A Revision of the Yoyetta abdominalis (Distant) Species Group of Cicadas (Hemiptera: Cicadidae: Cicadettinae), Introducing Eight New Species
Emery, David L.
Emery, Nathan J.
Popple, Lindsay W.
Records of the Australian Museum
2019
2019-10-02
71
7
277
347
8K2CP
Emery & Emery & Popple, 2019
Emery & Emery & Popple
2019
[963,1221,1423,1454]
Insecta
Cicadidae
Yoyetta
Animalia
Hemiptera
38
313
Arthropoda
species
subalpina
sp. nov.
Figs 2E, 3E, 26– 31, 52E, 53G
Holotype ♂, 18 kmE Cooma, NSW, 36°20'47"S 148°13'57"E, 10.xii.2010, L. Popple, 514-0013 ( AMK.559471). Paratypes NEW SOUTH WALES: 1♂ 1♀, Piccadilly Circus, Brindabella Ra., ACT, 35°21'S 148°48'E, 26.xii.1978, D. C. F. Rentz, ANIC Databaseno. 20 010822; 1♀, Snowy River, Mt Kosciusko, 4000FT, 12.xii.1931, L. F. Graham, ANIC Database No.20 010821; 1♂, Old AdaminabyNSW, 24.i.1967, T. G. Campbell, ANIC Databaseno. 20 010820; 2♀♀, NSW Snowy Plains, 19.ii.1969, M. G. Stanger, on Eucalypt, [ 1♀] ANIC Databaseno. 20 010823; 1♀, CoomaNSW, 10.i.[19]06, Sellar, W. W. Froggattcollection, ANIC Database No.20 010815; 1♂, Tinderry Mountains, Round Flat, netted, 1142 m, 25.xii.2014, 35°42'52.08"S 149°17'40.48"E, D. J. & R. L. Ferguson, ANIC Database No.20 010718; 1♂, Tinderry Mountains, Tinderry Rdverge, netted, 1160 m, 35°43'49.17"S 149°18'07.92"E, 21.xii.2014, D.J. Ferguson, ANIC Database No.20 010816 ( ANIC); 2♂♂ 4♀♀, Mt Kosciusko, NSW, 36°27'S 148°16'E, 18.i.1931( HEM 2554–58); 1♀, JindabyneNSW, 36°25'S 148°37'E, 17.i.1953, ( HEM2559) ( MMV); 4♂♂, 3♀♀, 1 kmS. of Brown Mountain, via Nimmitabel, 36°36'16"S 149°22'31"E, 29.xii.2009, L. W. Popple, 514-0003 to 514-0009; 1♂, Tom Groggin Road, 18 kmESE of Cooma, 36°20'48"S 148°13'57"E, 10.xii.2010, L. W. Popple, 514-0014 ( LWP); 1♀, 7 miles[SW of?] Countegany on Kybean Rd, 16.xii.[19]62, L. R. G[reenup]?, 105; 1♂ 8♀♀, Mt David, 30 kmSW of Oberon, 26.xii.1985, S. &. B. Underwood; 1♂ 2♀♀, 12 mi.from Cooma on Nimitabelle[sic.] Rd, 27.xii.1973, Euc.pauciflora, L. R. G[reenup]; 3♂♂ 2♀♀, Yarrangobilly Cavesroad, KosciuszkoNP, 35°33.258'S 148°30.842'E, 17.i.2011, 1305 m [elev.], K. Hill, D. Marshall, AU.NS.YAX, C. Simonlab voucher, Yoyetta“double-zit”; 1♂, same data; legs in ETOH, body pinned, 11.AU.NS.YAX.01 ( MSM); 1♂ 6♀♀, Piccadilly Circus, Brindabella Ranges, NSW, 35°21'34"S 148°47'59"E, 10–14.i.2002, D.Emery; 2♀♀, Kanangra Boyd National Park, NSW, 33°59'18"S 150°02'25"E, 8.i.2010, D. Emery& L. Popple(DE). VICTORIA: 1♂ Mt Buffalo Nat. Pk, 16.i.1966, T.Weir( UQIC); 1♀, 36°29'S 147°54'E, 8 kmSE by E of Nariel Vic., 16.i.1984, K. H.L. Key; 