Lispa uniseta Malloch, 1922: 389 Lispa uniseta Malloch, 1923 : 609 Malloch, 1925a : 335 Malloch, 1925b : 340 Lee, Crust & Sabrosky, 1956 : 338 Lispe uniseta Séguy, 1937 : 186 Pont, 1989 : 693 Studies on the Australian Muscidae (Diptera). VIII. The genus Lispe Latreille, 1797 Pont, Adrian C. Zootaxa 2019 2019-02-18 4557 1 1 232 Malloch Malloch [151,420,620,646] Insecta Anthomyiidae Lispe Animalia Diptera 191 192 Arthropoda species uniseta    Lispa unisetaMalloch, 1922: 389.    Lispa uniseta.—  Malloch, 1923: 609;  Malloch, 1925a: 335;  Malloch, 1925b: 340;  Lee, Crust & Sabrosky, 1956: 338.    Lispe uniseta.—  Séguy, 1937: 186;  Pont, 1989: 693.  Type-material.The holotype ♀, from Port Melbourne (V), is in BMNH and was examined in the course of this revision.   Diagnosis.This species and  L. vikhrevidiffer from other Australian  Lispeexcept  L. setigenaby the presence of a pdseta on hind tibia and an adon mid tibia; from  L. setigenathey differ by having only 1+2 dcsetae. Males of  L. unisetaand  L. vikhreviare easily separated by the characters given in the key (p. 17); most strikingly,  L. unisetahas the mid tarsomeres with simple shape and setation (cf.  L. vikhrevi, Fig. 406) and has a pvand pseta placed together at middle of fore tibia.   Material Examined.AUSTRALIA:  Northern Territory: 12 kmSW of Alroy Downs, at light, 10.iv.1976, D.H. Colless, 1♀( ANIC);  13 kmE by N of AndadoHS, Simpson Desert,  27–28.ix.1972, Z. Liepa, 1♂( ANIC).   Queensland: near Birdsville Racetrack,  1.ix.1997, J. & A. Skevington, C. Lambkin, S. Winterton, 1♀( QMBA);  Brisbane,  21.vii.1963, C. Speed, 1♀( QMBA);  Dynevor Lakes,  88 kmW Eulo,  150 m, 28.05S, 144.12E, G. Daniels,  27.ix.1991, 1♀( QMBA);  Roma,  15.vi.1976, J. Donaldson, 5♀( QDPC);  Townsville, on windows,  3.i.1970, P. Ferrar, 1♀( ANIC).   New South Wales: Argalong, Buccleugh, T. Vaughan-Sherrin, 1♀( USNM);  Billabong Creek, Conargo,  23–30.iv.1978, Z. Liepa, 4♀( ANIC& BMNH);  Billabong& Forest CreekJunction,  42 kmNE of Deniliquin,  2.v.1978, Z. Liepa, 1♀( ANIC);  Bronte, near Sydney,  17.vi.1956, D.K. McAlpine, 1♀( AMS);  23 kmSE by S of Byrock, at light,  5.iv.1976, D.H. Colless, 1♀( ANIC);   25 kmNW of Coonamble,  17.vi.1976, Z. Liepa, 2♂ 1♀( ANIC);   8 kmNE of Cudal,  25.xii.1976, G. Daniels, 1♀( AMS);   2.4 kmNE of Eugowra,  24.iv.1971, Z. Liepa, 2♀( ANIC);   24 kmN of Forbes,  29.xii.1976, Z. Liepa, 1♂( ANIC);  Gilmore Creek, Tumut,  23.xii.1973, Z. Liepa, 1♂( ANIC);  Goonoo Forest,  5 milesS Mendooran, m. v. lamp,  27.iii.1971, G. & A. Daniels, 3♀( QMBA);  Griffith, marshy creek,  15.xi.1956, W.W. Wirth, 1♀( USNM);  Gunnedah,  1.v.1925, 1♂( USNM);  Lachlan River,  35 kmW of Forbes,  26.xii.1976, Z. Liepa, 2♂( ANIC);  Lake Albert, near Wagga,  10.iv.1971, Z. Liepa, 1♀( ANIC);  Lake Cullulleraine,  36 milesW of Mildura,  14.i.1964, Z. Liepa, 4♂ 5♀( ANIC& BMNH);  Meroo Meadow,  15.xii.1926, B. Bertram, 1♀( AMS);  Merrylands,  25.xi.1964, M.I. Nikitin, 1♀( BMNH);  Murray River  50 milesW of Wentworth,  22.xi.1967, A. Neboiss, 1♀( MV);  Narrandera, 24 &  25.iii.1925, 1♂ 2♀( USNM) & 