temperate areas, the species name reflects the climatic situation where this species is typically found.  Distinguishing features.  Yoyettasubalpina sp. nov.displays concentric orange bands on the posterior margins of tergites 3–7; these are absent on  Y. denisoni, Y. kershawi stat. rev., comb. nov.,  Y. timothyi sp. nov.and  Y. grandis sp. nov.It is readily distinguished from  Y. abdominalis, Y. electrica sp. nov.,  Y. spectabilis sp. nov.and  Y. serrata sp. nov.by the following combination of characters: (1) large size with body length> 23 mm, (2) lack of symmetrical, orange triangular patterns on tergites 4–7, (3) ovipositors of females extend> 1 mmbeyond the posterior abdomen. The forewing basal membranes of  Y. regalis sp. nov.and  Y. subalpina sp. nov.are bright orange contrasting with the grey basal membranes of  Y.grandis sp. nov.in southern Victoriaand the dull orange basal membranes of specimens of  Y.grandis sp. nov.in the Grampians Ranges. Morphologically, male specimens of  Y. subalpina sp. nov.,  Y. verrens sp. nov.and  Y. regalis sp. nov.are difficult to differentiate, but the follow features can be used to achieve the correct identification. In  Y. subalpina sp. nov., the white colouration on the hind wing plaga is bold and extends to cover the entire jugum (cf. in  Y. verrens sp. nov.the colouration is dull white and restricted along the margins of anal cell 3 and vein 2A). In  Y. subalpina sp. nov., the sternites are predominantly orange, sometimes with diffuse black spots medially (cf. in  Y. regalis sp. nov.where the sternites have bold black medial markings that occupy more than half of each sternite). In female specimens, there is substantial colour variation morphologically (orangebrown to black) within species, but the ovipositors extend beyond the abdomen around 1.0 mm for  Y. subalpina sp. nov., 1–1.5 mmfor  Y. regalis sp. nov.,> 1.5 mmfor  Y. grandis sp. nov., and> 2 mmfor  Y. verrens sp. nov.The length of the ovipositor of  Y. regalis sp. nov.is> 8.8 mm, < 8.6 mmfor  Y. subalpina sp. nov., and> 10.0 mm for  Y. grandis sp. nov.  Calling song( Figs 29–31, 53G). The male calling song of  Y. subalpina sp. nov.is a simple repetition of phrases, each comprising an echeme followed by a macrosyllable ( Fig. 29A). Closer inspection of the echeme ( Fig. 29B) reveals that it comprises a series of macrosyllables that increase in amplitude and coalesce at the highest point of amplification, typically occurring between 0.16 and 0.43 sduration (n = 13). A short gap of 0.01– 0.04 sduration separates the echeme from the proceeding macrosyllable. The macrosyllable is typically louder than the echeme. It lasts between 0.05 and 0.09 s( Fig. 29C). Gaps of approximately 0.5 sduration separate each phrase. After warming up a little, males sometimes produce soft ticking (syllable trains) in the gaps between each phrase, with longer syllable trains between bouts of phrases ( Fig. 30). Males from most locations across the geographical distribution of this species had a frequency plateau of 8.4–11.1 kHz and a dominant frequency ranging between 9.4 and 10.1 kHz ( Fig. 31). The exceptions were males from Talleganda that produced a slightly lower frequency plateau, ranging from 7.1–9.7 kHz with a dominant frequency of 8.4–9.0 kHz (n = 4; Fig. 30). Within the  Y. abdominalisspecies group, the calling song of  Y. subalpina sp. nov.is most similar to  Y. verrens sp. nov.The main difference in the call of  Y. subalpina sp. nov.is the addition of a macrosyllable following the echeme, which gives it a distinct two-note signature (cf. the single echeme sweeping notes of  Y. verrens sp. nov.). 