1♀, 9 kmE of Mt Pendergast, E of Benambra, Vic., 1325m, 36°53'S 148°05'E, 29.xii.1983, K. H. L. Key, ANIC Database No.20 010819 ( ANIC); 3♂♂ 3♀♀, midway between Harrietville& St Bernard Hospice, 19 Dec 1933, A. Musgrave( AMK.307236–41); 1♂, Sommerville Ave, Lorne, Vic, 15.xii.2002, D. J. Hilton; 2♀♀, 1 kmSE Forrest, Otway Ranges, Vic, 38°54'39"S143°74'40"E, 5.ii.15, S. Emery& T. Corbin; 1♂, Kinglake, Vic, 37°28'11"S 145°20'12"E, 17.xii.2016, S. Emery& T. Corbin; 2♂♂, Country Fire Authority Station, Toolangi, 37°32'32"S 145°28'29"E, 11.i.2017, S. Emery& T. Corbin(to light); 2♂♂, same location, 23.i.2017, N., C. & D. Emery(DE); 2♂♂ 9♀♀, Mt Hotham Alpine Reserve, 1,830 m[elev.], 2.i.1976, M. S. & B. J. Moulds( 1♂ 1♀in cop); 4♂♂, same data, 1.i.1976, 1♂in cop; Mt Hotham Village, 36°59'S 147°11'E, 1–13.i.1990, G. R. Brown& M. A. Terras( MSM); 1♂, Kinglake, Vic, 37°32'S 145°20'E, 4.xii.1927, ( HEM2611); 5♀♀, Mt Buffalo, Vic, 36°43'S 146°46'E, 29.xii.1951( HEM 2612–16); 1♂, Country Fire Authority Station, Toolangi, 37°32'32"S 145°28'29"E, 27.xii.2018, S. Emery& T. Corbin(to light; T 22389); 5♂♂ 3♀♀, Allambee, Victoria, 14.i.2019, J. Creen( HEM5765–7[ ♀], HEM5768–72 [ ♂]) ( MMV). AUSTRALIAN CAPITAL TERRITORY. 1♀, 35°21S 148°48E, Piccadilly Circus, Brindabella Ra. A.C.T., 26.xii.1978, D. C. F. Rentz( ANIC); 1♂, Mt Franklin Rd, Brindabella Ranges, 35°32'S 148°46'E, 5.xii.2004, B.Kenway,514-0001; 1♂, Piccadilly Circus, Brindabella Range, 35°21'37"S 148°47'59"E, 28.xii.2009, L. W. Popple, 514-0002, LT2009371; 1♂ 1♀, 1.5 kmN. of Bull’s Head, 35°22'39"S 148°48'16"E, 15.i.2011, L. W. Popple, 514-0015, 514-0017, LT2009371; 1♀, Mt Franklin area, 35°29'23"S 148°45'48"E, 15.i.2011, L. W. Popple, 514-0018, LT2009371 ( LWP). Figure 26. Map of eastern Australia showing the geographical distribution of Yoyetta subalpina sp. nov.(open triangles), Y. serrata sp. nov.(closed squares) and Y. grandis sp. nov.(open circles). Additional records from song recordings NEW SOUTH WALES: Rocky Pic Road, Talleganda National Park, NSW, 35°37'12"S 149°29'54"E, 5.xii.2009, L. W. Popple. VICTORIA: Ridge Rd, Mt Dandenong, 600 m, 37°49'28"S 145°21'34"E, 15.xii.2017, T. Corbin.
Distribution, habitat and seasonality. Cool temperate alpine areas from Kanangra-Boyd National Park in central New South Walessouth to the Upper Yarra region in central Victoria, with an isolated population in the Otway Ranges ( Fig. 26). Adults occur mainly on smooth-barked eucalypts on the main trunks and upper branches. They can be encountered from December to February.
