Dasyhelea monticola Ingram and Macfie, 1931: 188
Wirth, 1974: 17
Borkent and Wirth, 1997: 56
Borkent and Spinelli, 2000: 25
Borkent and Spinelli, 2007: 60
A taxonomic revision of the Patagonian species of the Dasyhelea mutabilis species group with a phylogenetic analysis of New World species (Diptera: Ceratopogonidae)
Díaz, Florentina
Spinelli, Gustavo R.
Donato, Mariano
Ronderos, María M.
Journal of Natural History
2014
J. Nat. Hist.
2014-07-07
48
35 - 36
2117
2175
348PM
Ingram and Macfie, 1931
Ingram and Macfie Paramere
1931
[358,889,1444,1468]
Insecta
Ceratopogonidae
Dasyhelea
Animalia
Diptera
26
2142
Arthropoda
species
monticola
( Figures 7, 19A-B, 21)
Dasyhelea monticola Ingram and Macfie, 1931: 188(male, female, Chile); Wirth, 1974: 17(in catalogue of south USAspecies); Borkent and Wirth, 1997: 56(in World catalogue); Borkent and Spinelli, 2000: 25(in New World catalogue south of the USA); Borkent and Spinelli, 2007: 60(in Neotropical catalogue); Borkent, 2017: 67 (in online World catalogue).
Diagnosis Only species of Dasyheleain Patagonia with the following combination of characters: male paramere J-shaped. Female subgenital plate circular; cubital fork of wing distad to level of end of costa; spermatheca ovoid with oblique neck. Redescription of male.Similar to female with usual sexual differences. Antennal flagellum as in Figure 7A. Palpus ( Figure 7B) brown; segment 3 bearing scattered sensilla on basomesal surface; PR3.57–4.66 (4.14, n= 10). Scutellum with 7–8 large, 1–2 thinner setae. Wing ( Figure 19B) length 0.69–1.14 (0.94, n= 10) mm, width 0.24–0.36 (0.31, n= 10) mm; CR 0.38–0.44 (0.41, n= 10). Genitalia ( Figure 7C): tergite 9 rounded distally, reaching level of apex of gonocoxites, apicolateral process moderately short, stout, with conspicuous apical seta; cercus very small with 2–3 small setae; sternite 9 ( Figure 7D) 0.5 length of greatest width, posteromedian projection rounded, reaching one-third of aedeagus length. Gonocoxite stout, 1.8 × longer than greatest width, with short, blunt anteromedian process; gonostylus 0.9 length of gonocoxite, base moderately broad, slightly curved, tapering slightly distally, apex barely hooked, tip pointed. Paramere and gonocoxal apodemes ( Figure 7E) forming an asymmetrical structure; gonocoxal apodemes stout, right apodeme barely fused with paramere, left broadly fused with paramere; paramere greatly elongate, J-shaped. Aedeagus 0.9 length of greatest width, W/L ratio 0.50–0.60 (0.53, n= 7); anterior margin concave; basal arms heavily sclerotized, stout, recurved 60°, posteriorly directed; posterolateral arms stout, straight, parallel, tapering to slender, sharply pointed tips, each with mesal, subapical, outer triangular process; posteromedian projection short, slender, triangular, tapering distally, with simple, pointed tip. Redescription of female.Head dark brown. Eyes contiguous for distance of the width of 3 ommatidia. Antennal flagellum ( Figure 7F) pale; AR 0.85–1.03 (0.91, n= 10). Clypeus ( Figure 7G) with 4 pairs of setae. Palpus ( Figure 7H) pale brown; segment 3 with 2–3 subbasomesal capitate sensilla; PR. 3.20–4.00 (3.60, n= 10). Thorax. Scutum dark brown, scutellum pale brown with 6–7 large, 2–3 thinner setae; apex of hind tibia with 4–6 spines; foreleg TR 1.59–2.03 (1.70, n= 10), midleg TR 1.67–2.02 (1.82, n= 10), hind leg TR 1.77–2.00 (1.87, n= 10). Wing ( Figure 19A) length 0.75–0.96 (0.85, n= 10) mm, width 0.30–0.42 (0.37, n= 10) mm; CR 0.30–0.44 (0.39, n= 10); membrane slightly infuscated, densely covered with macrotrichia; radial cells obliterated; cubital fork slightly distad to level of apex of costa. Haltere dark brown, knob base whitish. Abdomen. Pale brown. Subgenital plate ( Figure 7I) circular with narrow lumen, without posteromedian projection; posterolateral arms moderately broad, distal portions slender, recurved nearly 90°. Spermatheca ( Figure 7J) ovoid heavily sclerotized, 60 × 36 µm, neck oblique, short, measuring 12 µm.