2010-12-10 AM L. Popple -36.346386 Cooma 19 148.2325 38 313 1 1 holotype 1978-12-26 D. C. F. Rentz & Database -35.35 Brindabella Ra. 1193 148.8 Piccadilly Circus 38 313 2 1 1 New South Wales paratype 1931-12-12 L. F. Graham & Database No. Mt Kosciusko Snowy River 38 313 1 1 New South Wales paratype 1967-01-24 T. G. Campbell & Database Old Adaminaby 38 313 1 1 New South Wales paratype 1969-02-19 M. G. Stanger & Eucalypt & Database Snowy Plains 38 313 3 3 New South Wales paratype Sellar, W. W & Froggatt & Database No. Cooma 38 313 1 1 New South Wales paratype 2014-12-25 D. J. & R. L. Ferguson & Database No. 1142 -35.714466 Round Flat 1 149.29459 Tinderry Mountains 38 313 1 1 New South Wales paratype 2014-12-21 ANIC D. J. Ferguson & Database No. 1160 -35.730324 Tinderry Rd 1 149.3022 Tinderry Mountains 38 313 1 1 New South Wales paratype 1931-01-18 2558 1931-01-18 HEM -36.45 Mt Kosciusko 1187 148.26666 38 313 6 4 2 New South Wales paratype 1953-01-17 MMV -36.416668 Jindabyne 1187 148.61667 38 313 HEM2559 1 1 New South Wales paratype 2009-12-29 L. W. Popple -36.604443 Nimmitabel 19 149.37527 Brown Mountain 38 313 7 3 4 New South Wales paratype 2010-12-10 LWP L. W. Popple 39 314 -36.346664 Tom Groggin Road 19 148.2325 38 313 1 1 New South Wales paratype Countegany on Kybean Rd 39 314 1 1 New South Wales paratype 1985-12-26 B. Underwood Mt David 39 314 9 8 1 New South Wales paratype 1973-12-27 Rd Cooma on Nimitabelle 39 314 3 2 1 New South Wales paratype 2011-01-17 Hill & D. Marshall & C. Simon -35.5543 Kosciuszko 1 148.51404 Yarrangobilly Caves 39 314 5 2 3 New South Wales paratype 2011-01-17 MSM Hill & D. Marshall & C. Simon -35.5543 Kosciuszko 1 148.51404 Yarrangobilly Caves 39 314 1 1 New South Wales paratype 2002-01-10 2002-01-14 2002-01-10 D. Emery -35.359444 Brindabella Ranges 19 148.79973 Piccadilly Circus 39 314 7 6 1 New South Wales paratype 2010-01-08 D. Emery & L. Popple -33.988335 Kanangra Boyd National Park 19 150.04028 39 314 2 2 New South Wales paratype [121,641,1651,1670] 1966-01-16 UQIC T. Weir Mt Buffalo Nat. Pk 39 314 1 1 Victoria paratype 1984-01-16 Key -36.483334 E of Nariel Vic. 1187 147.9 39 314 1 1 Victoria paratype 1983-12-29 ANIC K. H. L. Key & Database No. 1325 -36.883335 Vic. 1185 148.08333 Benambra 39 314 1 1 Victoria paratype 1933-12-19 AM Harrietville & St Bernard Hospice & Musgrave 39 314 6 3 3 Victoria paratype 2002-12-15 D. J. Hilton Sommerville Ave Vic Lorne 39 314 1 1 Victoria paratype S. Emery & T. Corbin -38.910835 Vic Otway Ranges 39 314 2 2 Victoria paratype 2016-12-17 S. Emery & T. Corbin -37.469723 Vic 19 145.33665 Kinglake 39 314 1 1 Victoria paratype 2017-01-11 S. Emery & T. Corbin -37.54222 Toolangi 19 145.47472 Country Fire Authority Station 39 314 2 2 Victoria paratype [176,646,1891,1910] 2017-01-23 Emery -37.54222 Toolangi 19 145.47472 Country Fire Authority Station 39 314 2 2 Victoria paratype 1976-01-02 M. S. & B. J. Moulds 1830 Mt Hotham Alpine Reserve 39 314 13 10 3 Victoria paratype [261,477,1939,1958] 1976-01-01 M. S. & B. J. Moulds 1830 Mt Hotham Alpine Reserve 39 314 4 4 Victoria paratype 1990-01-01 1990-01-13 1990-01-01 MSM G. R. Brown & M. A. Terras -36.983334 Mt Hotham Village 1184 147.18333 39 314 1 1 Victoria paratype 1927-12-04 HEM -37.533333 Vic 1181 145.33333 Kinglake 39 314 HEM2611 1 1 Victoria paratype [602,1255,1387,2006] 1951-12-29 2616 1951-12-29 HEM -36.716667 Vic 1185 146.76666 Mt Buffalo 39 314 5 5 Victoria paratype 2018-12-27 S. Emery & T. Corbin -37.54222 Toolangi 19 145.47472 Country Fire Authority Station 39 314 1 1 Victoria paratype 2019-01-14 MMV Creen Victoria 39 314 HEM5765-7 2 1 1 Victoria paratype 1978-12-26 ANIC Piccadilly Circus & Brindabella Ra. A. C. T. & D. C. F. Rentz -35.35 AUSTRALIAN CAPITAL TERRITORY 1193 148.8 39 314 1 1 Australian Capital Territory paratype 2004-12-05 B. Kenway -35.533333 Brindabella Ranges 1193 148.76666 Mt Franklin Rd 39 314 1 1 Australian Capital Territory paratype 2009-12-28 L. W. Popple -35.360275 Brindabella Range 19 148.79973 Piccadilly Circus 39 314 1 1 Australian Capital Territory paratype 2011-01-15 L. W. Popple -35.3775 Bull's Head 19 148.80444 39 314 2 1 1 Australian Capital Territory paratype 2011-01-15 LWP L. W. Popple -35.489723 Mt Franklin area 19 148.76334 39 314 1 1 Australian Capital Territory paratype 2009-12-05 L. W. Popple -35.62 Talleganda National Park 19 149.49834 Rocky Pic Road 39 314 1 New South Wales 2017-12-15 T. Corbin 600 -37.824444 Mt Dandenong 19 145.35945 Ridge Rd 39 314 1 Victoria