Description Male( Figs 2E, 3E, 27A– B, 27F, 51G, 52E). Headas wide as mesonotum, black, with an ochraceous diamond-shaped marking on midline, posterior to ocelli; ocelli pink to pale; dorsal postclypeus black with central ochraceous triangular marking, apex directed anteriorly; ventral postclypeus black with anterior brown marking on midline, and ochraceous, lateral and posterior margins; anteclypeus black, rostrum brown at base, becoming black at apex, reaching the middle of hind coxae; lora black with ochraceous anterior margin, gena black; eyes dull black; antennae black, supra-antennal plates black, ochraceous around base of pedicel. Thoraxpredominantly black. Pronotum black, some specimens with brown in fissures; pronotal collar black, posterior margin ochraceous in some specimens, margins of lateral angles light brown, paranota black, anterior angles ochraceous in some specimens. Mesonotum black; cruciform elevation arms black, lateral depressions tinged brown. Metanotum black. Legs.Coxae mainly black, fore coxae black with proximal ochraceous triangular stripe on posterior surface; mid and hind coxae black; coxal joint and membranes red, basisterna black; trochanters black with central segment orange-red; meracantha small, narrow, orange, black at base, pointed, minimally overlapping opercula; fore femora striped longitudinally orange-red and black, black along base of femoral spines femoral spines erect, black at base tending dark brown at tips; mid and hind femora black, orange-brown longitudinal Figure 28. Yoyetta subalpina sp. nov., illustration of male pygofer and internal genitalia; (a) viewed laterally from the left; (b)viewed ventrally; (c)aedeagus; and (d)apex of theca. Characters as depicted in Fig. 4. Specimen from Piccadilly Circus ( 35°22'S 148°48'E). stripe on medial aspect, distal femoral joint orange-red; fore tibiae black, brown at distal tip; mid tibiae black, with medial ochraceous stripe; hind tibiae black proximally ochraceous over distal half; fore tarsi black with brown distal margins; mid and hind tarsi blackish-brown becoming black towards claws; claws dark brown, black at tips. Wingswith fore wing costal veins black, browner central rib, becoming ochraceous distally; pterostigma mottled red; basal cell translucent with black anterior border; basal membranes bright orange; other venation dark brown to black; with eight apical cells; hind wing plagas white over entire anal cell 3 and medial third of ac2(v), thin along vein 2A, an irregular grey-black marking at proximal end of jugum, reddish stippling at base of ac2(v), veins 2A and 3A pale, remainder dark brown to black, with six apical cells. Opercula( Fig. 3E) medium, spatulate, following body axis ventrolaterally, depressed centrally, variably black over basal half, orange-brown across remainder, clearly separated. Timbals( Fig. 2E) with five distinct long ribs; long ribs 1–4 extending across surrounding membrane and fused dorsally along basal spur; long rib 5 independent of basal spur, comparatively shorter, extending ventrally across half of membrane; large ridged dome on posterior timbal plate extending across three-quarters of timbal; apodeme pit ovalshaped and conspicuous. Abdomen. Tergite 1 black, with orange-red membranes around timbal cavity; tergite 2 black; tergites 3–7 black with orange posterior margins, extending laterally to epipleurites, increasing on tergites 6–7 on either side of midline; tergite 8 shiny black, with ventral margins orange. Epipleurites with medial black strip, orange over reminder. Sternite II mainly black, with central orange spot; sternite III black medially, becoming orange laterally along posterior margin; sternites IV–VI orange, posterior halves becoming progressively reddish, diffuse black midline markings over anterior twothirds of each sternite, decreasing posteriorly, sternite VIII dull orange-red, with black pubescence. Figure 29. Typical male calling song structure of Yoyetta subalpina sp. nov., illustrated in waveform plots, including (A)two phrases; (B)expanded section from A showing a single phrase with an echeme followed by a long macrosyllable; (C)expanded section from B showing more detail from the echeme and the macrosyllable. The final subfigure (D)is a spectrogram displaying song frequency. The specimen was recorded in the field at Piccadilly Circus (35°22'S 148°48'E) by LWP using RS2 (see Methods and terminology). Figure 30. Comparative male calling song structure of Yoyetta subalpina sp. nov., illustrated in waveform plots, including (A)a 20 s section containing echemes, interspersed with a “soft ticking” segment; (B)expanded “soft ticking” section (from A) showing 10 syllables.The final subfigure (C)is a spectrogram displaying song frequency. The