Distribution( Figure 21) Argentina( Neuquen, Río Negro, Chubut, Santa Cruz, Tierra del Fuego), Chile(Malleco, Osorno).
Type material Holotypemale, allotypefemale, Chile, Llanquihue prov., Casa Pangue, 4/ 10 December 1926, F. and M. Edwards( BMNHexamined). Other specimens examined Argentina, Neuquen prov., Parque Nacional Lanin, lago Ruca-Choroy, 14 January 1994, G. Spinelli, 2 females( MLPA); 20 kmsouth Moquehue, 14 February 1994, G. Spinelli, 1 female( MLPA); lago Aluminé, 22 December 1996/ 3 January 1997, G. Spinelli, 1 female( MLPA); lago Huechulaufquen, 28 February 1998, G. Spinelli, 1 female( MLPA); arroyo camino a las Termas del Queñi, 7 February 1999, P. Marino, 2 females, sweep net( MLPA); Villa la Angostura, 11/ 15 January 2004, G. Spinelli, 1 male, at light ( MLPA). Río Negro prov., Parque Nacional Nahuel Huapi, Puerto Blest, 9 December 1992, G. Spinelli, 2 males, 6 females( MLPA); Rincónde Comi-Co, 41°08′35.1“ S, 67° 27′34.6″ W, 1000 m, 7 December 2006, G. Spinelli, 1 male, Malaise trap( MLPA); Parque Nacional Nahuel Huapi, Mallin la Cortadera, 41°05′13″ S, 71°48′26“ W, 769 m, 11/ 30 December 2006; Garré-Montes de Oca, 1 female, Malaise trap( MLPA); same data except 14 December 2006/ 8 January 2007, 3 males, 1 female( MLPA); same data except 8 January/ 3 February 2007, 8 males, 5 females( MLPA) 1 male, 1 female( BMNH), 1 male, 1 female, ( USNM), 1 male, 1 female, ( CNCI); Parque Nacional Nahuel Huapi, Mallinlos Patos, 41°15′48.6″ S, 71°17′50.3“ W, 1020 m, 20 December 2006/ 3 January 2007, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); Parque Nacional Nahuel Huapi, Mallin La Heladera, 41°05′13″ S, 71°48′26″ W, 769 m, 14 December 2006/ 8 January 2007, Garré-Montes de Oca, 2 males, Malaise trap( MLPA); same data except 8 January/ 3 February 2007, 1 male, 3 females( MLPA); Parque Nacional Nahuel Huapi, arroyo Vantitter, 41°11′53.8″ S, 71°29′47.3″ W, 1748 m, 24 January 2007, Garré-Montes de Oca, 1 male, sweep net( MLPA); Parque Nacional Nahuel Huapi, Laguna Schmoll, 41°11’36.7’’ S, 71°29’51.2’’ W, 1925 m, 24 January 2007, Garré-Montes de Oca, 2 females, sweep net( MLPA); Parque Nacional Nahuel Huapi, cerro Catedral, laguna Toncek, 41°11′52.2″ S, 71°29′31.2″ W, 1770 m, 20 February 2007, Cazorla-Donato, 1 male, 1 female, sweep net( MLPA); Parque Nacional Nahuel Huapi, Manso Superior, 41°14′8.1″ S, 71°46′58.5″ W, 845 m, 7 February/ 2 March 2007, Garré- Montes de Oca, 1 female, Malaise trap( MLPA); Parque Nacional Nahuel Huapi, arroyo Llum, 41°16′13.3″ S, 71°30′56.7″ W, 857 m, 4/ 25 January 2007, Garré-Montes de Oca, 1 male, 1 female, Malaise