Dasyhelea monticola Ingram and Macfie, 1931: 188 Wirth, 1974: 17 Borkent and Wirth, 1997: 56 Borkent and Spinelli, 2000: 25 Borkent and Spinelli, 2007: 60 A taxonomic revision of the Patagonian species of the Dasyhelea mutabilis species group with a phylogenetic analysis of New World species (Diptera: Ceratopogonidae) Díaz, Florentina Spinelli, Gustavo R. Donato, Mariano Ronderos, María M. Journal of Natural History 2014 J. Nat. Hist. 2014-07-07 48 35 - 36 2117 2175 348PM Ingram and Macfie, 1931 Ingram and Macfie Paramere 1931 [358,889,1444,1468] Insecta Ceratopogonidae Dasyhelea Animalia Diptera 26 2142 Arthropoda species monticola  ( Figures 7, 19A-B, 21)      Dasyhelea monticola Ingram and Macfie, 1931: 188(male, female, Chile);  Wirth, 1974: 17(in catalogue of south USAspecies);  Borkent and Wirth, 1997: 56(in World catalogue);  Borkent and Spinelli, 2000: 25(in New World catalogue south of the USA);  Borkent and Spinelli, 2007: 60(in Neotropical catalogue); Borkent, 2017: 67 (in online World catalogue).    Diagnosis Only species of  Dasyheleain Patagonia with the following combination of characters: male paramere J-shaped. Female subgenital plate circular; cubital fork of wing distad to level of end of costa; spermatheca ovoid with oblique neck.  Redescription of male.Similar to female with usual sexual differences. Antennal flagellum as in Figure 7A. Palpus ( Figure 7B) brown; segment 3 bearing scattered sensilla on basomesal surface; PR3.57–4.66 (4.14, n= 10). Scutellum with 7–8 large, 1–2 thinner setae. Wing ( Figure 19B) length 0.69–1.14 (0.94, n= 10) mm, width 0.24–0.36 (0.31, n= 10) mm; CR 0.38–0.44 (0.41, n= 10). Genitalia ( Figure 7C): tergite 9 rounded distally, reaching level of apex of gonocoxites, apicolateral process moderately short, stout, with conspicuous apical seta; cercus very small with 2–3 small setae; sternite 9 ( Figure 7D) 0.5 length of greatest width, posteromedian projection rounded, reaching one-third of aedeagus length. Gonocoxite stout, 1.8 × longer than greatest width, with short, blunt anteromedian process; gonostylus 0.9 length of gonocoxite, base moderately broad, slightly curved, tapering slightly distally, apex barely hooked, tip pointed. Paramere and gonocoxal apodemes ( Figure 7E) forming an asymmetrical structure; gonocoxal apodemes stout, right apodeme barely fused with paramere, left broadly fused with paramere; paramere greatly elongate, J-shaped. Aedeagus 0.9 length of greatest width, W/L ratio 0.50–0.60 (0.53, n= 7); anterior margin concave; basal arms heavily sclerotized, stout, recurved 60°, posteriorly directed; posterolateral arms stout, straight, parallel, tapering to slender, sharply pointed tips, each with mesal, subapical, outer triangular process; posteromedian projection short, slender, triangular, tapering distally, with simple, pointed tip.  Redescription of female.Head dark brown. Eyes contiguous for distance of the width of 3 ommatidia. Antennal flagellum ( Figure 7F) pale; AR 0.85–1.03 (0.91, n= 10). Clypeus ( Figure 7G) with 4 pairs of setae. Palpus ( Figure 7H) pale brown; segment 3 with 2–3 subbasomesal capitate sensilla; PR. 3.20–4.00 (3.60, n= 10). Thorax. Scutum dark brown, scutellum pale brown with 6–7 large, 2–3 thinner setae; apex of hind tibia with 4–6 spines; foreleg TR 1.59–2.03 (1.70, n= 10), midleg TR 1.67–2.02 (1.82, n= 10), hind leg TR 1.77–2.00 (1.87, n= 10). Wing ( Figure 19A) length 0.75–0.96 (0.85, n= 10) mm, width 0.30–0.42 (0.37, n= 10) mm; CR 0.30–0.44 (0.39, n= 10); membrane slightly infuscated, densely covered with macrotrichia; radial cells obliterated; cubital fork slightly distad to level of apex of costa. Haltere dark brown, knob base whitish. Abdomen. Pale brown. Subgenital plate ( Figure 7I) circular with narrow lumen, without posteromedian projection; posterolateral arms moderately broad, distal portions slender, recurved nearly 90°. Spermatheca ( Figure 7J) ovoid heavily sclerotized, 60 × 36 µm, neck oblique, short, measuring 12 µm.    