specimen was recorded in the field at Talleganda (35°37'S 149°30'E) by LWP using RS2 (see Methods and terminology). Genitalia( Fig. 28). Pygofer black, upper lobe mainly black, apex brown, basal lobe black; dorsal beak black, dark brown posterior margin, anal styles orange-red. Uncus reddishbrown; in lateral view beak-like and elongated; lobes in ventral view narrow, with rounded lateral termination; claspers clearly divided, short, bulbous, with apices tapering laterally.Aedeagus with pseudoparameres almost extending as far as theca; theca recurved ventrally at around 180° towards apex, with prominent transparent flange along distal margin of recurvature, smooth and broadening to> 3× width of theca; remainder of theca is short, with <20% of theca extending apically beyond margin of recurvature; apex blunt, with 2–3 cornuti on ventral surface and several small cornuti on dorsal side at tip. Female( Fig. 27C–E). Head, thoraxand abdomenof dark specimens similar to male. Paler, brown-coloured females apparently occur at a ratio of approximately 1:2. Headin dark specimens, similar to male; in paler specimens mainly brown with black markings around ocelli extending to the supra-antennal plate, brown at anterior margin, black spot at medial border of eye; dorsal postclypeus ochraceous centrally, dark brown laterally; ventral postclypeus black with central ochraceous spot anteriorly; anteclypeus and rostrum similar to male; gena variably ochraceous and black, lateral margins ochraceous posteriorly, black anteriorly. Thoraxin dark specimens, similar to male. In paler specimens; pronotum with black markings restricted to lateral and posterior margins of a broad ochraceous midline strip over central third, and variably along fissures and lateral margins; pronotal collar brown, with lateral angles black; mesonotum brown with black submedian and lateral sigilla, and a diffuse black central triangular marking with base around scutal depressions, apex directed anteriorly on midline to posterior third of submedian sigilla; cruciform elevation black on central, medial portion and lateral margins, yellow-ochraceous on posterior margins, lateral depressions ochraceous, anterior and posterior depressions dark brown to black; scutal depressions black; metanotum black. Abdomen.Colour variation similar to thorax. On dark specimens, similar to male; abdominal segment 9 black dorsally with ochraceous triangular markings either side of midline, tapering posteriorly, ending midway to dorsal beak, becoming orange laterally, reddish on ventral margin. On brown specimens; tergite 1 black, tergite 2 black on midline, orange merging to black laterally; tergites 3–8 black with orange posterior margins, increasing posteriorly, extending laterally to epipleurites; epipleurites with medial black strip, orange over remainder; sternite II black; sternites III–VI pale brown with midline black marking; sternite VII pale brown with anterior black spot either side of midline; abdominal segment 9 orange-brown with three longitudinal black stripes faint on midline and prominent either side of midline. On all specimens, dorsal beak orange to black at base, black at tip; ovipositor dark brown, becoming black at tip, extending < 1 mmbeyond apex of abdominal segment 9. Anal styles orange-red; ovipositor sheath dark brown to black. Measurements(in mm; range with mean in parentheses: 10 males, 8 females). Body length: male 25.0–25.1 (25.1); female 24.0–27.0 (26.0). Fore wing length: male 27.7– 28.5 (28.1); female 29.4–33.2 (31.9). Head width: male 7.0–7.6 (7.3); female 6.8–7.6 (7.4). Pronotum width: male 6.7–7.4 (7.1); female 6.5–8.4 (7.5). Abdomen width: male 7.4–8.0 (7.7); female 7.5–8.2 (7.8). Ovipositor length 7.6–8.4 (8.0).
Etymology. Since most specimens have been collected in cool temperate areas, the species name reflects the climatic situation where this species is typically found. Distinguishing features. Yoyettasubalpina sp. nov.displays concentric orange bands on the posterior margins of tergites 3–7; these are absent on Y. denisoni, Y. kershawi stat. rev., comb. nov., Y. timothyi sp. nov.and Y. grandis sp. nov.It is readily distinguished from Y. abdominalis, Y. electrica sp. nov., Y. spectabilis sp. nov.and Y. serrata sp. nov.by the following combination of characters: (1) large size with body length> 23 mm, (2) lack of symmetrical, orange triangular patterns on tergites 4–7, (3) ovipositors of females extend> 1 mmbeyond the posterior abdomen. The forewing basal membranes of Y. regalis sp. nov.and Y. subalpina sp. nov.are bright orange contrasting with the grey basal membranes of Y.grandis sp. nov.in southern Victoriaand the dull orange basal membranes of specimens of Y.grandis sp. nov.in the Grampians Ranges. Morphologically, male