trap( MLPA); Parque Nacional Nahuel Huapi, arroyo Blanco( Villa Traful), 40°39′03.4″ S, 71°24′45.8″ W, 822 m, 12 December 2007/ 7 March 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); same data except 21 January/ 7 February 2008, 1 female, ( MLPA); Parque Nacional Nahuel Huapi, arroyo Neuquenco, 40°28′48.3″ S, 71°36′44.1″ W, 809 m, 9/ 23 January 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); same data except 23 January/ 4 February 2008, 1 male, 2 females( MLPA); Parque Nacional Nahuel Huapi, laguna Mercedes, 40°52′43.4″ S, 71°34′41″ W, 899 m, 3 January/ 21 February 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); Parque Nacional Nahuel Huapi, río Pichitraful, 40°42′46″ S, 71°47′23″ W, 789 m, 4/ 23 February 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); Parque Nacional Nahuel Huapi, Lago Espejo, 40°38′49.3″ S, 71°42′12.7″ W, 903 m, 23 February/ 5 March 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA). Chubut prov., lago Fontana, 12 December 1995, G. Spinelli, 2 males, sweep net( MLPA); cerro Galera, 2/ 4 December 2002, G. Spinelli, 2 males, Malaise trap( MLPA); Puerto Patriada, 500 m Epuyén, 2/ 8 December 2008 P. Pessacq, 2 males, Malaise trap( MLPA); same data except 3/ 29 January 2009, 1 male( MLPA). Santa Cruz prov., lago del Desierto, 19 January 1995, G. Spinelli, 2 females, sweep net( MLPA); same data except 9 December 1996, 2 females( MLPA). Tierra del Fuego prov., Parque Nacional Tierra del Fuego, laguna Negra(turbera), 54°50′42.1″ S, 68°35′17.9″ W, 15 m, 1/ 4 December 2008, Donato-Spinelli, 2 males, Malaise trap( MLPA); same data except 4/ 7 December 2008, 4 males, 3 females( MLPA). Chile, Malleco prov., Parque Nacional Conguillio, 22 December 1984, J. A. Downes, 1 female, sweep net( JAD 1677/1) ( CNCI); Parque Nacional Nahuelbuta, 26 December 1984, J. A. Downes, 1 male, sweep net( JAD 1682/2) ( CNCI). Osorno prov., Pucatrihue, 1 December 1992, G. Spinelli, 1 male( MLPA); Parque Nacional Vicente Perez Rosales, Casa Pangue, 41°02′55″ S, 71°52′31.2″ W, 366 m, 18 February 2008, M. Donato, 2 females, sweep net( MLPA). Phylogenetic relationships Dasyhelea monticolais the sister species of D. suareziSpinelli and Ronderos. Theclade ( D. monticola, D. suarezi) is suppoted by the characters “parameres” [character 13 (1)]; “shape of posteromedian projection of aedeagus” [character 19 (2)] and “cubital fork (female)” [character 23 (1)].
Argentina
Neuquen, Rio Negro, Chubut, Santa Cruz, Tierra del Fuego
28
2144
1
Neuquen, Rio Negro, Chubut, Santa Cruz, Tierra del Fuego
1926-12-10
BMNH
F. and M. Edwards
Chile
Llanquihue prov.
