Review of the genus Omalium Gravenhorst, 1802 (Coleoptera: Staphylinidae: Omaliinae: Omaliini) of China Shavrin, Alexey V. Zootaxa 2025 2025-02-18 5588 4 501 543 Shavrin 2025 [225,460,1785,1812] Insecta Staphylinidae Omalium Animalia Coleoptera 22 523 Arthropoda species schuelkei sp. nov.  ( Figs 21, 48, 51, 56–58, 62)   Type material examined.  Holotype ♂:  CHINA: GANSU:‘CHINA, Gansu, Xinlong| Shan, cca  70 kmS| Lanzhou, 2225- |  2380m,  7.VIII.1994| A. Smetana[C32]’ <printed>, ‘HOLOTYPE |  Omalium|  schuelkei  sp. nov.| Shavrin A.V. des. 2025’ <red, printed> ( CNC).    Paratypes( 13 ♂♂, 15 ♀♀):  CHINA: GANSU: 5 ♂♂(one specimen dissected), 4 ♀♀(one specimen dissected): same data as that in the holotype ( 2 ♂♂, 2 ♀♀: cSh; 3 ♂♂, 3 ♀♀: CNC);  7 ♂♂( two specimensdissected), 7 ♀♀: ‘ CHINA: Gansu Prov.[CH11-27] | Lenglong Ling Mts., 60 km NNW| Honggu, Jin Sha Gorge, mixed forest | (  Picea, Populus, Betula) 36°51´56.9´´N, | 102°38´55.3´´E, litter, moss, mushrooms | sifted,  12.VII.2011, leg. M. Schülke’<printed>, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> ( 3 ♂♂, 2 ♀♀: cSh;  4 ♂♂, 6 ♀♀: cSch);  1 ♀: ‘  CHINA: Gansu province, | XIAHE(= Labrang) env., | 35°11.3´´N 102°30.6´E, |  3043 m( GPS), 19.– 22.VI. | 2005, J. Hájek, D. Král& | J. Růžičkaleg. [Ch 2]’ <printed>, ‘baited pitfall traps(fish | meat + cheese) with | ethylene glycol. | margin of a wet | coniferous forest’ <printed>, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> ( cSch);   SICHUAN: 1 ♂(dissected), 2 ♀♀: ‘ CHINA: N-Sichuan [CH12-21] | 49 km N Songpan, road S 301 km 114, | N Gonganglingpass, 33°04´31´´N, | 103°42´38´´E,  3230 m, spruce forest, | litter, moss & mushrooms sifted, |  9.VIII.2012, M. Schülke’<printed>, ‘Museum für Naturkunde | Berlin | Sammlung M. Schülke’ <printed> ( cSch);  1 ♀: ‘ CHINA: W-Sichuan1999 | Ganzi Tibet Aut. Pref., Luding Co.| W Erlangshan-Pass,  2600 m| 7 km SSE Luding, 29°51N| 102°15E, Nadelstreu, Pilze | 29. VI., leg. M. Schülke’<printed>, ‘Sammlung | M. Schülke | Berlin’ <printed>, ‘ Omalium[handwritten] | sp. [handwritten] | det. M. Schülke 2002’ <printed> ( cSch).All paratypeswith additional red printed label: ‘ PARATYPE|  Omalium|  schuelkei  sp. nov.| Shavrin A.V. des. 2025’.   Description.Measurements (n=29): HW: 0.55–0.60; HL: 0.36–0.38; OL: 0.15–0.16; TL: 0.05–0.07; AL ( holotype): 0.87; PL: 0.46–0.52; PWmax: 0.72–0.80; PWmin: 0.65–0.70; ESL: 0.85–0.97; EW: 0.95–1.05; MTbL ( holotype): 0.47; MTrL ( holotype): 0.27 (MTrL 1–4: 0.10; MTrL 5: 0.17); AW: 1.02–1.06; AedL: 0.60–0.62; BL: 2.65–3.50 ( holotype: 3.25). Habitus as in Fig. 51. Body reddish-brown, with darker head, pronotum and abdomen (lateral and basal portions of pronotum, elytra, paratergites and intersegmental membranes of abdomen in some paratypesdistinctly paler); antennomeres 6–11 or 7–11 brown; mouthparts, antennomeres 1–5 or 1–6 and legs yellow-brown; tarsi yellow. Punctation of head dense or sparse, fine, usually denser in middle, denser and coarser on infraorbital portions; punctation of neck dense, about as that in middle of head; punctation of pronotum moderately dense, usually finer and sparser in medioapical, lateral and mediobasal portions and denser in middle; abdominal tergites with indisinct, fine and sparse punctation. Anterior portion of clypeus with distinct transverse microsculpture, basal portions of clypeus with fine transverse and longitudinal meshes, middle portion sometimes with longitudinal meshes between anteocellar foveae, posterior parts of infraorbital portions usually with distinct longitudinal and diagonal microreticulation; middle part of neck without or with fine transverse or isodiametric meshes; scutellum with fine isodiametric sculpture; abdominal tergites with dense isodiametric microsculpture. Anterior and posterior margins of pronotum with row of short cuticular fringe, invisible in some specimens. Head 1.5 times as broad as long, with wide and moderately deep anteriomedian depressions, reaching level of anterior third of eyes; posteriolateral margins of clypeus subparallel, stretching posteriad toward level about middle length of eyes. Latero-apical margin between anterior margin of eyes and clypeus with distinct semicircular notch. Dorsal surface sometimes with fine irregular transverse elevations between punctures in middle, infraorbital portions with irregular elongate wrinkles in postocular portions. Anteocellar foveae suboval and deep, slightly convergent latero-apicad toward level of posterior third or middle length of eyes. Temples very short, from two to three