Coenosia
caledonica Bigot, 1885: 269
Crosskey, 1962: 445
Coenosia caledonica
Stein, 1919: 158
Curran, 1929: 7
Pygophora caledonica
Paramonov, 1961: 511
Crosskey, 1962: 445
Pont, 1989: 689
Shinonaga et al ., 1991: 333
The Muscidae (Diptera) of New Caledonia 2503
Couri, Marcia S.
Pont, Adrian C.
Daugeron, Christophe
Zootaxa
2010
2010-06-11
2503
1
1
61
6X32H
(Bigot)
Bigot
1885
[195,557,1375,1401]
Insecta
Muscidae
Pygophora
Animalia
Diptera
47
48
Arthropoda
species
caledonica
( Figs 35–37)
Coenosia? caledonica Bigot, 1885: 269, Holotypefemale, “ Nouvelle-Calédonie”, in BMNH ( Crosskey, 1962: 445). Coenosia caledonica; Stein, 1919: 158; Curran, 1929: 7. Pygophora caledonica; Paramonov, 1961: 511; Crosskey, 1962: 445; Pont, 1989: 689; Shinonaga et al., 1991: 333.
Diagnosis.Antenna yellow, postpedicel about 2.8 times as long as pedicel in females and longer in male, about 4.5 times; palpus yellowish-white; scutum with two fine brown vittae; legs yellow; wing clear in female, but with a brown spot at apex of veins R 2+ 3in male. Abdomen in male: syntergite 1+2 and tergite 3 yellow, tergites 4 and 5 grey with three brown spots. Abdomen in female with syntergite 1+2 and tergites 3–4 yellow, tergites 3 and 4 each with three brown spots, those on tergite 4 with grey areas between the spots, tergite 5 grey with 2 brown spots. Abdomen strongly modified in male, compressed laterally, and tergite 5 with a median keel, a tuft of long setae and ventrally with strong black setae on each side ( Fig. 35). Sternite 5 ( Fig. 36). Aedeagus in lateral view as in Fig. 37.
Material examined: MNHN. New Caledonia: Col d’Amieu: Sarraméa, 412 m, 21°4.407S 165°45.674E, piège Malaise, 5–19.xii.2008, 10 ♀(1 MNRJ, 2 BMNH); 11–25.i.2008, 34 ♀; 19.xii.2007– 11.i.2008, 14 ♀, T. Théry. Forêt de Thi: 4.i.1979, P. Fauran, piège lumineux, 1 ♀, Shinonaga det. (Mission D. et L. Matile, xi– xii.1983); x.1979, J. Chazeau, B. Sallé, piège de Malaise, 3 ♀, Shinonaga det.; 250 m, bord de ruisseau, 18– 28.xi.1983, L. Matile, piège de Malaise, 4 ♀, Shinonaga det. (Mission D. et L. Matile, xi–xii.1983). Hienghène: 510 m, 25.xi.1983, L. Matile, 1 ♀, Shinonaga det. (Mission D. et L. Matile, xi–xii.1983); 21–22.ii.1978, S. Shinonaga, 1 ♀, Shinonaga det. Nouméa: v.1955, 2 ♀, J. Rageau, Shinonaga det.; viii.1955, 1 ♂, 3 ♀, J. Rageau, Shinonaga det.; Anse Vata, vii.1958, 2 ♀, J. Rageau, Shinonaga