The New World ant parasitoid genus Orasema (Hymenoptera: Eucharitidae) Baker, Austin J. Heraty, John M. Zootaxa 2020 2020-11-30 4888 1 1 84 Baker & Heraty, 2020 Baker & Heraty 2020 [151,392,1795,1822] Insecta Eucharitidae Orasema Animalia Hymenoptera 67 68 Arthropoda species masonicki sp. nov.  ( Fig. 37)   Diagnosis.Distinguished from  O. heacoxiby features provided in the key.   Description. Female.Length 1.9–2.2 mm( Fig. 37A). Color.Scape brown. Mandible brown. Femora brown with strong iridescence. Gaster dark brown with strong blue-green iridescence. Head( Fig. 37B). Head in frontal view elongate; HW:HH = 0.7–1.0; longitudinal groove between eye and torulus absent; IOD:EH = 1.0–1.1; MS:EH = 0.3–0.6; malar depression weakly impressed adjacent to mouth; epistomal sulcus distinct and sharply defined. Occiput deeply emarginate in dorsal view. Pedicel small and globose. FL:HH = 0.6–0.9; F2L:F2W = 1.0–1.5, F2L:F3L = 1.0 ( Fig. 37D). Mesosoma( Fig. 37C, F, G). ML:MH = 1.4–1.6. Mesoscutal midlobe reticulate. Scutoscutellar sulcus broad, narrowly separated from transscutal articulation; frenum reticulate; axillula reticulate. Mesepisternum reticulate laterally, weakly reticulate ventrally. Upper mesepimeron weakly reticulate; lower mesepimeron reticulate; transepimeral sulcus weakly impressed. Metepisternum laterally reticulate. HCL:HCW = 1.2–1.8, reticulate; HFL:HFW = 4.6–6.3. FWL:FWW = 2.2–3.9, FWL:ML = 1.9–2.4. Metasoma.PTL:PTW = 2.5–3.4, PTL:HCL = 1.1–1.7, lateral margin rounded, ventral sulcus present with margins narrowly separated. Apical setae of hypopygium with several setae on each side of midline.   FIGURE 37.  Orasema masonicki. Holotype female (INBIOCRI002242095): A, habitus, lateral; B, head, anterior; C, mesosoma, lateral; D, antenna; F, mesosoma, dorsal; G, propodeum, posterior. Paratype male (INBIOCRI002133102): E, antenna.  Male.Length 2–2.1 mm. HW:HH = 0.9–1.1. Scape brown; FL:HH = 0.9–1.2; F2L:F2W = 0.8–1.4 ( Fig. 37E). Tibiae yellow. PTL:PTW = 5.5–7.5, PTL:HCL = 2.2–3.3.   Hosts.Unknown.  Plant associates.Unknown.   Distribution( Fig. 35). Costa Rica: AL, GU, HE. Collected January–May.   Material examined.   Holotype. COSTA RICA. Guanacaste:Est. Pitilla, 9 kmS Santa Cecilia, P.N. Guanacaste,  700m, 10°59’28”N, 85°25’39”W,  iii.1995, P. Rios, #4359 [ ♀, deposited in INBIO: INBIOCRI02242095].    Paratypes. COSTA RICA. Alajuela: Sect. San Ramon de Dos Rios,  620m, 10°52’54”N, 85°24’41”W,  16.i- 3.ii.1995, F.A. Quesada, #4400 [1³, INBIO: INBIOCRI02192938];  20.ii-5.iii.1995, #4401 [4³ 1?, INBIO: IN-BIOCRI02138790, INBIOCRI02138802, INBIOCRI02138811, INBIOCRI02138832, INBIOCRI02138835]; C. Cano, #4396 [15³ 4♀, INBIO: INBIOCRI02133093, INBIOCRI02133101–02, INBIOCRI02133144, INBIOC- RI02133155–56, INBIOCRI02133159, INBIOCRI02133164–66, INBIOCRI02133183, INBIOCRI02133195, IN-BIOCRI02133204, INBIOCRI02133209, INBIOCRI02133213–15, INBIOCRI02133225, INBIOCRI02133227];  