Subepimeria geodesiae Bellan-Santini, 1972: 225 Epimeria puncticulata Watling & Holman 1981: 213–215 E. ( Subepimeria ) iota Coleman 1998b: 223–224 Subepimeria geodesiae De Broyer 1983: 305 Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea) d’Acoz, Cédric d’Udekem Verheye, Marie L. European Journal of Taxonomy 2017 2017-10-17 359 1 553 6G5VH Bellan-Santini, 1972 Bellan-Santini 1972 [460,1128,267,294] Malacostraca Epimeriidae Epimeria GBIF Animalia Amphipoda 142 143 Arthropoda species geodesiae Subepimeria      Subepimeria geodesiae Bellan-Santini, 1972: 225, pls 33–34.    Epimeria puncticulata–  Watling & Holman 1981: 213–215(discussion, in part), not fig. 21 (=  E.( Subepimeria) iota sp. nov.);  Coleman 1998b: 223–224(in part).    Subepimeria geodesiae–  De Broyer 1983: 305(discussion).    Description Description based on the illustrations of Bellan-Santini (1972). ROSTRUM. Reaching tip of article 1 of peduncle of antenna 1, broad in lateral view. EYES. Large, rounded. PEREION–PLEOSOME TOOTH PATTERN. Pereionites 1–7 totally smooth; pleonite 1 toothless; pleonite 2 with small, posterodorsal tooth; pleonite 3 with posterodorsal tip forming a small, distinct rounded (very broadly triangular) lobe projecting backwards. COXAE 1–3. Tip subacute in coxae 1–2, blunt in coxa 3. COXA 4. Very narrow; anterodorsal border weakly convex, directly followed by very broadly rounded ventral lobe (anteroventral border absent); posteroventral border nearly straight (weakly concave); ratio length of posterodorsal border / length of posteroventral border: 0.5. COXA 5. Very broad, posteroventral corner with broadly rounded angular discontinuity. COXA 6. Posterior border regularly rounded. COXA 7. Posterior border nearly straight (very weakly convex); posteroventral corner broadly rounded. EPIMERAL PLATES 1–3. Posteroventral angle angulate in plate 1, produced into a small tooth in plate 2; produced into a medium-sized tooth in plate 3. UROSOME TOOTH PATTERN. Urosomite 1 with narrow triangular dorsal process; urosomite 3 with dorsolateral borders straight, with tip produced into a long tooth pointing obliquely. TELSON. Cleft on 0.25. GNATHOPODS 1–2. Carpus and propodus of normal slenderness; propodus not narrowing distally, and palm distinct. PEREIOPOD 5. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterodistal corner forming a blunt-tipped broadly triangular process (acute angle) projecting backwards; merus, carpus and propodus stout. PEREIOPOD 6. Basis of normal width, with posteroproximal process indistinct (reduced to very low proximal dilatation in continuity with the more distal part of the posterior border), with posterior border nearly straight and parallel to anterior border, with posterodistal corner forming a blunt-tipped broadly triangular process (acute but nearly squared angle) scarcely projecting backwards; merus, carpus and propodus stout.   Table 1.(continued on next page). Tentative table of character states and distribution for  Subepimeriaspecies.     GROUPS OF  SUBEPIMERIA     Epimeria (Subepimeria)group  adeliae    Epimeria (Subepimeria)group  puncticulata   Pereionite 7 with posterodorsal bump or reduced tooth without posterodorsal bump or tooth   Pleonite 1 with well-developed sharp tooth toothless or with slight bump or with trace of tooth    SUBEPIMERIAGROUP ADELIAE   E. (Subep.) adeliae sp. nov.   E. (Subep.)sp. 1   Pleonite 3 posterodorsal process distinctly triangular, forming a squared angle regularly rounded   Distribution Adélie Coast South Georgia, «Scotia Arc»    SUBEPIMERIAGROUP PUNCTICULATA   E. (Subep.) geodesiae  E. (Subep.) iota sp. nov.  E. (Subep.) puncticulata  E. (Subep.) teres sp. nov.  