1♂( AMS);  Nowra Creek,  8.xii.1926, B. Bertram, 1♂( AMS);  Nyngan,  30.v.1984, K.C. Khoo, 1♂( AMS);   45 kmSE of Nyngan,  22.ix.1975, Z. Liepa, 1♀( ANIC);  Packsaddle,  111 milesN of Broken Hill,  24.xi.1969, R.W. Matthews, 1♀( ANIC);  Salt Caves Dam, PilligaSF,  9.x.1988, D.K. McAlpine, 1♀( AMS); same locality, m. v. lamp,  8–9.x.1988, R. de Keyzer& K.C. Khoo, 1♀( AMS);  Tumut River,  12.8 kmSE of Gundagai,  24.xii.1973, Z. Liepa, 2♂( ANIC);   40 kmN of Walgett,  19.vi.1976, Z. Liepa, 1♀( ANIC);  Wallaroi Creek,  15 kmS of Condobolin,  27.xii.1976, Z. Liepa, 1♂ 1♀( ANIC).   Australian Capital Territory: Black Mtn, light trap,  19.i.1968, 3.ii.1969, 4.ii.1964, 21.ii.1969, 14.xi.1968, 25.xi.1959, 2 &  10.xii.1968, 13.xii.1966, 13.xii.1968, I.F.B. Common, 4♂, 6♀( ANIC);  Lake Burley, Griffin,  31.x.1963, D.H. Colless, 1♂ 2♀( ANIC);   2 milesE of Mt Coree,  19.xi.1968, D.H. Colless, 1♀( ANIC).   Victoria: Clarkefield,  18.xii.1953, A. Neboiss, 1♀( MV);  Lake Cullulleraine, W of Mildura,  26.xii.1966, Z. Liepa, 3♂ 5♀( ANIC& BMNH);   47 kmNW of Hopetoun,  6.ix.1974, Z. Liepa, 1♀( ANIC);  Ovens River,  14 milesE of Yarrawonga,  24.xii.1966, Z. Liepa, 1♀( ANIC);  Port Melbourne,  10.xii.1906, W. Wesché, holotype ♀( BMNH);  Tatyoon,  23.i.1973, E.A. Fonseca, 7♂ 5♀( BMNH);  Wyperfield Nat. Park,  51 kmNW of Hopetoun,  4–7.ix.1974, Z. Liepa, 2♂( ANIC).   South Australia: Koonchera Dunenear Koonchera Waterhole, 26.41S, 139.30E,  2.ix.1997, J. & A. Skevington, 1♂ 1♀( QMBA);  Pandiburra Bore, 26.45S, 139.24E,  2.ix.1997, C. Lambkin, 2♂ 2♀( QMBA).   Description ♂ ♀. Head(Fig. 394). Ground-colour black. Frons at broadest point 0.39–0.40, at lunula 0.31 ( ♂) or 0.33 ( ♀), of maximum head-width (frontal view). Fronto-orbital plate white to yellowish-white pruinose, brownish above, and subshining at vertex, and also subshining around ocellar tubercle; parafacial and face white to yellowish-white pruinose; gena and lower half of occiput silvery pruinose, occiput with a large brown patch in upper half. Frontal triangle narrow, yellowish-white. Parafacial narrow, half ( ♂) or slightly less than ( ♀) width of postpedicel; setulose along whole length. Antenna black, except tip of pedicel and base of postpedicel which are obscurely orange; postpedicel 2.75 times as long as broad, falling short of mouth-margin by at most 0.35 its length. Arista plumose, the hairing at widest point equal to width of postpedicel. Vibrissa strong. Gena moderate, 0.18 ( ♂) to 0.15 ( ♀) of vertical eye-length; genal setae strong, sparse. Palpus yellow, greatly swollen in apical half.  Thorax. Ground-colour black. Scutum mainly shining black and undusted outside the dcsetae; between the dcsetae rather densely yellowish-grey dusted, this running from neck to tip of scutellum, sometimes extending somewhat beyond the line of the dcespecially in ♀♀, and superimposed on this with a median dark brown vitta running through the acrrows, also from neck to tip of scutellum. Postpronotal lobe and pleura grey dusted. Anterior spiracle dark brown. Acrsetulae in 4–5 irregular rows at suture. Dc1+2, prstseta midway between suture and neck or closer to neck; the 2 postsetae strong, anterior one closer to suture than to posterior one. 2 pprn. 1+0 (– 1) ia, postseta absent or hair-like, the prstseta hair-like in ♀. 