times shorter than longitudinal length of eyes. Apical part of neck sometimes distinctly and narrowly depressed, with irregular short elevations between punctures. Ocelli located slightly below level of posterior margins of eyes; distance between ocelli 1.4–1.6 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about three times as long as penultimate segment. Antenna with slightly transverse antennomeres 6–7 and distinctly transverse 8–10; 5 indistinctly longer and broader than 4, 6 slightly broader than 5, 7 broader than 6, 8 slightly broader than 7, 9–10 slightly longer and distinctly broader than 8, apical antennomere about twice as long as penultimate antennomere. Pronotum 1.5 times as broad as long, 1.3 times as broad as head, widest in or slightly in front of middle, distinctly more narrowed posteriad than anteriad. Apical angles widely rounded, not protruded anteriad. Anterior margin somewaht straight or rounded, not protruded anteriad, sometimes indistinctly concave in middle. Surface of disc with two long, deep and moderately wide depressions. Each mediolateral portion sometimes with distinct narrow elongate elevations, slightly curved posteriad and reaching middle length of pronotum. Middle portion without elevations between punctures. Elytra 1.1 times as broad as long, 1.8 times as long as pronotum. Mediolateral surface of each elytron with irregular longitudinal and diagonal elevations.   FIGURES 62.Type localitiy of  Omalium schuelkeiin Sichuan, China (photograph: M. Schülke). Metatarsi slightly less than twice shorter than metatibia. Abdomen slightly broader than elytra. Male. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII widely sinuate.Aedeagus with wide basal portion, gradually narrowed toward median lobe; median lobe wide and moderately long, from widest basal part gradually narrowed toward small rounded apex; accessory plates short and relatively narrow, each with bilobed apex; parameres relatively narrow, distinctly broadened in preapical parts, reaching about middle of median lobe, each with two moderately long and two short apical setae; internal sac long and moderately narrow, with two small elongate sclerotized structures in basal part ( Fig. 56). Lateral aspect of the aedeagus as in Fig. 57; apical portion of median lobe (lateral view) with distinctly truncate ventrolateral margin ( Fig. 58). Female. Posterior margin of abdominal tergite VIII truncate. Posterior margin of abdominal sternite VIII truncate or rounded.Accessory sclerite with wide basal part, gradually anrrowed toward broadened and elongate median part with rounded apex ( Fig. 21). Spermatheca not recognized.   Comparative notes.Based on the general shape of the body and the aedeagus, the shape of the elliptic anteocellar foveae,  O. schuelkei  sp. nov.is similar to the Holarctic  O. oxyacanthae. The new species can be distinguished from it by the larger body (the body of  O. oxyacanthaevaries from 2.2 to 2.6 mm), denser and coarser punctation of the pronotum and the elytra, broader and shorter median lobe, slightly broader parameres, and different shape of the female accessory sclerite.   Distribution.  Omalium schuelkei  sp. nov.is known from several localities from Gansuand Sichuan, China( Fig. 48).   Etymology.Patronymic; the species is named to honour Michael Schülke ( Berlin), one of the collectors of the typespecimens.   Bionomics.Specimens were collected at elevations from 2225 to 3230 ma.s.l. and were taken by sifting litter, mosses and mushrooms in mixed (  Picea, Populus, Betula) and coniferous forests. One specimenfrom Xiahe was collected using pitfall traps. The photograph of the typelocalities in Sichuan as in Fig. 62( CH12-21). 5069426335 1994-08-07 CNC, cSh A. Smetana China 2380 22 524 Xinlong 22 523 48 24 24 GANSU holotype 5069426365 1994-08-07 CNC, cSh A. Smetana China 2380 23 524 Xinlong 22 523 48 24 24 GANSU holotype 5069426375 2011-07-12 cSh M. Schulke' China 23 524 36.865803 Honggu 1 102.64869 Lenglong Ling Mts. 23 523 21 9 10 Gansu Prov. 5069426322 2011-07-12 cSch M. Schulke' China 23 524 36.865803 Honggu 1 102.64869 Lenglong Ling Mts. 23 523 10 6 4 Gansu Prov. 5069426310 2005-06-19 2005-06-22 GPS, cSch J. Hajek & D. Kral & J. Ruzicka China 3043 23 524 35.11 XIAHE 102.51 Labrang 23 523 1 1 Gansu province 5069426361 2012-08-09 cSch M. Schulke' China 3230 23 524 33.075275 Gongangling 20 103.710556 Songpan 23 523 3 2 1 SICHUAN 5069426364 cSch M. Schulke' China Luding Co. 2600 23 524 29.85 W-Sichuan 1224 102.25 Erlangshan-Pass 23 523 1 1 Ganzi Tibet Aut. Pref