det. Env. [environs] de Nouméa: ix.1955, 2 ♀, J. Rageau, Shinonaga det. Mont Humboldt: 1350 m, 20–22.i.1987, R. et S. Tillier, 2 ♀, Shinonaga det. Rivière Bleue: Parc 4, 164c, 4–20.xii.1985, 1 ♀; 164d, 4 ♀(1 MNRJ); 291a, 01–15.ix.1986, 4 ♀, L.B.dL., J.C, A.S.T.; 291d, 4 ♀(1 BMNH, 1 OUMNH); 291e, 2 ♀; 289a, 23.iv–09.v.1986, 4 ♀, L.B.dL., J.C.; 292a, 01–14.viii.1986, 5 ♀, L.B.dL., J.C.; 292b, 1 ♀; 292c, 2 ♀; 294a, 11–23.iv.1986, 10 ♀, L.B.dL., J.C.; 294b, 2 ♀; 295a, 09– 26.v.1986, 2 ♀, L.B.dL., J.C.; 295d, 09–26.v.1986, 1 ♀; 298c, 20.vi–04.vii.1986, 3 ♀, L.B.dL., J.C.; 299a, 06– 20.ii.1986, 1 ♂, 14 ♀(1 BMNH, 1 OUMNH), L.B.dL., J.C.; 299b, 2 ♀; 300a, 23.i–06.ii.1986, 1 ♀, L.B.dL., J.C.; 301c, 23.i–06.ii.1986, 2 ♀, L.B.dL., J.C.; 302d, 08–23.i.1986, 1 ♀, L.B.dL., J.C.; 302e, 3 ♀; 302f, 1 ♀; 303b, 06–20.vi.1986, 2 ♀, L.B.dL., J.C.; 303c, 1 ♀; Parc 5, 18–20.ii.1986, L. B. de Larbogne & J. Chazeau, piège Malaise, 1 ♀, Shinonaga det.; 13–28.x.1986, 4 ♀; 150 m, 5–20.i.1987, forêt humide alluvions, L. B. de Larbogne, J. Chazeau, A. et S. Tillier, piège Malaise, 4 ♀, Shinonaga det.; 65a, 04–17.vii.1986, 3 ♀, L.B.dL., J.C.; 65c, 2 ♀; 168a, 19.xi–04.xii.1985, 3 ♀, L.B.dL., J.C.; 168b, 2 ♀; 228e, 8 ♀; 230a, 12–27.iii.1986, 1 ♀, L.B.dL., J.C.; 231a, 12–25.xi.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 232b, 20.ii–02.iii.1986, 9 ♀(1 OUMNH, 2 BPBM), L.B.dL., J.C.; 233a, 15–29.ix.1986, 2 ♀, L.B.dL., J.C., A.S.T.; 233b, 1 ♀; 234b, 29.ix–13.x.1986, 6 ♀, L.B.dL., J.C., A.S.T.; 228a, 25.xi–08.xii.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 228b, 2 ♀; 230b, 1 ♀; 230c, 1 ♀; 234a, 29.ix–13.x.1986, 2 ♀(1 MNRJ); 379a, 26.iii–09.iv.1987, 3 ♀, L.B.dL., J.C., A.S.T.; 380b, 09–22.iv.1986, 1 ♀, L.B.dL., J.C.; A.S.T.; 386a, 16.vi–07.vii. 1986, 1 ♀; L.B.dL., J.C., A.S.T. 386c, 2 ♀; 387a, 25.ii– 13.iii.1987, 1 ♀, L.B.dL., J.C.; A.S.T.; Parc6: 160 m, 13–28.x.1986, forêt humide sur alluvions L. B. de L. et J. Chazeau, 1 ♀, Shinonagadet.; Parc 7, 170 m, 19.xi–4.xii.1985, forêt humide sur pente, L.B.de L. & J.Chazeau, 1 ♀, Shinonaga det.; 27, 14.viii–01.ix.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 67a, 07–18.vii1986, 4 ♀, L.B.dL., J.C.; 248a, 29.ix–13.x.1986, 4 ♀, 248b, 1 ♀; 252, 01–04.viii.1986, 2 ♀; 253a, 15–29.ix.1986, 3 ♀, L.B.dL., J.C., A.S.T.; 255a, 26.v–06.vi.1986, 2 ♀; 262a, 01–15.ix.1986, 3 ♀; 262c, 1 ♀; 266a, 12–27.iii.1986, 1 ♀, L.B.dL., J.C.; 272a, 27.iii–11.iv.1986, 8 ♀, L.B.dL., J.C., A.S.T.; 272b, 3 ♀; 272c, 6 ♀; 273c, 11–25.ii.1986, 4 ♀; L.B.dL., J.C., A.S.T.; 265b, 11–23.iv.1986, 7 ♀; 268a, 08–23.i., 1986, 6 ♀(1 BPBM), L.B.dL., J.C.; 269a, 25.xi–08.xii.