E. (Subep.) urvillei sp. nov.   Rostrum shape, in lateral view broad broad very narrow very narrow narrow   Rostrum size (better seen in dorsal view) reaching tip of art. 1 of peduncle of antenna 1 not reaching tip of art. 1 of peduncle of antenna 1 — reaching tip of art. 1 of peduncle of antenna 1 not reaching tip of art. 1 of peduncle of antenna 1   Eye medium-sized, rounded large, broadly elliptic, nearly rounded medium-sized, large, broadly narrowly elliptic, nearly elliptic rounded large, narrowly elliptic   Coxa 4 anterior margin forming a weak with distinct curve, which angular dramatically discontinuity increases near of ventral 0.6 its tip with distinct angular discontinuity of ventral 0.6 forming a regular curve, anteroventral part very weakly arching backwards forming a regular curve, anteroventral part weakly arching backwards   Coxa 4 ventral corner forming a very forming a blunt broad curve squared angle forming a sharp squared angle forming a blunt acute angle forming a very distinct acute (nearly squared) angle   Basis of pereiopod 5, posterodistal corner produced into a broadly triangular tooth produced into a broadly triangular tooth produced into a narrowly triangular tooth produced into produced into a narrowly a narrowly triangular triangular tooth tooth   Table 1.Tentative table of character states and distribution for  Subepimeriaspecies.      SUBEPIMERIAGROUP PUNCTICULATA   E. (Subep.)  geodesiae  E. (Subep.)  iota sp. nov.  E. (Subep.)  puncticulata  E. (Subep.)  teres sp. nov.  E. (Subep.)  urvilleisp. nov.   Basis of pereiopod 6, posterodistal corner produced into a sharp squared angle produced into a sharp squared angle in adults, into a blunt one in immatures produced into a sharp acute (nearly squared) angle produced into a sharp acute (nearly squared) angle produced into an acute angle   Basis of pereiopod 7, posterodistal corner produced into a sharp tooth forming a squared angle produced into a blunt obtuse angle produced into a blunt obtuse angle produced into a sharp tooth forming a squared angle produced into a blunt obtuse angle   Posterodorsal corner of pleonite 3 with rounded or very broadly triangular protrusion pointing backwards with rounded or very broadly triangular protrusion very weakly pointing backwards with very broadly triangular protrusion pointing backwards with very broadly triangular protrusion pointing backwards with bluntly triangular protrusion strongly pointing backwards   Dorsal protrusion of urosomite 1 fairly strong fairly weak to weak weak fairly strong fairly strong   Distribution Adélie Coast Elephant Island to tip of Antarctic Peninsula  western Ross Sea western Weddell Sea Adélie Coast PEREIOPOD 7. Basis broad; posterior border weakly convex, with shallow concavity in distal 0.8, terminated into a sharp triangular tooth (forming a squared angle).   Body length  15 mm.    Distribution Adélie Coast: CapeGéodésie, 115–135 m, on coarse sand with abundant epifauna: hydroids, bryozoans and sponges ( Bellan-Santini 1972).    Remarks  Watling & Holman (1981)synonymised  Subepimeria geodesiaewith  Epimeria puncticulata, which they considered as a widely distributed variable species. Molecular data (COI, 28S) revealed different species within  E. puncticulata s. lat.( Verheye et al.2016a; this paper Fig. 342), leading to the resurrection of  Subepimeria, as a subgenus. The shape of coxa 4 of  E. geodesiaeas illustrated by Bellan-Santini (1972)is unique and does not fit with the descriptions of the holotypeof  E. puncticulata(K.H. Barnard 1930; Coleman 1994) or any other  Subepimeriaspecies.  Epimeria geodesiaeis considered here to be a valid species.