specimens of Y. subalpina sp. nov., Y. verrens sp. nov.and Y. regalis sp. nov.are difficult to differentiate, but the follow features can be used to achieve the correct identification. In Y. subalpina sp. nov., the white colouration on the hind wing plaga is bold and extends to cover the entire jugum (cf. in Y. verrens sp. nov.the colouration is dull white and restricted along the margins of anal cell 3 and vein 2A). In Y. subalpina sp. nov., the sternites are predominantly orange, sometimes with diffuse black spots medially (cf. in Y. regalis sp. nov.where the sternites have bold black medial markings that occupy more than half of each sternite). In female specimens, there is substantial colour variation morphologically (orangebrown to black) within species, but the ovipositors extend beyond the abdomen around 1.0 mm for Y. subalpina sp. nov., 1–1.5 mmfor Y. regalis sp. nov.,> 1.5 mmfor Y. grandis sp. nov., and> 2 mmfor Y. verrens sp. nov.The length of the ovipositor of Y. regalis sp. nov.is> 8.8 mm, < 8.6 mmfor Y. subalpina sp. nov., and> 10.0 mm for Y. grandis sp. nov. Calling song( Figs 29–31, 53G). The male calling song of Y. subalpina sp. nov.is a simple repetition of phrases, each comprising an echeme followed by a macrosyllable ( Fig. 29A). Closer inspection of the echeme ( Fig. 29B) reveals that it comprises a series of macrosyllables that increase in amplitude and coalesce at the highest point of amplification, typically occurring between 0.16 and 0.43 sduration (n = 13). A short gap of 0.01– 0.04 sduration separates the echeme from the proceeding macrosyllable. The macrosyllable is typically louder than the echeme. It lasts between 0.05 and 0.09 s( Fig. 29C). Gaps of approximately 0.5 sduration separate each phrase. After warming up a little, males sometimes produce soft ticking (syllable trains) in the gaps between each phrase, with longer syllable trains between bouts of phrases ( Fig. 30). Males from most locations across the geographical distribution of this species had a frequency plateau of 8.4–11.1 kHz and a dominant frequency ranging between 9.4 and 10.1 kHz ( Fig. 31). The exceptions were males from Talleganda that produced a slightly lower frequency plateau, ranging from 7.1–9.7 kHz with a dominant frequency of 8.4–9.0 kHz (n = 4; Fig. 30). Within the Y. abdominalisspecies group, the calling song of Y. subalpina sp. nov.is most similar to Y. verrens sp. nov.The main difference in the call of Y. subalpina sp. nov.is the addition of a macrosyllable following the echeme, which gives it a distinct two-note signature (cf. the single echeme sweeping notes of Y. verrens sp. nov.).
2010-12-10
AM
L. Popple
-36.346386
Cooma
19
148.2325
38
313
1
1
holotype
1978-12-26
D. C. F. Rentz & Database
-35.35
Brindabella Ra.
1193
148.8
Piccadilly Circus
38
313
2
1
1
New South Wales
paratype
1931-12-12
L. F. Graham & Database No.
Mt Kosciusko
Snowy River
38
313
1
1
New South Wales
paratype
1967-01-24
T. G. Campbell & Database
Old Adaminaby
38
313
1
1
New South Wales
paratype
1969-02-19
M. G. Stanger & Eucalypt & Database
Snowy Plains
38
313
3
3
New South Wales
paratype
Sellar, W. W & Froggatt & Database No.
Cooma
38
313
1
1
New South Wales
paratype
2014-12-25
D. J. & R. L. Ferguson & Database No.
1142
-35.714466
Round Flat
1
149.29459
Tinderry Mountains
38
313
1
1
New South Wales
paratype
2014-12-21
ANIC
D. J. Ferguson & Database No.
1160
-35.730324
Tinderry Rd
1
149.3022
Tinderry Mountains
38
313
1
1
New South Wales
paratype
1931-01-18
2558
1931-01-18
HEM
-36.45
Mt Kosciusko
1187
148.26666
38
313
6
4
2
New South Wales
paratype
1953-01-17
MMV
-36.416668
Jindabyne
1187
148.61667
38
313
HEM2559
1
1
New South Wales
paratype
2009-12-29
L. W. Popple
-36.604443
Nimmitabel
19
149.37527
Brown Mountain
38
313
7
3
4
New South Wales
paratype
2010-12-10
LWP
L. W. Popple
39
314
-36.346664
Tom Groggin Road
19
148.2325
38
313
1
1
New South Wales
paratype
Countegany on Kybean Rd
39
314
1
1
New South Wales
paratype
1985-12-26
B. Underwood
Mt David
39
314
9
8
1
New South Wales
paratype
1973-12-27
Rd
Cooma on Nimitabelle
39
314
3
2
1
New South Wales
paratype
2011-01-17
Hill & D. Marshall & C. Simon
-35.5543
Kosciuszko
1
148.51404
Yarrangobilly Caves
39
314
5
2
3
New South Wales
paratype
2011-01-17
MSM
Hill & D. Marshall & C. Simon
-35.5543
Kosciuszko
1
148.51404
Yarrangobilly Caves
39
314
1
1
New South Wales
paratype
2002-01-10
2002-01-14
2002-01-10
D. Emery
-35.359444
Brindabella Ranges
19
148.79973
Piccadilly Circus
39
314
7
6
1
New South Wales
paratype
2010-01-08
D. Emery & L. Popple
-33.988335
Kanangra Boyd National Park
19
150.04028
39
314
2
2
New South Wales
paratype
[121,641,1651,1670]
1966-01-16
UQIC
T. Weir
Mt Buffalo Nat. Pk
39
314
1
1
Victoria
paratype
1984-01-16
Key
-36.483334
E of Nariel Vic.