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 270a, 20.xii–12.xi.1986, 2 ♀, L.B.dL., J.C.; 270b, 1 ♀; 274a, 20.ii–12.iii.1986, 8 ♀, L.B.dL., J.C.; 275a, 06–20.ii.1986, 1 ♀, L.B.dL., J.C.; 275b, 1 ♀; 276a, 23.iv–09.v.1986, 4 ♀, L.B.dL., J.C.; 276b, 6 ♀; 276c, 12 ♀; 276d, 2 ♀; 355a, 20–31.i.1987, 3 ♂(1 MNRJ), 7 ♀(1 MNRJ), L.B.dL., J.C., A.S.T.; 356b, 22.iv–06.v.1987, 6 ♀, L.B.dL., J.C., A.S.T.; 357a, 31.i–12.ii.1987, 4 ♀, L.B.dL., J.C., A.S.T.; 358b, 15.ii– 13.03.1987, 7 ♀, L.B.dL., J.C., A.S.T.; 386b, 16.vi–07.vii.1986, 2 ♀(1 BPBM); L.B.dL., J.C.; Parc7, 174b, 08–22.ii.1989, 6 ♀, L.B.dL., J.C.; 178a, 24.viii–07.ix.1989, 3 ♀, L.B.dL., J.C.; 179a, 12–27.v.1987, 5 ♀, L.B.dL., J.C.; 179b, 3 ♀; 191a, 07–21.xi.1988, 2 ♀, L.B.dL., J.C.; 198, 01– 14.viii.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 199a, 01–15.ix.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 202a, 25.xi– 08.xii.1986, 1 ♀, L.B.dL., J.C., A.S.T.; maquis sur crête, 197a, 28.x– 12.11.1986, 1 ♀(1 OUMNH), L.B.dL., J.C., A.S.T; 197d, 3 ♀; 210b, 28.x–12.xi.1986, 3 ♀, L.B.dL., J.C., A.S.T.; 210c, 4 ♀; 211a, 12–25.xi.1986, 1 ♀, L.B.dL., J.C., A.S.T.; 211b, 2 ♀; 212b, 20.vi–04.vii.1986, 2 ♀, L.B.dL., J.C., A.S.T.; 447b, 09–22.iv.1987, 1 ♀, L.B.dL., J.C., A.S.T., maquis, Sut. C, 27b, 8–25.xii.1986, 2 ♀, L.B.dL., J.C., A.S.T.; forêt de transition à casuarinacées sur pente, 185a, 8–22.ii.1989, 2 ♀, L.B.dL., J.C.; 194c, 07– 21.11.1988, 3 ♀, L.B.dL., J. C. Tiwaka (Poindimié): Gal. Forestière, 20 m, 22–24.xi.1983, L. Matile, 3 ♀, Shinonaga det. (Mission D. et L. Matile, xi– xii.1983). NSMT. New Caledonia: Hienghène: 21–22.ii.1978, 1 ♀, S. Shinonaga; 1 ♀, H. Kurahashi. BMNH. New Caledonia: Nodata, holotype ♀of caledonica, from Bigot’s collection. Tontouta: 4.vi.1925, 1 ♂, P.A. Buxton. MHNG. New Caledonia: Sarraméa, Réserve du Col d’Amieu, 609 m, 21°34.914S, 165°46.376E, Malaise trap, 29.iii-24.iv.2006, 1 ♂, 3 ♀, & 10.iv-21.iv.2006, 1 ♂, C. Mille. BPBM. New Caledonia: Col d’Amieu: NE on route 5 slopes of Mont Rembai, 400–600 m, 23.ix.1979, 1 ♀, W. C. Gagné. Hienghène: 0–50 m, i.1969, 1 ♀; 0–100 m, i.1971, 1 ♀, N.L.H. Krauss. La Crouen: 31.i.1963, 1 ♂, 4 ♀, C.M. Yoshimoto & N.L.H. Krauss; 150 m, 20–22.iii.1968, 1 ♀, Malaise trap, J.L. Gressitt& T.C. Maa. Monts Koghis: 400–600 m, ii.1973, 1 ♀, N.L.H. Krauss; SW slope, 500 m, 19.viii. 1971, light trap, 1 ♂, J. Holloway; 500 m, 2.xii.1963, 1 ♀( MNHN), Malaise trap, R. Straatman. Monts Koghis Road: 100–300 m, i.1985, 1 ♀, N.L.H. Krauss. Nepoui Valley: vii–viii.1940, 1 ♀. Nassirah: 100 m, 20.iii.1968, 1 ♂, J.L. Gressitt& T.C. Maa. Nouméa: I.F.O., Anse Vata, 1.iv.1958, 2 ♂, 10 ♀, J. Rageau; ORSTOM ground, 5 m, 9.viii.1979, 1 ♂, 1 ♀, G.M. Nishida; 25.x.1958, 3 ♀; 27.x.1958, 1 ♀, C.R. Joyce. Pouébo: 20–100 m, 3.ii.1964, 1 ♀; 100 m, 25.i.1964, 1 ♀, R. Straatman. Sarraméa: 70–150 m, i.1971, 1 ♂; 12.ii.1963, 1 ♀( MNRJ), N.L.H. Krauss. Tao: 8–10.ii.1963, 1 ♀, Malaise trap, C.M. Yoshimoto & N.L.H. Krause. Thio: 11.xi.1958, 1 ♀, C.R. Joyce. Yahoue: 100 m, 1 ♀, 25.i.1964, 1 ♀; 60–100 m, ii.1980, 2 ♀; 12.ii.1962, 1 ♀( BMNH), N.L.H. Krauss. Between Plum & Yati: 25.iii.1968, 3 ♀, T.C. Maa. Isle of Pines: Point SW of Kuto, 0–5 m, 17.viii.1979, 1 ♀, 25 Gardens Roadside, G.A. Samuelson. Loyalty Islands: Mare: La Roche, iii.1959, 1 ♂, 1 ♀, N.L.H. Krauss.
Comments.The species was described from New Caledoniabased on one female. The holotypewas reviewed by Stein (1907a), Crosskey (1962)and Pont (2000). Paramonov (1961)described the male, recorded from Nouméa, and added some characters to the original description. The male was keyed but the author considered females impossible to distinguish from females of P. apicalisSchinerand P. australisMalloch. Crosskey (1962)redescribed the species. He examined only females, but keyed the male based on Paramonov’s (1961)description. In the key, the couplet that segregates this species from P. maculigerais based only on distribution (from New Caledoniaor not). Comparison of the male abdomen of both species shows that they are very similar, especially the keel with long setae on tergite 4, but the long setae on the ventral margin of tergite 4 are all long in P. caledonicaand the straight setae on the disc of the epandrium are also more numerous and longer in this species (compare Fig. 10with Crosskey’s (1962)Fig. 47). The species was very abundant in the material examined, but almost all the specimens were females: of the 400 specimensexamined, only 12 were males.
Distribution. New Caledonia( New Caledonia, Isle of Pines, Loyalty Is),? Vanuatu.