20.ii-3.iii.1995, #4398 [10³ 8♀, INBIO: INBIOCRI02174676–77, INBIOCRI02174679–80, INBIOCRI02174682, INBIOCRI02174686–87, INBIOCRI02174691, INBIOCRI02174714, INBIOCRI02174730, INBIOCRI02174755, INBIOCRI02174760, INBIOCRI02174776, INBIOCRI02174783, INBIOCRI02174787–88, INBIOCRI02174793, INBIOCRI02174800];  11-15.iv.1994, C. Moraga, #2840 [1³, INBIO: INBIOCRI01780190];  13-28.iii.1994, K. Tay-lor, #2763[4³ 1♀, INBIO:INBIOCRI01712287, INBIOCRI01712554, INBIOCRI01712730, INBIOCRI01712733, INBIOCRI01760375].   Guanacaste:Est. Pitilla,  9 kmS Santa Cecilia, P.N. Guanacaste,  700m, 10°59’28”N, 85°25’39”W,  2-19.iii.1992, P. Rios [ 1♀, INBIO: INBIOCRI00420518]. Sector Las Pailas,  800m, 10°46’31”N, 85°20’59”W,  12.iv-3.v.1995, K. Taylor, #4810 [2³ 1♀, INBIO: INBIOCRI02425608, INBIOCRI02425621, IN-BIOCRI02425633];  16-30.iii.1995, #4811 [8³, INBIO: INBIOCRI02403459, INBIOCRI02403595, INBIOC-RI02403602–03, INBIOCRI02403778, INBIOCRI02403840, INBIOCRI02403851, INBIOCRI02403878].   Heredia:Send. Terciopelo, Est. Magsaysay, P.N.Braulio Carrillo,  220m, 10°24’2”N, 84°02’53”W, 1991, malaise trap[1³, INBIO: INBIOCRI01194079].   Etymology.Named in honor of Paul Masonick, an entomologist who has helped to collect specimens of  Orasema, including those of the  heacoxigroup.  Unplaced  Orasemaspecies ( Figs 38–44)   Discussion.Several species could not be confidently placed to species group, especially those lacking molecular phylogenetic data. Placing these taxa into species groups described herein would make the species group key more difficult to use, so we prefer to treat them as unplaced until more data can be collected. 2988974368 1995-03 P. N. Guanacaste & P. Rios Costa Rica 700 10.991112 Santa Cecilia 21 -85.4275 69 70 1 1 Guanacaste holotype 2988974329 1994-03-13 1995-03-05 1994-03-13 IN- F. A. Quesada & C. Cano & C. Moraga Costa Rica 620 10.881667 Sect. San Ramon de Dos Rios 21 -85.41139 69 70 IN-BIOCRI02138790, INBIOCRI02138802, INBIOCRI02138811, INBIOCRI02138832, INBIOCRI02138835, IN-BIOCRI02133204, INBIOCRI02133209, INBIOCRI02133213-15, INBIOCRI02133225, INBIOCRI02133227 13 13 Alajuela paratype 2988974309 1992-03-02 1992-03-19 1992-03-02 IN- P. Rios Costa Rica 700 10.775277 Guanacaste 21 -85.349724 69 70 BIOCRI02425633 1 2 Guanacaste paratype 2988974457 1995-04-12 1995-05-03 1995-04-12 IN- K. Taylor Costa Rica 800 10.775277 Sector Las Pailas 21 -85.349724 69 70 BIOCRI02425633 1 1 Guanacaste paratype 2988974318 1995-03-16 1995-03-30 1995-03-16 IN- Costa Rica 700 10.775277 Sector Las Pailas 21 -85.349724 69 70 IN-BIOCRI02425633 1 1 Guanacaste paratype 2988974384 1992-03-02 1995-05-03 1992-03-02 IN- Costa Rica 700 10.775277 Sector Las Pailas 21 -85.349724 69 70 BIOCRI02425633 1 2 Guanacaste paratype 2988974442 Est. Magsaysay, P. N. Costa Rica 220 10.400556 Heredia 21 -84.04806 69 70 1 Heredia paratype