1187
147.9
39
314
1
1
Victoria
paratype
1983-12-29
ANIC
K. H. L. Key & Database No.
1325
-36.883335
Vic.
1185
148.08333
Benambra
39
314
1
1
Victoria
paratype
1933-12-19
AM
Harrietville & St Bernard Hospice & Musgrave
39
314
6
3
3
Victoria
paratype
2002-12-15
D. J. Hilton
Sommerville Ave
Vic
Lorne
39
314
1
1
Victoria
paratype
S. Emery & T. Corbin
-38.910835
Vic
Otway Ranges
39
314
2
2
Victoria
paratype
2016-12-17
S. Emery & T. Corbin
-37.469723
Vic
19
145.33665
Kinglake
39
314
1
1
Victoria
paratype
2017-01-11
S. Emery & T. Corbin
-37.54222
Toolangi
19
145.47472
Country Fire Authority Station
39
314
2
2
Victoria
paratype
[176,646,1891,1910]
2017-01-23
Emery
-37.54222
Toolangi
19
145.47472
Country Fire Authority Station
39
314
2
2
Victoria
paratype
1976-01-02
M. S. & B. J. Moulds
1830
Mt Hotham Alpine Reserve
39
314
13
10
3
Victoria
paratype
[261,477,1939,1958]
1976-01-01
M. S. & B. J. Moulds
1830
Mt Hotham Alpine Reserve
39
314
4
4
Victoria
paratype
1990-01-01
1990-01-13
1990-01-01
MSM
G. R. Brown & M. A. Terras
-36.983334
Mt Hotham Village
1184
147.18333
39
314
1
1
Victoria
paratype
1927-12-04
HEM
-37.533333
Vic
1181
145.33333
Kinglake
39
314
HEM2611
1
1
Victoria
paratype
[602,1255,1387,2006]
1951-12-29
2616
1951-12-29
HEM
-36.716667
Vic
1185
146.76666
Mt Buffalo
39
314
5
5
Victoria
paratype
2018-12-27
S. Emery & T. Corbin
-37.54222
Toolangi
19
145.47472
Country Fire Authority Station
39
314
1
1
Victoria
paratype
2019-01-14
MMV
Creen
Victoria
39
314
HEM5765-7
2
1
1
Victoria
paratype
1978-12-26
ANIC
Piccadilly Circus & Brindabella Ra. A. C. T. & D. C. F. Rentz
-35.35
AUSTRALIAN CAPITAL TERRITORY
1193
148.8
39
314
1
1
Australian Capital Territory
paratype
2004-12-05
B. Kenway
-35.533333
Brindabella Ranges
1193
148.76666
Mt Franklin Rd
39
314
1
1
Australian Capital Territory
paratype
2009-12-28
L. W. Popple
-35.360275
Brindabella Range
19
148.79973
Piccadilly Circus
39
314
1
1
Australian Capital Territory
paratype
2011-01-15
L. W. Popple
-35.3775
Bull's Head
19
148.80444
39
314
2
1
1
Australian Capital Territory
paratype
2011-01-15
LWP
L. W. Popple
-35.489723
Mt Franklin area
19
148.76334
39
314
1
1
Australian Capital Territory
paratype
2009-12-05
L. W. Popple
-35.62
Talleganda National Park
19
149.49834
Rocky Pic Road
39
314
1
New South Wales
2017-12-15
T. Corbin
600
-37.824444
Mt Dandenong
19
145.35945
Ridge Rd
39
314
1
Victoria