1+1 sa, both strong. 1 strong and 1 weaker proepisternal setae. 1 strong proepimeral, equal to the strong proepisternal, with 3–4 adjacent setulae. Anepisternum with 1 short setula in upper anterior corner; posterior row with 3 strong and 3 weak setae. 3 strong kepst, lower one equidistant from upper two. Meron bare. Scutellum with disc densely setulose, with a few setulae below level of the strong setae; ventral surface bare.  Legs. Black, extreme tips of tibiae orange. Fore coxa of ♂with a long, flattened, drum-stick seta at tip of posterior edge (Fig. 395). Fore femur bare on avsurface; pvrow complete, the setae mostly very long in ♂; pdrow complete. Fore tibia with 1 submedian pvseta, and just basad of it in ♂with an additional strong pseta, this sometimes present in ♀too; with d, pand pvapical setae, and also with a short ( ♂) or long ( ♀) avapical. Fore tarsomeres without modifications. Mid femur bare ventrally, but the setulae slightly elongated, especially the pvnear base; 1 aat middle; 2 ppreapicals. Mid tibia with 1 submedian pand apicad of it with 1 ad, both strong; with strong d, a, v, pv, and weaker pdapicals. Mid tarsomeres without modifications. Hind coxa without a seta on posterior apical margin. Hind femur on pvsurface usually with 2 setae in basal half, subequal to femoral depth, and in ♂with a short preapical seta; avsurface setulose, in ♂with 2–3 setulae developed as short setae but much shorter than femoral depth; adrow complete; 1 pdpreapical. Hind tibia with 1 ad, 1 avand 1 pd; dand adpreapical setae present, the adshifted to ain ♂; 1 ( ♂) or 2 ( ♀) avbut no pvapicals. Hind tarsomeres not modified.  Wing. Clear. Tegula black, basicosta orange. Crossvein r-m placed slightly beyond the point where vein R1 enters costa; dm-cu weakly sinuous, longer than apical section of vein CuA1. Vein M running straight to wingmargin. Calypters white, with creamy to yellow margins. Knob of haltere dark brown.  Abdomen. Ground-colour black. Dust on tergites light grey: syntergite 1+2 mostly dusted; tergites 3–5 each with a pair of antero-lateral spots that hardly extend on to sides of tergites in ♂, and tergites 3–4 each with a weakly indicated dusted vitta on posterior half (Fig. 396), these better developed in ♀than in ♂. ♂epandrium matt, black. ♀ovipositor with exposed tergites dusted. Sternites 1–5 grey dusted, but sternite 5 subshining in ♂. Tergites 3–5 each with a strong lateral discal seta in ♂, and tergite 5 with 3 pairs of erect marginals, these weak in ♀. Sternites 2–5 with only sparse setulae (Fig. 397).  