Distribution( Figure 21)   Argentina( Neuquen, Río Negro, Chubut, Santa Cruz, Tierra del Fuego), Chile(Malleco, Osorno).    Type material   Holotypemale, allotypefemale, Chile, Llanquihue prov., Casa Pangue, 4/  10 December 1926, F. and M. Edwards( BMNHexamined).   Other specimens examined   Argentina, Neuquen prov., Parque Nacional Lanin, lago Ruca-Choroy,  14 January 1994, G. Spinelli, 2 females( MLPA);   20 kmsouth Moquehue,  14 February 1994, G. Spinelli, 1 female( MLPA); lago Aluminé,  22 December 1996/  3 January 1997, G. Spinelli, 1 female( MLPA); lago Huechulaufquen,  28 February 1998, G. Spinelli, 1 female( MLPA); arroyo camino a las Termas del Queñi,  7 February 1999, P. Marino, 2 females, sweep net( MLPA);  Villa la Angostura, 11/  15 January 2004, G. Spinelli, 1 male, at light ( MLPA).  Río Negro prov., Parque Nacional Nahuel Huapi, Puerto Blest,  9 December 1992, G. Spinelli, 2 males, 6 females( MLPA);  Rincónde Comi-Co, 41°08′35.1“ S, 67° 27′34.6″ W,  1000 m,  7 December 2006, G. Spinelli, 1 male, Malaise trap( MLPA);  Parque Nacional Nahuel Huapi, Mallin la Cortadera, 41°05′13″ S, 71°48′26“ W,  769 m, 11/  30 December 2006; Garré-Montes de Oca, 1 female, Malaise trap( MLPA); same data except  14 December 2006/  8 January 2007, 3 males, 1 female( MLPA); same data except 8 January/  3 February 2007, 8 males, 5 females( MLPA)  1 male, 1 female( BMNH),  1 male, 1 female, ( USNM),  1 male, 1 female, ( CNCI);  Parque Nacional Nahuel Huapi, Mallinlos Patos, 41°15′48.6″ S, 71°17′50.3“ W,  1020 m,  20 December 2006/  3 January 2007, Garré-Montes de Oca, 1 female, Malaise trap( MLPA);  Parque Nacional Nahuel Huapi, Mallin La Heladera, 41°05′13″ S, 71°48′26″ W,  769 m,  14 December 2006/  8 January 2007, Garré-Montes de Oca, 2 males, Malaise trap( MLPA); same data except 8 January/  3 February 2007, 1 male, 3 females( MLPA);  Parque Nacional Nahuel Huapi, arroyo Vantitter, 41°11′53.8″ S, 71°29′47.3″ W,  1748 m,  24 January 2007, Garré-Montes de Oca, 1 male, sweep net( MLPA);  Parque Nacional Nahuel Huapi, Laguna Schmoll, 41°11’36.7’’ S, 71°29’51.2’’ W,  1925 m,  24 January 2007, Garré-Montes de Oca, 2 females, sweep net( MLPA);  Parque Nacional Nahuel Huapi, cerro Catedral, laguna Toncek, 41°11′52.2″ S, 71°29′31.2″ W,  1770 m,  20 February 2007, Cazorla-Donato, 1 male, 1 female, sweep net( MLPA);  Parque Nacional Nahuel Huapi, Manso Superior, 41°14′8.1″ S, 71°46′58.5″ W,  845 m, 7 February/  2 March 2007, Garré- Montes de Oca, 1 female, Malaise trap( MLPA);  Parque Nacional Nahuel Huapi, arroyo Llum, 41°16′13.3″ S, 71°30′56.7″ W,  857 m, 4/  25 January 2007, Garré-Montes de Oca, 1 male, 1 female, Malaise trap( MLPA);  Parque Nacional Nahuel Huapi, arroyo Blanco( Villa Traful), 40°39′03.4″ S, 71°24′45.8″ W,  822 m,  12 December 2007/  7 March 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); same data except 21 January/  7 February 2008, 1 female, ( MLPA);  Parque Nacional Nahuel Huapi, arroyo Neuquenco, 40°28′48.3″ S, 71°36′44.1″ W,  809 m, 9/  23 January 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA); same data except 23 January/  4 February 2008, 1 male, 2 females( MLPA);  Parque Nacional Nahuel Huapi, laguna Mercedes, 40°52′43.4″ S, 71°34′41″ W,  899 m, 3 January/  21 February 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA);  Parque Nacional Nahuel Huapi, río Pichitraful, 40°42′46″ S, 71°47′23″ W,  789 m, 4/  23 February 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA);  Parque Nacional Nahuel Huapi, Lago Espejo, 40°38′49.3″ S, 71°42′12.7″ W,  903 m, 23 February/  5 March 2008, Garré-Montes de Oca, 1 female, Malaise trap( MLPA).  