Casa Pangue
29
2145
2
holotype
1994-01-14
MLPA
G. Spinelli
Argentina
Ruca-Choroy
Parque Nacional Lanin
29
2145
2
2
Neuquen
1994-02-14
MLPA
Spinelli
Argentina
20 km south Moquehue
29
2145
1
1
Neuquen prov.
1996-12-22
1997-01-03
1996-12-22
MLPA
G. Spinelli
Argentina
Alumine
29
2145
1
1
Neuquen prov.
1998-02-28
MLPA
G. Spinelli
Argentina
Huechulaufquen
29
2145
1
1
Neuquen prov.
1999-02-07
MLPA
Marino
Argentina
Termas del Queni
29
2145
2
2
Neuquen prov.
2004-01-15
MLPA
Spinelli
Argentina
Villa la Angostura
29
2145
1
1
Neuquen prov.
1992-12-09
MLPA
G. Spinelli
Argentina
Puerto Blest
Parque Nacional Nahuel Huapi
29
2145
8
6
2
Rio Negro
2006-12-07
MLPA
G. Spinelli
Argentina
1000
-41.143085
Rincon
1
-67.45961
29
2145
1
1
Rio Negro prov.
2006-12-30
MLPA
Argentina
Parque Nacional Nahuel Huapi
769
-41.086945
Garre-Montes de Oca
19
-71.80723
Mallin la Cortadera
29
2145
1
1
Rio Negro prov.
2006-12-14
2007-01-08
2006-12-14
MLPA
Argentina
Parque Nacional Nahuel Huapi
769
-41.086945
Garre-Montes de Oca
19
-71.80723
Mallin la Cortadera
29
2145
4
1
3
Rio Negro prov.
2007-02-03
2007-01-08
2006-12-14
MLPA
Argentina
Parque Nacional Nahuel Huapi
769
-41.086945
January
19
-71.80723
Mallin la Cortadera
29
2145
13
5
8
Rio Negro prov.
[413,722,803,827]
2007-02-03
2007-01-08
2006-12-14
BMNH
Argentina
Parque Nacional Nahuel Huapi
769
true
-41.086945
January
19
-71.80723
Mallin la Cortadera
29
2145
2
1
1
Rio Negro prov.
[737,1050,803,827]
USNM
Argentina
29
2145
2
1
1
Rio Negro prov.
CNCI
Argentina
29
2145
2
1
1
Rio Negro prov.
2006-12-20
2007-01-03
2006-12-20
MLPA
Argentina
Parque Nacional Nahuel Huapi
1020
-41.2635
Patos
1
-71.2973
Mallin
29
2145
1
1
Rio Negro prov.
2006-12-14
2007-01-08
2006-12-14
MLPA
Argentina
Parque Nacional Nahuel Huapi
769
-41.086945
Garre-Montes de Oca
19
-71.80723
Mallin La Heladera
29
2145
2
2
Rio Negro prov.
2007-02-03
2007-01-08
2006-12-14
MLPA
Argentina
Parque Nacional Nahuel Huapi
769
-41.086945
January
19
-71.80723
Mallin La Heladera
29
2145
4
3
1
Rio Negro prov.
2007-01-24
MLPA
Argentina
Parque Nacional Nahuel Huapi
1748
-41.19828
Garre-Montes de Oca
1
-71.49647
Vantitter
29
2145
1
1
Rio Negro prov.
2007-01-24
MLPA
Argentina
Parque Nacional Nahuel Huapi
1925
-41.193527
Garre-Montes de Oca
1
-71.49755
Laguna Schmoll
29
2145
2
2
Rio Negro prov.
2007-02-20
MLPA
Cazorla-Donato
Argentina
Parque Nacional Nahuel Huapi
1770
-41.197834
Toncek
1
-71.492
Catedral
29
2145
2
1
1
Rio Negro prov.