Ƌ terminalia. Epandrium separated from tergite 5 by a narrow syntergosternite 8, which has 2 spiracles; strongly produced ventrally (Fig. 398). Tergite 6 and sternite 6 absent. Surstylus absent, i.e. fused to epandrium without trace (Fig. 398). Cercal plate only narrowly divided ventrally (Fig. 399). Hypandrium attached at two points: to a rod originating from wall of epandrium close to upper outer edge of cercal plate; and directly to a protuberance at lower anterior corner of epandrium. Phallic complex (Fig. 400): hypandrium narrow, branching posteriorly. the upper branch forming a bridge over base of phallapodeme, the lower branch meeting postgonite; praegonite absent, postgonite long, curved; phallapodeme connected to hypandrium by a pair of bridges; phallus broadly triangular, short, simple, strongly sclerotised, juxta withdrawn, without spinules, without flanking processes.   ♀ovipositor( Figs 401–403). Tergite 6 divided dorsally, weakly sclerotised medially; tergite 7 complete; tergite 8 divided dorsally, each half displaced laterally, with a saucer-like depression on lower posterior part. Sternites 6 and 7 each represented by several setae and bristle-dots; sternite 8 represented by 2 small plates concealed in the invaginated membrane beneath the tip of segment 8. Epiproct V-shaped, with 1 long seta, connected to the cercus. Hypoproct large, extended posteriorly, with several stout spines at apex. Cercus plate-like, subtriangular, with several spines posteriorly. 3 spermathecae.  Measurements. Wing-length, 3.0– 3.5 mm( ♂), 3.5–5.0 mm ( ♀). Body-length, 4.5–5.0 mm ( ♂), 5.0– 6.5 mm( ♀).   Biology.Adults have been collected near rivers and around creeks.   Distribution.Known only from Australia(NT, Q, NSW, ACT, V, SA). 2004851380 [459,1433,1142,1167] 1972-09-27 1972-09-28 1972-09-27 ANIC Z. Liepa Simpson Desert 191 192 1 1 2004851425 [151,1431,1178,1203] 1997-09-01 QMBA J. & A. Skevington & C. Lambkin & S. Winterton Birdsville Racetrack 191 192 1 1 Queensland 2004851468 [151,695,1214,1239] 1963-07-21 QMBA C. Speed Brisbane 191 192 1 1 Queensland 2004851443 1991-09-27 QMBA 150 -28.05 Daniels 740 144.12 Dynevor Lakes 191 192 1 1 Queensland 2004851444 [572,1122,1250,1275] 1976-06-15 QDPC J. Donaldson Roma 191 192 5 5 Queensland 2004851393 1970-01-03 ANIC P. Ferrar Townsville 191 192 1 1 Queensland 2004851503 [534,1432,1286,1311] USNM Buccleugh, T & Vaughan-Sherrin Argalong 191 192 1 1 New South Wales 2004851504 [151,993,1322,1347] 1978-04-23 1978-04-30 1978-04-23 ANIC, BMNH Z. Liepa Conargo Billabong Creek 191 192 4 4 New South Wales 2004851479 1978-05-02 ANIC Billabong & Z. Liepa Forest Creek 191 192 1 1 New South Wales 2004851422 1956-06-17 AMS D. K. McAlpine Sydney Bronte 191 192 1 1 New South Wales 2004851371 [249,1106,1394,1419] 1976-04-05 ANIC D. H. Colless Byrock 191 192 1 1 New South Wales 2004851451 1976-06-17 ANIC Liepa 25 km NW of Coonamble 191 192 3 1 2 New South Wales 2004851455 [604,1256,1430,1455] 1976-12-25 AMS Daniels 8 km NE of Cudal 191 192 1 1 New South Wales 2004851411 1971-04-24 ANIC Liepa 2.4 km NE of Eugowra 191 192 2 2 New South Wales 2004851469 [647,1252,1466,1491] 1976-12-29 ANIC Liepa 24 km N of Forbes 191 192 1 1 New South Wales 2004851447 1973-12-23 ANIC Z. Liepa Tumut Gilmore Creek 191 192 1 1 New South Wales 2004851474 1971-03-27 QMBA G. & A. Daniels Goonoo