Chubut prov., lago Fontana,  12 December 1995, G. Spinelli, 2 males, sweep net( MLPA); cerro Galera, 2/  4 December 2002, G. Spinelli, 2 males, Malaise trap( MLPA);  Puerto Patriada,  500 m Epuyén, 2/  8 December 2008 P. Pessacq, 2 males, Malaise trap( MLPA); same data except 3/  29 January 2009, 1 male( MLPA).  Santa Cruz prov., lago del Desierto,  19 January 1995, G. Spinelli, 2 females, sweep net( MLPA); same data except  9 December 1996, 2 females( MLPA).  Tierra del Fuego prov., Parque Nacional  Tierra del Fuego, laguna Negra(turbera), 54°50′42.1″ S, 68°35′17.9″ W,  15 m, 1/  4 December 2008, Donato-Spinelli, 2 males, Malaise trap( MLPA); same data except 4/  7 December 2008, 4 males, 3 females( MLPA).   Chile, Malleco prov., Parque Nacional Conguillio,  22 December 1984, J. A. Downes, 1 female, sweep net( JAD 1677/1) ( CNCI);  Parque Nacional Nahuelbuta,  26 December 1984, J. A. Downes, 1 male, sweep net( JAD 1682/2) ( CNCI).  Osorno prov., Pucatrihue,  1 December 1992, G. Spinelli, 1 male( MLPA);  Parque Nacional Vicente Perez Rosales, Casa Pangue, 41°02′55″ S, 71°52′31.2″ W,  366 m,  18 February 2008, M. Donato, 2 females, sweep net( MLPA).   Phylogenetic relationships   Dasyhelea monticolais the sister species of  D. suareziSpinelli and Ronderos. Theclade (  D. monticola,  D. suarezi) is suppoted by the characters “parameres” [character 13 (1)]; “shape of posteromedian projection of aedeagus” [character 19 (2)] and “cubital fork (female)” [character 23 (1)]. Argentina Neuquen, Rio Negro, Chubut, Santa Cruz, Tierra del Fuego 28 2144 1 Neuquen, Rio Negro, Chubut, Santa Cruz, Tierra del Fuego 1926-12-10 BMNH F. and M. Edwards Chile Llanquihue prov. Casa Pangue 29 2145 2 holotype 1994-01-14 MLPA G. Spinelli Argentina Ruca-Choroy Parque Nacional Lanin 29 2145 2 2 Neuquen 1994-02-14 MLPA Spinelli Argentina 20 km south Moquehue 29 2145 1 1 Neuquen prov. 1996-12-22 1997-01-03 1996-12-22 MLPA G. Spinelli Argentina Alumine 29 2145 1 1 Neuquen prov. 1998-02-28 MLPA G. Spinelli Argentina Huechulaufquen 29 2145 1 1 Neuquen prov. 1999-02-07 MLPA Marino Argentina Termas del Queni 29 2145 2 2 Neuquen prov. 2004-01-15 MLPA Spinelli Argentina Villa la Angostura 29 2145 1 1 Neuquen prov. 1992-12-09 MLPA G. Spinelli Argentina Puerto Blest Parque Nacional Nahuel Huapi 29 2145 8 6 2 Rio Negro 2006-12-07 MLPA G. Spinelli Argentina 1000 -41.143085 Rincon 1 -67.45961 29 2145 1 1 Rio Negro prov. 2006-12-30 MLPA Argentina Parque Nacional Nahuel Huapi 769 -41.086945 Garre-Montes de Oca 19 -71.80723 Mallin la Cortadera 29 2145 1 1 Rio Negro prov. 2006-12-14 2007-01-08 2006-12-14 MLPA Argentina Parque Nacional Nahuel Huapi 769 -41.086945 Garre-Montes de Oca 19 -71.80723 Mallin la Cortadera 29 2145 4 1 3 Rio Negro prov. 2007-02-03 2007-01-08 2006-12-14 MLPA Argentina Parque Nacional Nahuel Huapi 769 -41.086945 January 19 -71.80723 Mallin la Cortadera 29 2145 13 5 8 Rio Negro prov. [413,722,803,827] 2007-02-03 2007-01-08 2006-12-14 BMNH Argentina Parque Nacional Nahuel Huapi 769 true -41.086945 January 19 -71.80723 