2008-12-05
2008-12-19
2008-12-05
MNHN
Malaise
New Caledonia
412
-21.07345
Col d'Amieu
1
165.76123
Sarramea
47
48
1
[503,983,1957,1983]
2008-01-11
2008-01-25
2008-01-11
MNRJ, BMNH
New Caledonia
47
48
10
10
1979-01-04
New Caledonia
Thery. Foret de Thi
47
48
14
14
1983-11-25
New Caledonia
510
Hienghene
48
49
1
1987-01-20
1987-01-22
1987-01-20
New Caledonia
Mont Humboldt
1350
Rageau
48
49
2
2
[699,1126,375,402]
1986-09-01
1986-09-15
1986-09-01
MNRJ
New Caledonia
48
49
4
4
[222,617,413,439]
BMNH, OUMNH
New Caledonia
48
49
4
4
[285,937,525,551]
BMNH, OUMNH
D.L. Bickel & J. Chazeau
New Caledonia
48
49
15
14
1
1986-09-15
1986-09-29
1986-09-15
OUMNH, BPBM
D.L. Bickel & J. Chazeau
New Caledonia
48
49
9
9
[422,873,861,887]
1987-03-26
1987-04-09
1987-03-26
MNRJ
New Caledonia
48
49
2
2
[289,936,935,962]
1986-10-13
1986-10-28
1986-10-13
D.L. Bickel & J. Chazeau & A. et S. Tillier
New Caledonia
160
Parc
48
49
1
1
[265,876,973,999]
1985-11-19
1985-12-04
1985-11-19
New Caledonia
Shinonaga
7
Parc
48
49
1
1
1986-11-25
1986-12-08
1986-11-25
BPBM
D.L. Bickel & J. Chazeau
New Caledonia
48
49
6
6
[1060,1239,1271,1298]
MNRJ
New Caledonia
48
49
3
3
1987-04-22
1987-05-06
1987-04-22
MNRJ
D.L. Bickel & J. Chazeau & A. et S. Tillier
New Caledonia
48
49
7
7
1989-02-08
1989-02-22
1989-02-08
BPBM
D.L. Bickel & J. Chazeau
New Caledonia
Parc
48
49
2
2
OUMNH
D.L. Bickel & J. Chazeau & J.A. Scopoli
New Caledonia
48
49
1
1
1983-11-22
1983-11-24
1983-11-22
D.L. Bickel
New Caledonia
20
Gal. Forestiere
48
49
3
3
[198,628,1757,1783]
NSMT
New Caledonia
Hienghene
48
49
1
1925-06-04
BMNH
P. A. Buxton
New Caledonia
No
48
49
2
1
1
holotype
2006-03-29
2006-04-24
2006-03-29
MHNG
Reserve du Col d'Amieu & C. Mille
New Caledonia
609
-21.5819
Sarramea
1
165.77293
Reserve du Col d'Amieu
48
49
5
3
2
1979-09-23
BPBM
Col d'Amieu
New Caledonia
500
Mont Rembai
48
49
1
1
[733,980,1981,2007]
1971-01
BPBM
Col d'Amieu
New Caledonia
50
true
Mont Rembai
48
49
1
1
1963-01-31
BPBM
New Caledonia
49
50
Krauss. La Crouen
48
49
5
4
1
1973-02
BPBM
J. L. Gressitt
New Caledonia
500
Maa. Monts Koghis
49
50
1
1
[151,698,226,252]
1963-12-02
MNHN
Holloway
New Caledonia
500
49
50
1
1
[297,874,263,290]
1940-07
1940-08-31
1940-07
MNHN
New Caledonia
Krauss. Nepoui Valley
49
50
1
1
1958-04-01
MNHN
J. L. Gressitt & T. C. Maa. Noumea
New Caledonia
Anse Vata
49
50
12
10
2
[637,815,338,364]
1958-10-27
MNHN
J. L. Gressitt & T. C. Maa. Noumea
New Caledonia
true
Anse Vata
49
50
1
1
1964-01-25
MNHN
J. L. Gressitt & T. C. Maa. Noumea
New Caledonia
100
true
Anse Vata
49
50
1
1
[956,1244,375,402]
1963-02-12
MNRJ
J. L. Gressitt & T. C. Maa. Noumea
New Caledonia
100
true
Anse Vata
49
50
1
1
1963-02-08
1963-02-10
1963-02-08
MNRJ
New Caledonia
Krauss. Tao
49
50
1
1
[923,1235,450,476]
1962-02-12
1963-02-10
1963-02-08
BMNH
New Caledonia
true
Krauss. Tao
49
50
1
1
[624,1428,487,514]
1979-08-17
BMNH
T. C. Maa.
New Caledonia
3
SW of Kuto
49
50
1
1
Isle of Pines
[151,1244,525,551]
1959-03
BMNH
Gardens Road & G. A. Samuelson. & La Roche
New Caledonia
Loyalty Islands
49
50
2
1
1
Province des Iles