2007-03-02
MLPA
Argentina
Parque Nacional Nahuel Huapi
845
-41.235584
February
1
-71.78292
Manso Superior
29
2145
1
1
Rio Negro prov.
2007-01-25
MLPA
Argentina
Parque Nacional Nahuel Huapi
857
-41.27036
Garre-Montes de Oca
1
-71.51575
Llum
29
2145
2
1
1
Rio Negro prov.
2007-12-12
2008-03-07
2007-12-12
MLPA
Argentina
Parque Nacional Nahuel Huapi
822
-40.650948
Villa Traful
1
-71.41273
Blanco
29
2145
1
1
Rio Negro prov.
2008-02-07
2008-03-07
2007-12-12
MLPA
Argentina
Parque Nacional Nahuel Huapi
822
-40.650948
January
1
-71.41273
Blanco
29
2145
1
1
Rio Negro prov.
2008-01-23
MLPA
Argentina
Parque Nacional Nahuel Huapi
809
-40.480083
Garre-Montes de Oca
1
-71.61225
Neuquenco
29
2145
1
1
Rio Negro prov.
2008-02-04
MLPA
Argentina
Parque Nacional Nahuel Huapi
809
-40.480083
January
1
-71.61225
Neuquenco
29
2145
3
2
1
Rio Negro prov.
2008-02-21
MLPA
Argentina
Parque Nacional Nahuel Huapi
899
-40.87872
January
11
-71.57806
Mercedes
29
2145
1
1
Rio Negro prov.
2008-02-23
MLPA
Argentina
Parque Nacional Nahuel Huapi
789
-40.71278
Garre-Montes de Oca
19
-71.78972
rio Pichitraful
29
2145
1
1
Rio Negro prov.
2008-03-05
MLPA
Argentina
Parque Nacional Nahuel Huapi
903
30
2146
-40.64703
February
1
-71.70352
Lago Espejo
29
2145
1
1
Rio Negro prov.
1995-12-12
MLPA
G. Spinelli
Argentina
Fontana
30
2146
2
2
Chubut
2002-12-04
MLPA
G. Spinelli
Argentina
Galera
30
2146
2
2
Chubut prov.
2008-12-08
MLPA
P. Pessacq
Argentina
500
Epuyen
Puerto Patriada
30
2146
2
2
Chubut prov.
[411,1027,296,320]
2009-01-29
MLPA
P. Pessacq
Argentina
500
Epuyen
Puerto Patriada
30
2146
1
1
Chubut prov.
1995-01-19
MLPA
G. Spinelli
Argentina
Desierto
30
2146
2
2
Santa Cruz
[258,897,359,383]
1996-12-09
MLPA
G. Spinelli
Argentina
Desierto
30
2146
2
2
Santa Cruz prov.
Parque Nacional
Argentina
Tierra del Fuego prov.
30
2146
1
Tierra del Fuego - Antartida e Islas del Atlantico Sur
2008-12-04
MLPA
Donato-Spinelli
Argentina
15
-54.845028
Negra
1
-68.58831
30
2146
2
2
Tierra del Fuego - Antartida e Islas del Atlantico Sur
[257,1015,455,479]
2008-12-07
MLPA
Donato-Spinelli
Argentina
15
-54.845028
Negra
1
-68.58831
30
2146
7
3
4
Tierra del Fuego
1984-12-22
CNCI
J. A. Downes
Chile
Malleco prov.
Parque Nacional Conguillio
30
2146
JAD 1677
1
1
1984-12-26
CNCI
J. A. Downes
Chile
Parque Nacional Nahuelbuta
30
2146
JAD 1682
1
1
1992-12-01
MLPA
G. Spinelli
Chile
Osorno prov.
Pucatrihue
30
2146
1
1
2008-02-18
MLPA
M. Donato
Chile
366
-41.04861
Parque Nacional Vicente Perez Rosales
15
-71.875336
Casa Pangue
30
2146
2
2