Mallin la Cortadera 29 2145 2 1 1 Rio Negro prov. [737,1050,803,827] USNM Argentina 29 2145 2 1 1 Rio Negro prov. CNCI Argentina 29 2145 2 1 1 Rio Negro prov. 2006-12-20 2007-01-03 2006-12-20 MLPA Argentina Parque Nacional Nahuel Huapi 1020 -41.2635 Patos 1 -71.2973 Mallin 29 2145 1 1 Rio Negro prov. 2006-12-14 2007-01-08 2006-12-14 MLPA Argentina Parque Nacional Nahuel Huapi 769 -41.086945 Garre-Montes de Oca 19 -71.80723 Mallin La Heladera 29 2145 2 2 Rio Negro prov. 2007-02-03 2007-01-08 2006-12-14 MLPA Argentina Parque Nacional Nahuel Huapi 769 -41.086945 January 19 -71.80723 Mallin La Heladera 29 2145 4 3 1 Rio Negro prov. 2007-01-24 MLPA Argentina Parque Nacional Nahuel Huapi 1748 -41.19828 Garre-Montes de Oca 1 -71.49647 Vantitter 29 2145 1 1 Rio Negro prov. 2007-01-24 MLPA Argentina Parque Nacional Nahuel Huapi 1925 -41.193527 Garre-Montes de Oca 1 -71.49755 Laguna Schmoll 29 2145 2 2 Rio Negro prov. 2007-02-20 MLPA Cazorla-Donato Argentina Parque Nacional Nahuel Huapi 1770 -41.197834 Toncek 1 -71.492 Catedral 29 2145 2 1 1 Rio Negro prov. 2007-03-02 MLPA Argentina Parque Nacional Nahuel Huapi 845 -41.235584 February 1 -71.78292 Manso Superior 29 2145 1 1 Rio Negro prov. 2007-01-25 MLPA Argentina Parque Nacional Nahuel Huapi 857 -41.27036 Garre-Montes de Oca 1 -71.51575 Llum 29 2145 2 1 1 Rio Negro prov. 2007-12-12 2008-03-07 2007-12-12 MLPA Argentina Parque Nacional Nahuel Huapi 822 -40.650948 Villa Traful 1 -71.41273 Blanco 29 2145 1 1 Rio Negro prov. 2008-02-07 2008-03-07 2007-12-12 MLPA Argentina Parque Nacional Nahuel Huapi 822 -40.650948 January 1 -71.41273 Blanco 29 2145 1 1 Rio Negro prov. 2008-01-23 MLPA Argentina Parque Nacional Nahuel Huapi 809 -40.480083 Garre-Montes de Oca 1 -71.61225 Neuquenco 29 2145 1 1 Rio Negro prov. 2008-02-04 MLPA Argentina Parque Nacional Nahuel Huapi 809 -40.480083 January 1 -71.61225 Neuquenco 29 2145 3 2 1 Rio Negro prov. 2008-02-21 MLPA Argentina Parque Nacional Nahuel Huapi 899 -40.87872 January 11 -71.57806 Mercedes 29 2145 1 1 Rio Negro prov. 2008-02-23 MLPA Argentina Parque Nacional Nahuel Huapi 789 -40.71278 Garre-Montes de Oca 19 -71.78972 rio Pichitraful 29 2145 1 1 Rio Negro prov. 2008-03-05 MLPA Argentina Parque Nacional Nahuel Huapi 903 30 2146 -40.64703 February 1 -71.70352 Lago Espejo 29 2145 1 1 Rio Negro prov. 1995-12-12 MLPA G. Spinelli Argentina Fontana 30 2146 2 2 Chubut 2002-12-04 MLPA G. Spinelli Argentina Galera 30 2146 2 2 Chubut prov. 2008-12-08 MLPA P. Pessacq Argentina 500 Epuyen Puerto Patriada 30 2146 2 2 Chubut prov. [411,1027,296,320] 2009-01-29 MLPA P. Pessacq Argentina 500 Epuyen Puerto Patriada 30 2146 1 1 Chubut prov. 1995-01-19 MLPA G. Spinelli Argentina Desierto 30 2146 2 2 Santa Cruz [258,897,359,383] 1996-12-09 MLPA G. Spinelli Argentina Desierto 30 2146 2 2 Santa Cruz prov. Parque Nacional Argentina Tierra del Fuego prov. 30 2146 1 Tierra del Fuego - Antartida e Islas del Atlantico Sur 2008-12-04 MLPA Donato-Spinelli Argentina 15 -54.845028 Negra 1 -68.58831 30 2146 2 2 Tierra del Fuego - Antartida e Islas del Atlantico Sur [257,1015,455,479] 2008-12-07 MLPA Donato-Spinelli Argentina 15 -54.845028 Negra 1 -68.58831 30 2146 7 3 4 Tierra del Fuego 1984-12-22 CNCI J. A. Downes Chile Malleco prov. Parque Nacional Conguillio 30 2146 JAD 1677 1 1 1984-12-26 CNCI J. A. Downes Chile Parque Nacional Nahuelbuta 30 2146 JAD 1682 1 1 1992-12-01 MLPA G. Spinelli Chile Osorno prov. Pucatrihue 30 2146 1 1 2008-02-18 MLPA M. Donato Chile 366 -41.04861 Parque Nacional Vicente Perez Rosales 15 -71.875336 Casa Pangue 30 2146 2 2