Forest 191 192 3 3 New South Wales 2004851424 [422,1139,1538,1563] 1956-11-15 USNM W. W. Wirth Griffith 191 192 1 1 New South Wales 2004851409 1925-05-01 USNM Gunnedah 191 192 1 1 New South Wales 2004851501 [270,1116,1574,1599] 1976-12-26 ANIC Z. Liepa Lachlan River 191 192 2 2 New South Wales 2004851462 1971-04-10 ANIC Z. Liepa Wagga Lake Albert 191 192 1 1 New South Wales 2004851418 1964-01-14 ANIC, BMNH Z. Liepa Lake Cullulleraine 191 192 9 5 4 New South Wales 2004851448 [258,851,1646,1671] 1926-12-15 AMS B. Bertram Meroo Meadow 191 192 1 1 New South Wales 2004851427 [862,1432,1646,1671] 1964-11-25 BMNH M. I. Nikitin Merrylands 191 192 1 1 New South Wales 2004851493 [151,1004,1682,1707] 1967-11-22 MV A. Neboiss Murray River 191 192 1 1 New South Wales 2004851472 1925-03-25 USNM, AMS Narrandera 191 192 4 2 2 New South Wales 2004851410 [436,1010,1718,1743] 1926-12-08 AMS B. Bertram Nowra Creek 191 192 1 1 New South Wales 2004851432 1984-05-30 AMS K. C. Khoo Nyngan 191 192 1 1 New South Wales 2004851488 [251,931,1754,1779] 1975-09-22 ANIC Liepa 45 km SE of Nyngan 191 192 1 1 New South Wales 2004851359 1969-11-24 ANIC R. W. Matthews Packsaddle 191 192 1 1 New South Wales 2004851415 [619,1366,1790,1815] 1988-10-09 AMS D. K. McAlpine Pilliga Salt Caves Dam 191 192 1 1 New South Wales 2004851438 1988-10-08 1988-10-09 1988-10-08 AMS R. de Keyzer & K. C. Khoo Pilliga Salt Caves Dam 191 192 1 1 New South Wales 2004851452 1973-12-24 ANIC Z. Liepa Tumut River 191 192 2 2 New South Wales 2004851467 [543,1150,1862,1887] 1976-06-19 ANIC Liepa 40 km N of Walgett 191 192 1 1 New South Wales 2004851399 1976-12-27 ANIC Z. Liepa Wallaroi Creek 191 192 2 1 1 New South Wales 2004851483 19. i. 1968, 3. ii. 1969, 4. ii. 1964, 21. ii. 1969, 14. xi. 1968, 25. xi. 1959 10. xii. 1968, 13. xii. 1966, 13. xii. 1968 19. i. 1968, 3. ii. 1969, 4. ii. 1964, 21. ii. 1969, 14. xi. 1968, 25. xi. 1959 ANIC I. F. B. Common Black Mtn 191 192 10 6 4 Australian Capital Territory 2004851470 [463,1167,1970,1995] 1963-10-31 ANIC D. H. Colless Griffin Lake Burley 191 192 3 2 1 Australian Capital Territory 2004851453 1968-11-19 ANIC Colless 2 miles E of Mt Coree 191 192 1 1 Australian Capital Territory 2004851491 [645,1356,2006,2031] 1953-12-18 MV A. Neboiss Clarkefield 191 192 1 1 Victoria 2004851401 1966-12-26 ANIC, BMNH Z. Liepa 192 193 Lake Cullulleraine 191 192 8 5 3 Victoria 2004851450 1974-09-06 ANIC Liepa 47 km NW of Hopetoun 192 193 1 1 Victoria 2004851403 [402,1242,188,213] 1966-12-24 ANIC Z. Liepa Ovens River 192 193 1 1 Victoria 2004851431 1906-12-10 BMNH W. Wesche Port Melbourne 192 193 1 1 Victoria holotype 2004851463 [699,1296,223,248] 1973-01-23 BMNH E. A. Fonseca Tatyoon 192 193 12 5 7 Victoria holotype 2004851445 1974-09-04 1974-09-07 1974-09-04 ANIC Z. Liepa Wyperfield Nat. Park 192 193 2 2 Victoria holotype 2004851484 1997-09-02 QMBA J. & A. Skevington -26.41 Koonchera Waterhole 745 139.3 Koonchera Dune 192 193 2 1 1 South Australia holotype 2004851347 1997-09-02 QMBA C. Lambkin -26.45 Pandiburra Bore 745 139.24 192 193 4 2 2 South Australia holotype